Blueberry

Source: Wikipedia, the free encyclopedia.
This article is about the North American blueberry. For the Eurasian blueberry, see Bilberry. For other uses, see Blueberry (disambiguation).

Blueberry
Vaccinium caesariense
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Ericales
Family: Ericaceae
Genus: Vaccinium
Section: Vaccinium sect. Cyanococcus
Rydb.
Species

See text

Blueberries showing various stages of maturation. IG = Immature Green, GP = Green Pink, BP = Blue Pink, and R = Ripe.

Blueberry is a widely distributed and widespread group of

huckleberries and Madeira blueberries.[1] Commercial blueberries—both wild (lowbush) and cultivated (highbush)—are all native to North America. The highbush varieties were introduced into Europe during the 1930s.[2]

Blueberries are usually prostrate shrubs that can vary in size from 10 centimeters (4 inches) to 4 meters (13 feet) in height. In commercial production of blueberries, the species with small, pea-size berries growing on low-level bushes are known as "lowbush blueberries" (synonymous with "wild"), while the species with larger berries growing on taller, cultivated bushes are known as "highbush blueberries". Canada is the leading producer of lowbush blueberries, while the United States produces some 40% of the world supply of highbush blueberries.

Origin and history of cultivation

Flowers on a cultivated blueberry bush
Fresh blueberries

The genus Vaccinium has a mostly circumpolar distribution, with species mainly present in North America, Europe, and Asia.[1] Many commercially available species with English common names including "blueberry" are from North America,[3] particularly Atlantic Canada and the northeastern United States for wild (lowbush) blueberries, and several US states and British Columbia for cultivated (highbush) blueberries.[4][5] First Nations peoples of Canada consumed wild blueberries for millennia.[4] Highbush blueberries were first cultivated in New Jersey around the beginning of the 20th century.[5][3]

North American native species of blueberries are grown commercially in the Southern Hemisphere in Australia, New Zealand and South American nations. The Colombian or Andean blueberry,

bilberries
, which in many languages have a name that translates to "blueberry" in English.

Description

Five species of blueberries grow wild in Canada, including Vaccinium myrtilloides, Vaccinium angustifolium and Vaccinium corymbosum, which grow on forest floors or near swamps.[7] Wild (lowbush) blueberries are not planted by farmers, but rather are managed on berry fields called "barrens".[4]

Wild blueberries reproduce by

cross pollination, with each seed producing a plant with a different genetic composition, causing within the same species differences in growth, productivity, color, leaf characteristics, disease resistance, flavor, and other fruit characteristics.[7] The mother plant develops underground stems called rhizomes, allowing the plant to form a network of rhizomes creating a large patch (called a clone) which is genetically distinct.[7] Floral and leaf buds develop intermittently along the stems of the plant, with each floral bud giving rise to 5–6 flowers and the eventual fruit.[7] Wild blueberries prefer an acidic soil between 4.2 and 5.2 pH and only moderate amounts of moisture.[7] They have a hardy cold tolerance in their Canadian range and the U.S. state of Maine.[7] Fruit productivity of lowbush blueberries varies by the degree of pollination, genetics of the clone, soil fertility, water availability, insect infestation, plant diseases and local growing conditions.[7] Wild (lowbush) blueberries have an average mature weight of 0.3 grams (1⁄128 oz).[7]

Highbush (cultivated) blueberries prefer sandy or

lanceolate, and 1–8 cm (1⁄2–3+1⁄4 in) long and 0.5–3.5 cm (1⁄4–1+3⁄8 in) broad. The flowers
are bell-shaped, white, pale pink or red, sometimes tinged greenish.

The fruit is a berry 5–16 mm (3⁄16–5⁄8 in) in diameter with a flared crown at the end; they are pale greenish at first, then reddish-purple, and finally uniformly blue when ripe.[8] They are covered in a protective coating of powdery epicuticular wax, colloquially known as the "bloom".[7] They generally have a sweet taste when mature, with variable acidity.[7][8] Blueberry bushes typically bear fruit in the middle of the growing season: fruiting times are affected by local conditions such as climate, altitude and latitude, so the time of harvest in the northern hemisphere can vary from May to August.[7][8]

Species

Note: habitat and range summaries are from the Flora of New Brunswick, published in 1986 by Harold R. Hinds, and Plants of the Pacific Northwest coast, published in 1994 by Pojar and MacKinnon.

Some other blue-fruited species of Vaccinium:

  • Wild blueberry in autumn foliage, Pilot Mountain, North Carolina, in October
    Wild blueberry in autumn foliage, Pilot Mountain, North Carolina, in October
  • A maturing 'Polaris' blueberry (Vaccinium corymbosum)
    A maturing 'Polaris' blueberry (Vaccinium corymbosum)
  • A selection of blueberries, showing the typical sizes of the berries. The scale is marked in centimeters.
    A selection of blueberries, showing the typical sizes of the berries. The scale is marked in centimeters.

The lowbush varieties are V. angustifolium, V. boreale, V. mytilloides, V. pallidum, and V. angustifolium × V. corymbosum. They are still grown in a similar manner to pre-Columbian semi-wild cultivation, i.e.

slash and burn. The highbush varieties are darrowii and corymbosum. Rabbiteye (V. ashei/V. virgatum) is considered different from both high- and lowbush.[3]

Identification

Commercially offered blueberries are usually from species that naturally occur only in eastern and north-central North America. Other sections in the genus are native to other parts of the world, including the Pacific Northwest and southern United States,[10] South America, Europe and Asia. Other wild shrubs in many of these regions produce similar-looking edible berries, such as huckleberries and whortleberries (North America) and bilberries (Europe). These species are sometimes called "blueberries" and are sold as blueberry jam or other products.

The names of blueberries in languages other than English often translate as "blueberry", e.g.

Scots blaeberry and Norwegian blåbær. Blaeberry, blåbær and French myrtilles usually refer to the European native bilberry (V. myrtillus), while bleuets refers to the North American blueberry. Russian голубика ("blue berry") does not refer to blueberries, which are non-native and nearly unknown in Russia, but rather to their close relatives, bog bilberries
(V. uliginosum).

Cyanococcus blueberries can be distinguished from the nearly identical-looking bilberries by their flesh color when cut in half. Ripe blueberries have light green flesh, while bilberries, whortleberries and huckleberries are red or purple throughout.

Blueberries, raw
A dish of blueberries
Nutritional value per 100 g (3.5 oz)
Energy240 kJ (57 kcal)
14.49 g
Sugars9.96 g
Dietary fiber2.4 g
0.33 g
0.74 g
Niacin (B3)
3%
0.418 mg
Pantothenic acid (B5)
2%
0.124 mg
Vitamin B6
3%
0.052 mg
Folate (B9)
2%
6 μg
Vitamin C
11%
9.7 mg
Vitamin E
4%
0.57 mg
Vitamin K
16%
19.3 μg
MineralsQuantity
%DV†
Calcium
0%
6 mg
Iron
2%
0.28 mg
Magnesium
1%
6 mg
Manganese
15%
0.336 mg
Phosphorus
1%
12 mg
Potassium
3%
77 mg
Sodium
0%
1 mg
Zinc
2%
0.165 mg
Other constituentsQuantity
Water84 g
Link to USDA Database entry
†Percentages estimated using US recommendations for adults,[11] except for potassium, which is estimated based on expert recommendation from the National Academies.[12]

Culinary use

Blueberries are sold fresh or are processed as

breakfast cereals
.

Blueberry jam is made from blueberries, sugar, water, and fruit pectin. Blueberry sauce is a sweet sauce prepared using blueberries as a primary ingredient.

Blueberry wine is made from the flesh and skin of the berries, which is fermented and then matured; usually the lowbush variety is used.

Nutrients

Blueberries consist of 14%

dietary mineral manganese, vitamin C, vitamin K and dietary fiber (table). Generally, nutrient contents of blueberries are a low percentage of the DV (table). A 100-gram serving provides a relatively low amount of food energy – 240 kilojoules (57 kcal) – with a glycemic load
of 6.

Phytochemicals and research

Blueberries contain anthocyanins, other polyphenols and various phytochemicals under preliminary research for their potential biological effects.[13] Most polyphenol studies have been conducted using the highbush cultivar of blueberries (V. corymbosum), while content of polyphenols and anthocyanins in lowbush (wild) blueberries (V. angustifolium) exceeds values found in highbush cultivars.[14]

Cultivation

Blueberry harvester in Michigan

Blueberries may be cultivated, or they may be picked from semiwild or wild bushes. In North America, the most common cultivated species is V. corymbosum, the

northern highbush blueberry. Hybrids of this with other Vaccinium species adapted to southern U.S. climates are known collectively as southern highbush blueberries.[15]

So-called "wild" (lowbush) blueberries, smaller than cultivated highbush ones, have intense color. The

Atlantic provinces westward to Quebec and southward to Michigan and West Virginia. In some areas, it produces natural "blueberry barrens", where it is the dominant species covering large areas. Several First Nations communities in Ontario
are involved in harvesting wild blueberries.

"Wild" has been adopted as a marketing term for harvests of managed native stands of lowbush blueberries. The bushes are not planted or selectively bred, but they are pruned or burned over every two years, and pests are "managed".[16]

Numerous highbush

cranberry bogs at Whitesbog in the New Jersey Pine Barrens. His work doubled the size of some strains' fruit, and by 1916, he had succeeded in cultivating blueberries, making them a valuable crop in the Northeastern United States.[18][20] For this work he received the George Roberts White Medal of Honor from the Massachusetts Horticultural Society
.

The

the Carolinas to the Gulf Coast states. Production of rabbiteye blueberries was a focus in Texas in the early 21st century.[21] Other important species in North America include V. pallidum, the hillside or dryland blueberry. It is native to the eastern U.S., and common in the Appalachians and the Piedmont
of the Southeast. Sparkleberry, V. arboreum, is a common wild species on sandy soils in the Southeast.

Successful blueberry cultivation requires attention to

acidity) measurements in the acidic range.[22][23][24]

Blueberry bushes often require supplemental fertilization,[23] but over-fertilization with nitrogen can damage plant health, as evidenced by nitrogen-burn visible on the leaves.[22][23]

Growing regions

Worldwide highbush blueberry growing areas (data from 2008)

Significant production of highbush blueberries occurs in

Washington. The production of southern highbush varieties occurs in California, as varieties originating from University of Florida, Connecticut, New Hampshire, North Carolina State University and Maine have been introduced. Peru, Spain, and Mexico also have significant production, as of 2018 (see Production).[25]

United States

In 2018,

Georgia, Michigan, New Jersey, California, and North Carolina.[27]

Hammonton, New Jersey, claims to be the "Blueberry Capital of the World",[28] with over 80% of New Jersey's cultivated blueberries coming from this town.[29] Every year the town hosts a large festival, which draws thousands of people to celebrate the fruit.[30]

Maine is known for its wild blueberries,[31] but the state's lowbush (wild) and highbush blueberries combined account for 10% of all blueberries grown in North America. Some 44,000 hectares (110,000 acres) are farmed, but only half of this acreage is harvested each year due to variations in pruning practices.[32] The wild blueberry is the official fruit of Maine.[33]

Canada

Wild blueberry fields in Nova Scotia, Canada

Canadian production of wild and cultivated blueberries in 2015 was 166,000 tonnes valued at $262 million, the largest fruit crop produced nationally accounting for 29% of all fruit value.[34]

British Columbia was the largest Canadian producer of cultivated blueberries, yielding 70,000 tonnes in 2015,[34] the world's largest production of blueberries by region.[35]

Québec are also major producers.[37] Nova Scotia recognizes the wild blueberry as its official provincial berry,[38] with the town of Oxford, Nova Scotia known as the Wild Blueberry Capital of Canada.[39]

Québec is a major producer of wild blueberries, especially in the regions of

Saguenay-Lac-Saint-Jean (where a popular name for inhabitants of the regions is bleuets, or "blueberries") and Côte-Nord, which together provide 40% of Québec's total provincial production. This wild blueberry commerce benefits from vertical integration of growing, processing, frozen storage, marketing and transportation within relatively small regions of the province.[40] On average, 80% of Québec wild blueberries are harvested on farms (21 million kilograms (23,000 short tons)), the remaining 20% being harvested from public forests (5 million kilograms (5,500 short tons)).[40] Some 95% of the wild blueberry crop in Québec is frozen for export out of the province.[40]

Vaccinium meridionale, a wild species found in the Andes[citation needed]

Europe

Highbush blueberries were first introduced to Germany, Sweden and the Netherlands in the 1930s, and have since been spread to numerous other countries of Europe.[2] V. corymbosum only began to be cultivated in Romania in a few years leading up to 2018 and rapidly increased in production and sales in that time (as with berries in general). As of 2018 it remains relatively unmolested by pests and diseases (see Diseases below).[41]

Southern Hemisphere

In the Southern Hemisphere, Brazil, Chile, Argentina, Peru, Uruguay, New Zealand, Australia, South Africa,[42] and Zimbabwe grow blueberries commercially.[43]

In Brazil, blueberries are produced in the states of Rio Grande do Sul, Santa Catarina, Paraná, São Paulo and Minas Gerais.[44]

Blueberries were first introduced to Australia in the 1950s, but the effort was unsuccessful. In the early 1970s, the Victorian Department of Agriculture imported seed from the U.S. and a selection trial was started. This work was continued into the mid-1970s when the Australian Blueberry Growers' Association was formed.[45]

In the 21st century, the industry grew in

forced labor, blueberries were listed among the goods produced in such working conditions in Argentina.[48]

Pests and diseases

Diseases

Main article: List of Vaccinium diseases

As of 2018 V. corymbosum remains relatively unmolested by pests and diseases in Romania, with

Microsphaera penicillata var. vaccinii, and various viruses being the most common.[41]

Pest management

Pesticides

DDT began to be used in blueberry soon after its discovery in 1939, and a few years later in the mid-1940s research began into its use in North America.[3]

Because "wild" is a marketing term generally used for all low-bush blueberries, it is not an indication that such blueberries are free from pesticides.[49]

Insecticide modes of action must be varied to avoid encouraging resistance in the invasive pest Drosophila suzukii.[3]

Some

Kaolin clay for Rhagoletis mendax also reduced effectiveness of Diachasma alloeum, its parasitoid. The pest predator Harpalus erraticus maintains greater abundance with selective insecticides rather than broad-spectrum MoAs.[3]

Integrated pest management

Blueberries are naturally relatively unmolested by

cicadellids (vectoring the phytoplasma that causes blueberry stunt). Managing pests down to the cosmetic level is necessary in this fruit because they are a premium type product.[3]

Changes in locale and environment – to new geographies, and into

Operophtera brumata is a pest of blueberries and birches which is successfully parasitized by Cyzenis albicans despite the lack of historical, natural contact between the two. The same results were obtained with Scirtothrips citri and Beauveria bassiana. Results are available for Choristoneura rosaceana and overwhelming numbers of Trichogramma minutum, and Cyclocephala longula overwhelmed by Steinernema scarabaei. This has also been attempted with flower thrips and potential predators but with inconclusive results.[3]

International quarantine

Rhagoletis mendax is a

phytosanitary regimes of some countries around the world.[3]

Resistant cultivars

Insect resistance was not a priority in

resistant than V. corymbosum to Scaphytopius magdalensis. V. ashei is less resistant than V. darrowii to Prodiplosis vaccinia. There is variation between cultivars of V. ashei in resistance to Oberea myops. There is variation in resistance among cultivars of V. corymbosum to Acrobasis vaccinii and Popillia japonica. Wild V. spp. have greater resistance than highbush cultivars to I. pepperi. There is significant variation between highbush cultivars in abundance of various Tephritidae, thrips, and Homalodisca vitripennis.[3]

Production

Blueberry production – 2021
Country Production
(tonnes)
 United States 351,130
 Peru 227,971
 Canada 146,551
 Chile 122,795
 Mexico 66,482
 Spain 61,230
World 1,113,261
Source:
FAOSTAT of the United Nations[50]

In 2021, world production of blueberries (lowbush and highbush combined) was 1.1 million tonnes, led by the United States with 32% of global production, Peru with 20%, and Canada with 13% (table).[50]

In 2019, Canada was the largest producer of wild blueberries, mainly in

Atlantic provinces,[51] but Canadian production of wild blueberries decreased since 2017 by transitioning to the more profitable cultivated highbush blueberries.[52] British Columbia produced 93% of the Canadian highbush blueberry crop in 2019.[51]

Regulations

Canada

Canada No. 1 blueberries are all similar in size, shape, weight, and color—the total product can be no more than ten percent off-color and three percent otherwise defective.[53]

See also

Wikimedia Commons has media related to Blueberries.

References

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Further reading

External links
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