Bradysaurus

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Bradysaurus
Temporal range:
Ma
Skeleton in the
Field Museum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Parareptilia
Order: Procolophonomorpha
Clade: Pareiasauria
Family:
Pareiasauridae
Subfamily: Bradysaurinae
von Huene, 1948
Genus: Bradysaurus
Watson
, 1914
Type species
Bradysaurus baini
Seeley, 1892
Species
  • B. baini Seeley, 1892
  • B. seeleyi Haughton and Boonstra, 1929
Synonyms
  • Bradysaurus broomi Haughton and Boonstra, 1929
  • Bradysaurus vanderbyli Haughton and Boonstra, 1929
  • Pareiasaurus baini Pareiasaurus baini Seeley, 1892
  • Platyoropha broom Haughton and Boonstra, 1929

Bradysaurus was a large, early and common

pareiasaur. They possessed a covering of armoured scutes, likely serving as defense against their main predators, the gorgonopsians
.

Fossils of Bradysaurus are known from the

Period
.

Description

B. baini
B. seeleyi

Bradysaurus was 2.5–3 m (8 ft 2 in – 9 ft 10 in) in length

phalangeal
count being 2,3,3,3,2 on the fore-foot and 2,3,3,4,3 on the hind. The whole body is protected by dermal scutes, although these are not as thick or heavy as in more advanced forms.

Classification and species

Bradysaurus is the only member of the subfamily Bradysaurinae. It is the most primitive known pareiasaur and can be considered a good ancestral type from which the others developed. Its large dimensions show that, even very early in their evolutionary history, these strange animals had already attained an optimal size. Even later, more advanced forms, like Scutosaurus, were no larger. The advantage of large size was to provide defense against predators and to maintain a stable body temperature (gigantothermy).

Kuhn 1969 lists no fewer than nine species for this genus, but this is certainly an excessive number. Boonstra 1969 distinguishes only four species on the basis of tooth structure, two of which Kuhn places in the genus Embrithosaurus. The genera Brachypareia, Bradysuchus, Koalemasaurus, and Platyoropha are synonyms of Bradysaurus.

B. baini (Seeley, 1892) is from the Tapinocephalus zone, Lower Beaufort Beds, Karoo basin, South Africa. This is the type species for the genus. The quadra-jugal region (cheek-bones) were only moderately developed. The snout was broad and rounded and there were 15 or 16 pairs of overlapping teeth in each jaw. This animal could be considered a generic early pareiasaur. According to Lee, 1997, the available material of B. baini lacks distinguishing autapomorphies or characteristics.

B. seeleyi (Haughton and Boonstra, 1929) is from the Tapinocephalus zone, Lower Beaufort Beds, Karoo basin, South Africa. This is a less common form. Boonstra, 1969, considered this a valid species of Bradysaurus and Lee, 1997, considers this animal a sister group to more advanced pareiasaurs. B. seelyi seems to be closely related to Nochelesaurus and Embrithosaurus. In contrast to the more numerous but similarly sized B. baini, the cheekbones were heavy and greatly enlarged. There were 19 or 20 pairs of strongly overlapping teeth on each jaw.

References

  1. hdl:11573/1634310.{{cite journal}}: CS1 maint: multiple names: authors list (link
    )
  • Boonstra, L. D. 1969, "The Fauna of the Tapinoephalus Zone (Beaufort Beds of the Karoo)," Ann. S. Afr. Mus. 56 (1) 1-73, pp. 29–32
  • Edwin H. Colbert, 1965, The Age of Reptiles, The World Naturalist, Weidenfeld and Nicolson, London, pp. 52–3
  • Barry Cox, R.J.G.Savage, Brian Gardiner, Dougal Dixon, 1988 Illustrated Encyclopaedia of Dinosaurs and Prehistoric Animals
  • Carroll Lane Fenton and Mildred Adams Fenton, 1958, The Fossil Book, Doubleday & Co., Garden City, New York, p. 306
  • Kuhn, O, 1969, Cotylosauria, part 6 of Handbuch der Palaoherpetologie (Encyclopedia of Palaeoherpetology), Gustav Fischer Verlag, Stuttgart & Portland,
  • Lee, MSY (1997), Pareiasaur phylogeny and the origin of turtles. Zool. J. Linnean Soc., 120: 197-280

External links