Araeoscelidia
Araeoscelidans | |
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Life restoration (top) and skull reconstruction (bottom) of Petrolacosaurus kansensis
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Diapsida |
Order: | †Araeoscelidia Williston, 1913 |
Genera | |
Araeoscelidia or Araeoscelida is a
Description
Araeoscelidians were small animals (less than one meter in length) looking somewhat like lizards, though they are only distantly related to true lizards. They differ from other, earlier
Genera
Araeoscelidia includes well-known genera such as Araeoscelis Williston 1910,[1][2] Petrolacosaurus Lane 1945[3][4] and Spinoaequalis,[5][6] known from virtually complete skeletons. Zarcasaurus,[7] Aphelosaurus[8][9][10] and Kadaliosaurus[11] belong to this clade but are known only from post-cranial remains and a mandible fragment for Zarcasaurus.
The genus Dictybolos has been included in Araeoscelidia by Olson (1970)[12] but this inclusion has been criticized e.g., by Evans (1988),[13] especially since Olson also included distantly related groups such as protorosaurs and mesosaurs.
New specimens have been discovered in the United States state of Oklahoma,[14][15] but lack a scientific description as of 2023.
Phylogeny
The majority of phylogenetic studies recover araeoscelidians as the most basal group of diapsids; however, Simões et al. (2022) recover them as stem-amniotes instead, as the sister group to the clade including Captorhinidae and Protorothyris archeri.[16]
Stratigraphic and geographic distribution
Araeoscelidia are known from the Late Carboniferous in the United States (Petrolacosaurus, Spinoaequalis) to the Early Permian in France (Aphelosaurus), Germany (Kadaliosaurus) and the United States (Dictybolos, Zarcasaurus, Araeoscelis, Halgaitosaurus[17]). Apart from araeoscelidans, only one other diapsid is known before the Late Permian: Orovenator from the Early Permian of Oklahoma.[18]
References
- ^ Vaughn 1955
- ^ Reisz, Berman & Scott 1984
- ^ Peabody 1952
- ^ Reisz 1981
- ^ deBraga & Reisz 1995
- ^ deBraga & Rieppel 1997
- ^ Brinkman, Berman & Eberth 1984
- ^ Gervais 1859
- ^ Thévenin 1910
- ^ Falconnet & Steyer 2007
- ^ Credner 1889
- ^ Olson 1970
- ^ Evans 1988
- ^ May & Hall 2002
- ^ Swanson & Carlson 2002
- PMID 35984885.
- ISSN 0097-4463.
- ^ Reisz, Modesto & Scott 2011
Further reading
- Carroll, Robert L. (1988). Vertebrate Paleontology and Evolution. New York: W.H. Freeman and Co. ISBN 0-7167-1822-7.
- ISBN 0-632-05614-2.
- deBraga, M. & Reisz, R. R. (1995). "A new diapsid reptile from the uppermost Carboniferous (Stephanian) of Kansas". Palaeontology. 38: 199–212.
- deBraga, M. & Rieppel, O. (1997). "Reptile phylogeny and the interrelationships of turtles". Zoological Journal of the Linnean Society. 120 (3): 281–354. .
- Brinkman, D. B.; Berman, D. S. & Eberth, D. Z. (1984). "A new araeoscelid reptile, Zarcasaurus tandyderus, from the Culter Formation, (Lower Permian) of north-central New Mexico" (PDF). New Mexico Geology. 6 (2): 34–39.
- Credner, H. (1889). "Die Stegocephalen und Saurier aus dem Rothliegenden des Plauen'schen Grundes bei Dresden. 8 – Kadaliosaurus priscus Cred". Zeitschrift der Deutschen Geologischen Gesellschaft. 41: 319–342.
- Evans, S. E. (1988). "The early history and relationships of the Diapsida". In Benton, M. J. (ed.). The phylogeny and classification of the tetrapods. Vol. 1: Amphibians, Reptiles, Birds. Systematics Association Special Volume 35 A. Oxford: Clarendon Press. pp. 221–260. ISBN 0-19-857705-2.
- Falconnet, J. & Steyer, J.-S. (2007). "Revision, osteology and locomotion of Aphelosaurus, an enigmatic reptile from the Lower Permian of France". Journal of Morphology (abstract of the 8th International Congress of Vertebrate Morphology, Paris, July 2007): 38.
- Gervais, P. (1859). Zoologie et paléontologie française (2nd ed.). Paris: Bertrand.
- .
- May, W. J. & Hall, J. D. (2002). "Geology and vertebrate fauna of a new site in the Wellington Formation (Lower Permian) of Northern Oklahoma". Oklahoma Geology Notes. 62 (2): 63–66.
- Olson, E. C. (1970). "New and little known genera and species of vertebrates from the Lower Permian of Oklahoma". Fieldiana: Geology. 18: 359–434.
- Peabody, F. E. (1952). "Petrolacosaurus kansensis Lane, a Pennsylvanian reptile from Kansas". University of Kansas Paleontological Contributions. 10: 1–41. hdl:1808/3785.
- Reisz, R. R. (1981). "A diapsid reptile from the Pennsylvanian of Kansas". Special Publication of the Museum of Natural History, University of Kansas. 7: 1–74.
- Reisz, R. R.; Berman, D. S. & Scott, D. (1984). "The anatomy and relationships of the Lower Permian reptile Araeoscelis". Journal of Vertebrate Paleontology. 4 (1): 57–67. .
- Reisz, Robert R.; Modesto, Sean P.; Scott, Diane M. (2011). "A new Early Permian reptile and its significance in early diapsid evolution". Proceedings of the Royal Society B. 278 (1725): 3731–3737. PMID 21525061.
- Swanson, B. A. & Carlson, K. J. (2002). "Walk, Wade, or Swim? Vertebrate Traces on an Early Permian Lakeshore" (PDF). S2CID 130593903.
- Thévenin, A. (1910). "Les plus anciens quadrupèdes de France". Annales de Paléontologie. 5: 1–65.
- Vaughn, P. P. (1955). "The Permian reptile Araeoscelis restudied". Bulletin of the Harvard Museum of Comparative Zoology. 113: 305–467.