Anhangueria
| Anhanguerians | |
|---|---|
| |
| Snouts of the hamipterids Iberodactylus (left) and Hamipterus (right)
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anhanguerid Guidraco
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Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Order: | †Pterosauria |
| Suborder: | †Pterodactyloidea |
| Clade: | †Ornithocheirae |
| Clade: | †Anhangueria Rodrigues & Kellner, 2013 |
| Subgroups | |
Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade
Cimoliopterus cuvieri.[1]
Classification
Anhangueria originally only contained the
genera Brasileodactylus, Camposipterus, Cearadactylus, Ludodactylus as well as the family Anhangueridae,[1] however, recent analyses had recovered the family Hamipteridae within this clade as well.[2][3][4][5][6]
In 2014, paleontologist
phylogenetic analysis
by Andres and colleagues is shown below:
Lanceodontia
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Later, in 2019, a different phylogenetic analysis, this time conducted by Borja Holgado and colleagues, focused on derived
pterodactyloids with particular emphasis on anhanguerians. In this analysis, Anhangueria consisted of Camposipterus and the families Anhangueridae and Hamipteridae, the clade Ornithocheirae was recovered as a more inclusive group consisting of Ornithocheirus and Cimoliopterus as basal members, as well as the Anhangueria in the most derived position. This classification is supported by the expansion of the premaxillary tip (a spoon-like expansion at the end of the snout) with a high jaw end.[2] Many analyses afterwards have followed this concept,[3][5][10] although some had recovered Camposipterus outside the Anhangueria (within the clade Targaryendraconia), but still within the Ornithocheirae.[4][6]
The cladogram of the analysis by Holgado and colleagues in shown below:
Caulkicephalus trimicrodon
Anhanguera (G and H), and Uktenadactylus
(I and J)| Ornithocheirae |
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