Bengal tiger
Bengal tiger | |
---|---|
Adult male in Kanha Tiger Reserve, India | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Suborder: | Feliformia |
Family: | Felidae |
Subfamily: | Pantherinae |
Genus: | Panthera |
Species: | P. tigris |
Subspecies: | P. t. tigris
|
Population: | Bengal tiger |
Range of Bengal tiger in red |
The Bengal tiger is a
The tiger is estimated to have been present in the
The Bengal tiger's historical range covered the Indus River valley until the early 19th century, almost all of India, Pakistan, southern Nepal, Bangladesh, Bhutan, and southwestern China. Today, it inhabits India, Bangladesh, Nepal, Bhutan, and southwestern China.[6] India's tiger population was estimated at 2,603–3,346 individuals by 2018.[9] Around 300–500 individuals are estimated in Bangladesh by 2015,[8] 355 in Nepal by 2022,[10] and 90 individuals in Bhutan by 2015.[11]
Taxonomy
Felis tigris was the
The
Results of a genetic analysis of 32 tiger samples indicate that the Bengal tiger samples grouped into a different clade than the Siberian tiger samples.[16]
Genetic ancestry
The Bengal tiger is defined by three distinct
Characteristics
The Bengal tiger's coat is yellow to light orange, with stripes ranging from dark brown to black; the belly and the interior parts of the limbs are white, and the tail is orange with black rings. The
The greatest skull length of a tiger is 351 mm (13.8 in) in males and 293 mm (11.5 in) in females.[21] It has exceptionally stout teeth. Its canines are 7.5 to 10 cm (3.0 to 3.9 in) long and thus the longest among all cats.[22]
Body weight and size
Males and female Bengal tigers in Panna Tiger Reserve reach a head-to-body length of 183–211 cm (72–83 in) and 164–193 cm (65–76 in) respectively, including a tail about 85–110 cm (33–43 in) long. Total length ranges from 283 to 311 cm (111 to 122 in) for male tigers and 255–285 cm (100–112 in) for female tigers.[23] They typically range from 90–110 cm (35–43 in) in shoulder height.[24]
Subadult males weigh between 130 and 170 kg (290 and 370 lb) and reach 200–260 kg (440–570 lb) when adult; subadult females weigh 80–100 kg (180–220 lb) and reach between 110 and 180 kg (240 and 400 lb) when adult.[25] In central India, 42 adult male Bengal tigers weighed on average 190 kg (420 lb) with a range of 167–234 kg (368–516 lb); their total length was 282 cm (111 in) with a range of 267–312 cm (105–123 in), and their average shoulder height was 99 cm (39 in); 39 adult female Bengal tigers weighed an average of 132 kg (291 lb) with a maximum of 156 kg (344 lb) and an average total length of 254 cm (100 in) ranging from 239 to 277 cm (94 to 109 in).[26] Several scientists indicated that adult male Bengal tigers in the Terai consistently attain more than 227 kg (500 lb) of body weight. Seven adult males captured in Chitwan National Park in the early 1970s had an average weight of 235 kg (518 lb) ranging from 200 to 261 kg (441 to 575 lb), and that of the females was 140 kg (310 lb) ranging from 116 to 164 kg (256 to 362 lb).[27] Two male tigers captured in Chitwan National Park in the 1980s exceeded weights of 270 kg (600 lb) and are the largest free ranging tigers reported to date.[28]
The smallest recorded weights for Bengal tigers are from the Bangladesh Sundarbans, where adult females weigh 75–80 kg (165–176 lb).[29]
Three tigresses from the Bangladesh Sundarbans had a mean weight of 76.7 kg (169 lb). The oldest female weighed 75 kg (165 lb) and was in a relatively poor condition at the time of capture. Their skulls and body weights were distinct from those of tigers in other habitats, indicating that they may have adapted to the unique conditions of the mangrove habitat. Their small sizes are probably due to a combination of intense intraspecific competition and small size of prey available to tigers in the Sundarbans, compared to the larger deer and other prey available to tigers in other parts.
In the beginning of the 20th century, a male tiger was shot in central India with a head and body length of 221 cm (87 in) between pegs, a chest girth of 150 cm (59 in), a shoulder height of 109 cm (43 in) and a tail length of 81 cm (32 in), which was perhaps bitten off by a rival male. This specimen could not be weighed, but it was estimated to weigh about 272 kg (600 lb).[26] A male weighing 259 kg (570 lb) was shot in northern India in the 1930s.[32] A male tiger shot in Nepal weighed 320 kg (710 lb) and measured 328 cm (10 ft 9 in) 'over the curves'.[33] The heaviest wild tiger was possibly a huge male killed in 1967 at the foothills of the Himalayas. It weighed 388.7 kg (857 lb) after eating a buffalo calf; it measured 323 cm (127 in) in total length between pegs, and 338 cm (133 in) over curves. Without eating the calf beforehand, it would have likely weighed at least 324.3 kg (715 lb). This specimen is on exhibition in the Mammals Hall of the Smithsonian Institution.[34] In the Central Provinces of India, a male tiger shot weighed 317 kg (699 lb) and measured 3.02 m (9 ft 11 in).[35]
The Bengal tiger rivals the Siberian tiger in average weight.[36]
Distribution and habitat
In 1982, a sub-
In Pakistan, Khairpur was the last stronghold of the tiger by the late 19th century; the last individuals were shot in 1906 in Bahawalpur in the Indus Riverine jungles.[40]
India
In the 20th century, Indian censuses of wild tigers relied on the individual identification of footprints known as pug marks – a method that has been criticised as deficient and inaccurate. Camera traps are now being used in many sites.[41]
Good tiger habitats in subtropical and
During a tiger census in 2008, camera trap and sign surveys using
- in the Sohagi Barwa Sanctuary and in Valmiki National Park;
- in the ;
- in the Giddalur;
- in the Kudremukh-Dandeli;
- in the Brahmaputra flood plains and northeastern hills tigers live in an area of 4,230 km2 (1,630 sq mi) in several patchy and fragmented forests;
- in the Sundarbans National Park tigers live in about 1,586 km2 (612 sq mi) of mangrove forest.
Manas-Namdapha, Orang-Laokhowa and Kaziranga-Meghalaya are Tiger Conservation Units in northeastern India, stretching over at least 14,500 km2 (5,600 sq mi) across several protected areas.[39] Tigers are also present in Pakke Tiger Reserve.[43] In the Mishmi Hills, tigers were recorded in 2017 up to an elevation of 3,630 m (11,910 ft) in snow.[44] Ranthambore National Park hosts India's westernmost tiger population.
As of 2014, the Indian tiger population was estimated to range over an area of 89,164 km2 (34,426 sq mi) and number 2,226 adult and subadult tigers older than one year. About 585 tigers were present in the Western Ghats, where
In May 2018, a tiger was recorded in Sahyadri Tiger Reserve for the first time in eight years.[47] In February 2019, a tiger was sighted in Gujarat's Lunavada area in Mahisagar district, and found dead shortly afterwards.[48] Officials assumed that it originated in Ratapani Tiger Reserve and travelled about 300 km (190 mi) over two years. It probably died of starvation. In May 2019, camera traps recorded tigers in Mhadei Wildlife Sanctuary and Bhagwan Mahaveer Sanctuary and Mollem National Park, the first records in Goa since 2013.[49][50]
The tigers in the Sundarbans in India and Bangladesh are the only ones in the world inhabiting mangrove forests.[45] The population in the Indian Sundarbans was estimated as 86–90 individuals in 2018.[9]
Bangladesh
In Bangladesh, tigers are now relegated to the forests of the Sundarbans and the Chittagong Hill Tracts.[51] The Chittagong forest is contiguous with tiger habitat in India and Myanmar, but the tiger population is of unknown status.[52]
As of 2004, population estimates in Bangladesh ranged from 200 to 419 individuals, most of them in the Sundarbans.
Since 2007, tiger monitoring surveys have been carried out every year by WildTeam in the Bangladesh Sundarbans to monitor changes in the Bangladesh tiger population and assess the effectiveness of conservation actions. This survey measures changes in the frequency of tiger track sets along the sides of tidal waterways as an index of relative tiger abundance across the Sundarbans landscape.[56] By 2009, the tiger population in the Bangladesh Sundarbans was estimated as 100–150 adult females or 335–500 tigers overall. Female home ranges, recorded using Global Positioning System collars, were some of the smallest recorded for tigers, indicating that the Bangladesh Sundarbans could have one of the highest densities and largest populations of tigers anywhere in the world. They are isolated from the next tiger population by a distance of up to 300 km (190 mi). Information is lacking on many aspects of Sundarbans tiger ecology, including relative abundance, population status, spatial dynamics, habitat selection, life history characteristics, taxonomy, genetics, and disease. There is also no monitoring program in place to track changes in the tiger population over time, and therefore no way of measuring the response of the population to conservation activities or threats. Most studies have focused on the tiger-human conflict in the area, but two studies in the Sundarbans East Wildlife sanctuary documented habitat-use patterns of tigers, and abundances of tiger prey, and another study investigated tiger parasite load. Some major threats to tigers have been identified. The tigers living in the Sundarbans are threatened by habitat destruction, prey depletion, highly aggressive and rampant intraspecific competition, tiger-human conflict, and direct tiger loss.[30]
By 2017, this population was estimated at 84–158 individuals.[57] A rising sea-level due to climate change is projected to cause a severe loss of suitable habitat for this population in the following decades, around 50% by 2050 and 100% by 2070.[58]
Nepal
The tiger population in the Terai of Nepal is split into three isolated subpopulations that are separated by cultivation and densely settled habitat. The largest population lives in
From February to June 2013, a camera trapping survey was carried out in the Terai Arc Landscape, across an area of 4,841 km2 (1,869 sq mi) in 14 districts. The country's tiger population was estimated at 163–235 breeding adults comprising 102–152 tigers in the Chitwan-Parsa protected areas, 48–62 in Bardia-Banke National Parks and 13–21 in Shuklaphanta National Park.[60] Between November 2017 and April 2018, the third nationwide survey for tiger and prey was conducted in the Terai Arc Landscape; the country's population was estimated at 220–274 tigers.[61]
Bhutan
In Bhutan, tigers have been documented in 17 of 18
Royal Manas and Jigme Singye Wangchuck National Parks form the largest contiguous tiger conservation area in Bhutan representing subtropical to alpine habitat types.[64] In 2010, camera traps recorded a tiger pair at elevations of 3,000 to 4,100 m (9,800 to 13,500 ft). As of 2015, the tiger population in Bhutan was estimated at 89 to 124 individuals in a survey area of 28,225 km2 (10,898 sq mi).[65]
In 2008, a tiger was recorded at an elevation of 4,200 m (13,800 ft) in Jigme Dorji National Park, which is the highest elevation record of a tiger known to date.[66] In 2017, a tiger was recorded for the time in Bumdeling Wildlife Sanctuary. It probably used a wildlife corridor to reach northeastern Bhutan.[67]
Bhutan's tiger population was estimated at 90 individuals comprising 60 females and 30 males with a population density estimate of 0.19–0.31 tigers per 100 km2 (39 sq mi) by March 2015.[11]
China
The presence of the Bengal tiger in southeastern Tibet Autonomous Region, China was investigated in 1995 when the loss of livestock was high in Mêdog County due to a large predator. Tiger paw prints were found on pastures around several villages.[68] One tiger was shot in 1996, and about 4–5 tigers were reported by officials in the area by 1999.[69] About 8–12 tigers were thought to remain in this area a decade later.[70] A camera trapping and interview survey during 2013–2018 in nine potential sites in Mêdog County revealed that only 1–3 non-resident individuals might be entering the area south of the Yarlung Tsangpo river, but only during the dry season from October to March.[71] In early 2019, a Bengal tiger was photographed twice at an elevation of 2,000 m (6,600 ft) in a broadleaved forest in Yarlung Tsangpo Grand Canyon National Nature Reserve.[72]
Ecology and behaviour
The basic social unit of the tiger is composed of a female and her offspring. Adult animals congregate only temporarily when special conditions permit, such as plentiful supplies of food. Otherwise, they lead solitary lives, hunting individually for the forest and grassland animals upon which they prey. Resident adults of either sex maintain home ranges, confining their movements to definite habitats within which they satisfy their needs and those of their cubs, which include prey, water and shelter. In this site, they also maintain contact with other tigers, especially those of the opposite sex. Those sharing the same ground are well aware of each other's movements and activities.
In the Panna Tiger Reserve, an adult radio-collared male tiger moved 1.7 to 10.5 km (1.1 to 6.5 mi) between locations on successive days in winter, and 1 to 13.9 km (0.62 to 8.64 mi) in summer. His home range was about 200 km2 (77 sq mi) in summer and 110 km2 (42 sq mi) in winter. Included in his home range were the much smaller home ranges of two females, a tigress with cubs and a subadult tigress. They occupied home ranges of 16 to 31 km2 (6.2 to 12.0 sq mi).[74]
The home ranges occupied by adult male residents tend to be mutually exclusive, even though one of these residents may tolerate a transient or sub-adult male at least for a time. A male tiger keeps a large territory in order to include the home ranges of several females within its bounds, so that he may maintain mating rights with them. Spacing among females is less complete. Typically there is partial overlap with neighbouring female residents. They tend to have core areas, which are more exclusive, at least for most of the time. Home ranges of both males and females are not stable. The shift or alteration of a home range by one animal is correlated with a shift of another. Shifts from less suitable habitat to better ones are made by animals that are already resident. New animals become residents only as vacancies occur when a former resident moves out or dies. There are more places for resident females than for resident males.[18]
During seven years of camera trapping, tracking, and observational data in Chitwan National Park, six to nine breeding tigers, two to sixteen non-breeding tigers, and six to twenty young tigers of less than one year of age were detected in the study area of 100 km2 (39 sq mi). One of the resident females left her territory to one of her female offspring and took over an adjoining area by displacing another female; and a displaced female managed to re-establish herself in a neighbouring territory made vacant by the death of the resident. Of 11 resident females, 7 were still alive at the end of the study period, two disappeared after losing their territories to rivals, and two died. The initial loss of two resident males and subsequent take over of their home ranges by new males caused social instability for two years. Of four resident males, one was still alive and three were displaced by rivals. Five litters of cubs were killed by infanticide, two litters died because they were too young to fend for themselves when their mothers died. One juvenile tiger was presumed dead after being photographed with severe injuries from a deer
Hunting and diet
The tiger is a
Bengal tigers occasionally hunt and kill predators such as
Results of scat analyses indicate that the tigers in Nagarahole National Park preferred prey weighing more than 176 kg (388 lb) and that on average tiger prey weighed 91.5 kg (202 lb). The prey species included chital, sambar, wild pig and gaur. Gaur remains were found in 44.8% of all tiger scat samples, sambar remains in 28.6%, wild pig remains in 14.3% and chital remains in 10.4% of all scat samples.[85] In Bandipur National Park, gaur and sambar together also constituted 73% of tiger diet.[77]
In most cases, tigers approach their victim from the side or behind from as close a distance as possible and grasp the prey's throat to kill it. Then they drag the carcass into cover, occasionally over several hundred metres, to consume it. The nature of the tiger's hunting method and prey availability results in a "feast or famine" feeding style: they often consume 18–40 kg (40–88 lb) of meat at one time.[2] In one study, tigresses from Nepal made about 40–50 kills a year and ate a minimum of 5–6.3 kg (11–14 lb) of meat a day. Two males made about 40–50 kills in a year and ate 6.3–7.8 kg (14–17 lb) of meat a day at the least.[86]
If injured, old or weak, or when its regular prey species become scarce, tigers often attack humans and become man-eaters.[87]
Reproduction and lifecycle
The tiger in India has no definite mating and birth seasons. Most young are born in December and April.
Males reach maturity at 4–5 years of age, and females at 3–4 years. A Bengal comes into heat at intervals of about 3–9 weeks, and is receptive for 3–6 days. After a gestation period of 104–106 days, 1–4 cubs are born in a shelter situated in tall grass, thick bush or in caves. Newborn cubs weigh 780 to 1,600 g (1.72 to 3.53 lb) and they have a thick woolly fur that is shed after 3.5–5 months. Their eyes and ears are closed. Their milk teeth start to erupt at about 2–3 weeks after birth, and are slowly replaced by permanent dentition from 8.5 to 9.5 weeks of age onwards. They suckle for 3–6 months, and begin to eat small amounts of solid food at about 2 months of age. At this time, they follow their mother on her hunting expeditions and begin to take part in hunting at 5–6 months of age. At the age of 2–3 years, they slowly start to separate from the family group and become transient, looking out for an area, where they can establish their own home range. Young males move farther away from their native home range than young females. Once the family group has split, the mother comes into heat again.[2]
Threats
None of the Tiger Conservation Landscapes within the Bengal tiger range is large enough to support an effective population size of 250 individuals. Habitat losses and the extremely large-scale incidences of poaching are serious threats to the species' survival.[8]
The
Poaching
The most significant immediate threat to the existence of wild tiger populations is the
The illicit demand for bones and body parts from wild tigers for use in Traditional Chinese medicine is the reason for the unrelenting poaching pressure on tigers on the Indian subcontinent. For at least a thousand years, tiger bones have been an ingredient in traditional medicines that are prescribed as a muscle strengthener and treatment for rheumatism and body pain.[94]
Between 1994 and 2009, the Wildlife Protection Society of India has documented 893 cases of tigers killed in India, which is just a fraction of the actual poaching and trade in tiger parts during those years.[95] In 2004, all the tigers in India's
Human–tiger conflict
The Indian subcontinent has served as a stage for intense confrontations between tigers and people. At the beginning of the 19th century tigers were so numerous, that the killing of tigers was officially rewarded in many localities. The Terai region supported large numbers of tigers that were pushed into marginal habitat after the 1950s, when the conversion of natural habitat for paddy fields increased. Marauding tigers began to take a toll of human life in areas bordering cultivation. They are thought to have followed domestic livestock that wintered in the plains when they returned to the hills in the spring, and then being left without prey when the herds dispersed back to their respective villages. These tigers were the old, the young and the disabled. All suffered from some disability, mainly caused either by gunshot wounds or porcupine quills.[101]
In the Sundarbans, 10 out of 13-man-eaters recorded in the 1970s were males, and they accounted for 86% of the victims. These man-eaters have been grouped into the confirmed or dedicated ones who go hunting especially for human prey; and the opportunistic ones, who do not search for humans but will, if they encounter a man, attack, kill and devour him. In areas where opportunistic man-eaters were found, the killing of humans was correlated with their availability, most victims being claimed during the honey-gathering season.[102] Tigers in the Sunderbans presumably attacked humans who entered their territories in search of wood, honey or fish, thus causing them to defend their territories. The number of tiger attacks on humans may be higher outside suitable areas for tigers, where numerous people are present, but with little wild prey for tigers.[103]
In Nepal, the incidence of man-eating tigers has been only sporadic. In Chitwan National Park no cases were recorded before 1980. In the following few years, 13 people have been killed and eaten in the park and its environs. In the majority of cases, man-eating appeared to have been related to an intra-specific competition among male tigers.[101] An interview survey with 499 local people in Chitwan revealed that lower caste Hindus and people with less than eight years of formal education had negative attitudes to the tiger; most of them owned livestock and had heard about tigers attacking people and livestock.[104]
In December 2012, a tiger was shot by the
Conservation efforts
An area of special interest lies in the "Terai Arc Landscape" in the
The WWF partnered with Leonardo DiCaprio to form a global campaign, "Save Tigers Now", with the ambitious goal of building political, financial and public support to double the wild tiger population by 2022.[108] Save Tigers Now started its campaign in 12 different WWF Tiger priority landscapes, since May 2010.[109]
This population of tigers has been assessed at the local level in several countries. It is listed as
In April 2023, India signed a memorandum of understanding with Cambodia to assist the country with the tiger's reintroduction. At least 90 acres (36 ha) of the Cardamom Mountains of Tatai Wildlife Sanctuary could be used to host tigers that are imported from India.[115] The last tiger in Cambodia was photographed in 2007 by a camera trap. In 2016, the Cambodian government declared that the Indochinese tiger population was "functionally extinct".[116][117]
In India
In 1973,
Through this initiative the population decline was reversed initially, but has resumed in recent years; India's tiger population decreased from 3,642 in the 1990s to just over 1,400 from 2002 to 2008.[120]
The Indian
Following the revelation that only 1,411 Bengal tigers existed in the wild in India, down from 3,600 in 2003, the Indian government set up eight new
In January 2008, the Government of India launched a dedicated anti-poaching force composed of experts from Indian police, forest officials and various other environmental agencies.[126] Ranthambore National Park is often cited as a major success by Indian officials against poaching.[127]
Kuno-Palpur in Madhya Pradesh was supposed receive Asiatic lions from Gujarat. Since no lion has been transferred from Gujarat to Madhya Pradesh so far, it may be used as a sanctuary for the tiger instead.[128][129]
In captivity
Bengal tigers have been captive bred since 1880 and widely crossed with tigers from other range countries.[130]
In July 1976,
Indian zoos bred tigers for the first time at the
In Bangladesh
WildTeam is working with local communities and the Bangladesh Forest Department to reduce human-tiger conflict in the Bangladesh Sundarbans. For over 100 years people, tigers, and livestock have been injured and killed in the conflict; in recent decades up to 50 people, 80 livestock, and 3 tigers have been killed in a year. Now, through WildTeam's work, there is a boat-based Tiger Response team that provides first aid, transport, and body retrieval support for people being killed in the forest by tigers. WildTeam has also set up 49 volunteer Village Response Teams that are trained to save tigers that have strayed into the village areas and would be otherwise killed. These village teams are made up of over 350 volunteers, who are also now supporting anti-poaching work and conservation education/awareness activities. WildTeam also works to empower local communities to access the government funds for compensating the loss/injury of livestock and people from the conflict. To monitor the conflict and assess the effectiveness of actions, WildTeam have also set up a human-tiger conflict data collection and reporting system.
In Nepal
In May 2010, Banke National Park was established with an area of 550 km2 (210 sq mi).[136] The government aimed at doubling the country's tiger population by 2022 at the Global Tiger Summit in 2010. The tiger population reached 355 in 2022, almost tripling the population of 121 in 2009.[137][138]
"Re-wilding" project in South Africa
In 2000, the Bengal tiger re-wilding project Tiger Canyons was started by John Varty, who together with the zoologist Dave Salmoni trained captive-bred tiger cubs how to stalk, hunt, associate hunting with food and regain their predatory instincts. They claimed that once the tigers proved that they can sustain themselves in the wild, they would be released into a free-range sanctuary of South Africa to fend for themselves.[139]
The project has received controversy after accusations by their investors and conservationists of manipulating the behaviour of the tigers for the purpose of a film production, Living with Tigers, with the tigers believed to be unable to hunt.[140][141] Stuart Bray, who had originally invested a large sum of money in the project, claimed that he and his wife, Li Quan, watched the film crew "[chase] the prey up against the fence and into the path of the tigers just for the sake of dramatic footage."[140][141]
The four tigers involved in this project have been confirmed to be crossbred Siberian–Bengal tigers, which should neither be used for breeding nor being released into the Karoo. Tigers that are not genetically pure will not be able to participate in the tiger Species Survival Plan, as they are not used for breeding, and are not allowed to be released into the wild.[142]
In culture
The tiger is one of the animals displayed on the
Today, the tiger is the national animal of Bangladesh and India. Bangladeshi banknotes feature a tiger. The political party Muslim League of Pakistan uses the tiger as its election symbol.[146] Tipu Sultan, who ruled Mysore in late 18th-century India, was also a great admirer of the animal. The famed 18th-century automaton, Tipu's Tiger was also created for him.[147] The tiger was the dynastic symbol of this dynasty.[148] During the Indian Rebellion of 1857, Punch ran a political cartoon showing the Indian rebels as a tiger attacking a victim.[149]
In arts
- The main antagonist of The Jungle Book, Shere Khan, is a Bengal tiger.[150]
- The Kumaon Division.[151]
- In the fantasy adventure novel Life of Pi and in its 2012 film adaptation, a Bengal tiger named Richard Parker is the lead character.[152]
- The Bengal Tiger at the Baghdad Zoo is based on a real story of a tiger that escaped from Baghdad Zoo in 2003 and haunts the streets of Baghdad seeking the meaning of life.[153][154][155]
- The BBC on tigers in Bhutan.[156]
- The 2014 Roar – Tigers of the Sundarbans is about a white Bengal tigress in the Sundarbans.[157]
Notable individuals
Notable Bengal tigers include the
See also
- Prehistoric tigers: Panthera tigris soloensis
- Panthera tigris trinilensis
- Panthera tigris acutidens
- Holocene extinction
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Further reading
- Schnitzler, A.; Hermann, L. (2019). "Chronological distribution of the tiger Panthera tigris and the Asiatic lion Panthera leo persica in their common range in Asia". S2CID 202040786.
External links
- "Panthera t. tigris". Cat Specialist Group.
- "Tiger conservation in the Bangladesh Sundarbans". WildTeam. Archived from the original on 6 October 2020. Retrieved 26 May 2021.
- "Bengal Tiger". Panthera. Archived from the original on 24 September 2015. Retrieved 24 October 2011.
- Li, Y. & H. Liqiang (2019). "Bengal tigers found in Tibet, with plenty of prey". China Daily.
- "The four faces of the Bengal tiger". Guardian News and Media Limited.
- "Himalayan black bear killed by tiger". Big Cat Rescue. 12 November 2009.