Archaeidae

Source: Wikipedia, the free encyclopedia.

Assassin spiders
Temporal range: Middle Jurassic–Recent
Austrarchaea sp.
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Infraorder: Araneomorphae
Superfamily: Palpimanoidea
Family: Archaeidae
C. L. Koch & Berendt, 1854
Diversity
5 genera, 90 species

Archaeidae, also known as assassin spiders and pelican spiders, is a spider family with about ninety described species in five genera.[1] It contains small spiders, ranging from 2 to 8 millimetres (0.079 to 0.315 in) long, that prey exclusively on other spiders.[2] They are unusual in that they have "necks", ranging from long and slender to short and thick. The name "pelican spider" refers to these elongated jaws and necks used to catch their prey. Living species of Archaeidae occur in South Africa, Madagascar and Australia, with the sister family Mecysmaucheniidae occurring in southern South America and New Zealand.[1]

Assassin spiders were first known from 40 million-year-old amber fossils which were found in Europe in the 1840s and were not known to have living varieties until 1881, when the first living assassin spider was found in Madagascar.[3]

The fossil record of this family was first identified from

Afarchaea grimaldii was described from Cretaceous Burmese amber aged between 88 and 95 million years, extending the record of this group considerably, the oldest fossil known of the group is Patarchaea muralis from the Middle Jurassic (Oxfordian/Callovian) of Inner Mongolia, China.[4]

Taxonomy

The family Archaeidae was erected in 1854 by

C. L. Koch and Berendt[1] for one genus, Archaea, initially with three extinct species,[3] all found in amber from the Baltic Sea or Bitterfeld in Saxony-Anhalt
, Germany. No living species are placed in this genus.

Extant genera

As of April 2019[update], the World Spider Catalog accepts the following genera:[1]

Fossils

Fossils found in amber, particularly from the Baltic and Myanmar (Burma), have been assigned to a number of extant and extinct genera placed in the family Archaeidae.[5] The extinct species

Mya.[6] Jurarchaea zherikhini Eskov, 1987 was previously considered a member of this family. Still, it is more likely a holarchaeid or a pararchaeid.[citation needed
]

Phylogeny

A 2012 Bayesian phylogenetic analysis of 15 Archaeidae species, using combined molecular and morphological data, produced the cladogram shown below. Species representing modern genera found in Africa and Madagascar (Afrarchaea and Eriauchenius) were not resolved into monophyletic groups; Zephyrarchaea had not then been split off from Austrarchaea. The species found in European amber formed a clade, whereas Burmesarchaea grimaldii, from Burmese amber, appeared to be basal to modern genera.[7]

Archaeidae

Austrarchaea (including Zephyrarchaea)

Afrarchaea + Eriauchenius

Burmesarchaea grimaldii

Archaea paradoxa

Baltarchaea conica

Myrmecarchaea sp.

Patarchaea muralis

See also

References

  1. ^ a b c d "Family: Archaeidae C. L. Koch & Berendt, 1854". World Spider Catalog. Natural History Museum Bern. Retrieved 2019-04-19.
  2. ^ Wood HM, Scharff N (2018). "A review of the Madagascan pelican spiders of the genera Eriauchenius O.". ZooKeys.
  3. ^ a b Koch, C.L. & Berendt, G.C. (1854). Die I'm Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt [The organic remains of antiquity found in amber] (in German). Berlin.{{cite book}}: CS1 maint: location missing publisher (link)
  4. S2CID 73616235
    .
  5. ^ Dunlop, J.A.; Penney, D. & Jekel, D. (2016), "A summary list of fossil spiders and their relatives, version 17.5" (PDF), World Spider Catalog, Natural History Museum Bern, retrieved 2016-10-13
  6. S2CID 56446480
    , retrieved 2016-10-13
  7. .