Camel

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Camel
Temporal range: Pliocene–Recent[1]
A one-humped camel
Dromedary
(Camelus dromedarius)
A shaggy two-humped camel
Bactrian camel
(Camelus bactrianus)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Camelidae
Tribe: Camelini
Genus: Camelus
Linnaeus, 1758
Type species
Camelus dromedarius [6]

Linnaeus, 1758
Species
Distribution of camels worldwide
Synonyms
List

A camel (from

critically endangered
.

The word camel is also used informally in a wider sense, where the more correct term is "camelid", to include all seven species of the family

Bering land bridge into Asia during the late Miocene
, around 6 million years ago.

Taxonomy

Extant species

Three species are

Genus CamelusLinnaeus, 1758 – nine species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bactrian camel

Camelus bactrianus

Linnaeus, 1758
Domesticated; Central Asia, including the historical region of Bactria.
Map of range
Size:

Habitat:

Diet:
Dromedary / Arabian camel

Camelus dromedarius

Linnaeus, 1758
Domesticated; the Middle East, Sahara Desert, and South Asia; introduced to Australia
Map of range
Size:

Habitat:

Diet:
Wild Bactrian camel

Camelus ferus

Przewalski, 1878
Remote areas of
northwest China and Mongolia
Map of range
Size:

Habitat:

Diet:
 CR 


Biology

The average life expectancy of a camel is 40 to 50 years.[12] A full-grown adult dromedary camel stands 1.85 m (6 ft 1 in) at the shoulder and 2.15 m (7 ft 1 in) at the hump.[13] Bactrian camels can be a foot taller. Camels can run at up to 65 km/h (40 mph) in short bursts and sustain speeds of up to 40 km/h (25 mph).[14] Bactrian camels weigh 300 to 1,000 kg (660 to 2,200 lb) and dromedaries 300 to 600 kg (660 to 1,320 lb). The widening toes on a camel's hoof provide supplemental grip for varying soil sediments.[15]

The male dromedary camel has an organ called a dulla in his throat, a large, inflatable sac that he extrudes from his mouth when in

ejaculates three or four times within a single mating session.[18] Camelids are the only ungulates to mate in a sitting position.[19]

Ecological and behavioral adaptations

Camel humps store fat for when food is scarce. If a camel uses the fat, the hump will become limp and droop

Camels do not directly store water in their humps; they are reservoirs of fatty tissue. When this tissue is metabolized, it yields a greater mass of water than that of the fat processed. This fat metabolization, while releasing energy, causes water to evaporate from the lungs during respiration (as oxygen is required for the metabolic process): overall, there is a net decrease in water.[20][21]

A portrait of a camel with a visibly thick mane
A camel's thick coat is one of its many adaptations that aid it in desert-like conditions.
A leashed pack camel
A camel in Somalia, which has the world's largest camel population[22]

Camels have a series of physiological adaptations that allow them to withstand long periods of time without any external source of water.[23] The dromedary camel can drink as seldom as once every 10 days even under very hot conditions, and can lose up to 30% of its body mass due to dehydration.[24] Unlike other mammals, camels' red blood cells are oval rather than circular in shape. This facilitates the flow of red blood cells during dehydration[25] and makes them better at withstanding high osmotic variation without rupturing when drinking large amounts of water: a 600 kg (1,300 lb) camel can drink 200 L (53 US gal) of water in three minutes.[26][27][failed verification]

Camels are able to withstand changes in

cardiac failure results from circulatory disturbance.[27]

When the camel exhales,

nostrils and is reabsorbed into the body as a means to conserve water.[31] Camels eating green herbage can ingest sufficient moisture in milder conditions to maintain their bodies' hydrated state without the need for drinking.[32]

Domesticated camel calves lying in sternal recumbency, which aids heat loss

The camel's thick coat insulates it from the intense heat radiated from desert sand; a shorn camel must sweat 50% more to avoid overheating.[33] During the summer the coat becomes lighter in color, reflecting light as well as helping avoid sunburn.[27] The camel's long legs help by keeping its body farther from the ground, which can heat up to 70 °C (158 °F).[34][35] Dromedaries have a pad of thick tissue over the sternum called the pedestal. When the animal lies down in a sternal recumbent position, the pedestal raises the body from the hot surface and allows cooling air to pass under the body.[29]

Camels' mouths have a thick leathery lining, allowing them to chew thorny desert plants. Long eyelashes and ear hairs, together with nostrils that can close, form a barrier against sand. If sand gets lodged in their eyes, they can dislodge it using their translucent

third eyelid (also known as the nictitating membrane). The camels' gait and widened feet help them move without sinking into the sand.[34][36]

The

renal corpuscles have a smaller diameter, which reduces surface area for filtration. These two major anatomical characteristics enable camels to conserve water and limit the volume of urine in extreme desert conditions.[38] Camel urine comes out as a thick syrup, and camel faeces are so dry that they do not require drying when used to fuel fires.[39][40][41][42]

The camel

Genetics

The

flow sorted camel chromosomes, building on the fact that camels have 37 pairs of chromosomes (2n=74), and found that the karyotype consisted of one metacentric, three submetacentric, and 32 acrocentric autosomes. The Y is a small metacentric chromosome, while the X is a large metacentric chromosome.[53]

Skull of an F1 hybrid camel, Museum of Osteology, Oklahoma

The hybrid camel, a hybrid between Bactrian and dromedary camels, has one hump, though it has an indentation 4–12 cm (1.6–4.7 in) deep that divides the front from the back. The hybrid is 2.15 m (7 ft 1 in) at the shoulder and 2.32 m (7 ft 7 in) tall at the hump. It weighs an average of 650 kg (1,430 lb) and can carry around 400 to 450 kg (880 to 990 lb), which is more than either the dromedary or Bactrian can.[54]

According to molecular data, the wild Bactrian camel (C. ferus) separated from the domestic Bactrian camel (C. bactrianus) about 1 million years ago.

cloven hooves.[61][62] Like the mule, camas are sterile, despite both parents having the same number of chromosomes.[60]

Evolution

The earliest known camel, called Protylopus, lived in North America 40 to 50 million years ago (during the Eocene).[18] It was about the size of a rabbit and lived in the open woodlands of what is now South Dakota.[63][64] By 35 million years ago, the Poebrotherium was the size of a goat and had many more traits similar to camels and llamas.[65][66] The hoofed Stenomylus, which walked on the tips of its toes, also existed around this time, and the long-necked Aepycamelus evolved in the Miocene.[67] The split between the tribes Camelini, which contains modern camels and Lamini, modern llamas, alpacas, vicuñas, and guanacos, is estimated to have occurred over 16 million years ago.[68]

The ancestor of modern camels,

guanacos and related animals.[18][63][64] Populations of Paracamelus continued to exist in the North American Arctic into the Early Pleistocene.[72][73] This creature is estimated to have stood around nine feet (2.7 metres) tall. The Bactrian camel diverged from the dromedary about 1 million years ago, according to the fossil record.[74]

The last camel native to North America was

Quaternary extinction event, coinciding with the migration of humans from Asia at the end of the Pleistocene, around 13–11,000 years ago.[75][76]

Domestication

A camel carrying supplies, Tang dynasty
A man on a camel, Tang dynasty
Woman on a camel breastfeeding, Tang dynasty

Like horses, camels originated in North America and eventually spread across Beringia to Asia. They survived in the Old World, and eventually humans domesticated them and spread them globally. Along with many other megafauna in North America, the original wild camels were wiped out during the spread of the first indigenous peoples of the Americas from Asia into North America, 10 to 12,000 years ago; although fossils have never been associated with definitive evidence of hunting.[75][76]

Most camels surviving today are domesticated.[42][77] Although feral populations exist in Australia, India and Kazakhstan, wild camels survive only in the wild Bactrian camel population of the Gobi Desert.[12]

History

When humans first domesticated camels is disputed. Dromedaries may have first been domesticated by humans in Somalia or South Arabia sometime during the 3rd millennium BC, the Bactrian in central Asia around 2,500 BC,[18][78][79][80] as at Shar-i Sokhta (also known as the Burnt City), Iran.[81] A study from 2016, which genotyped and used world-wide sequencing of modern and ancient mitochondrial DNA (mtDNA), suggested that they were initially domesticated in the southeast Arabian Peninsula,[82] with the Bactrian type later being domesticated around Central Asia.[83]

Martin Heide's 2010 work on the domestication of the camel tentatively concludes that humans had domesticated the Bactrian camel by at least the middle of the third millennium somewhere east of the Zagros Mountains, with the practice then moving into Mesopotamia. Heide suggests that mentions of camels "in the patriarchal narratives may refer, at least in some places, to the Bactrian camel", while noting that the camel is not mentioned in relationship to Canaan.[84] Heide and Joris Peters reasserted that conclusion in their 2021 study on the subject.[85]

In 2009-2013, excavations in the Timna Valley by Lidar Sapir-Hen and Erez Ben-Yosef discovered what may be the earliest domestic camel bones yet found in Israel or even outside the Arabian Peninsula, dating to around 930 BC. This garnered considerable media coverage, as it is strong evidence that the stories of Abraham, Jacob, Esau, and Joseph were written after this time.[86][87]

The existence of camels in Mesopotamia—but not in the eastern Mediterranean lands—is not a new idea. The historian Richard Bulliet did not think that the occasional mention of camels in the Bible meant that the domestic camels were common in the Holy Land at that time.[88] The archaeologist William F. Albright, writing even earlier, saw camels in the Bible as an anachronism.[89]

The official report by Sapir-Hen and Ben-Joseph says:

The introduction of the dromedary camel (Camelus dromedarius) as a pack animal to the southern Levant ... substantially facilitated trade across the vast deserts of Arabia, promoting both economic and social change (e.g., Kohler 1984; Borowski 1998: 112–116; Jasmin 2005). This ... has generated extensive discussion regarding the date of the earliest domestic camel in the southern Levant (and beyond) (e.g., Albright 1949: 207; Epstein 1971: 558–584; Bulliet 1975; Zarins 1989; Köhler-Rollefson 1993; Uerpmann and Uerpmann 2002; Jasmin 2005; 2006; Heide 2010; Rosen and Saidel 2010; Grigson 2012). Most scholars today agree that the dromedary was exploited as a pack animal sometime in the early Iron Age (not before the 12th century [BC])

and concludes:

Current data from copper smelting sites of the Aravah Valley enable us to pinpoint the introduction of domestic camels to the southern Levant more precisely based on stratigraphic contexts associated with an extensive suite of radiocarbon dates. The data indicate that this event occurred not earlier than the last third of the 10th century [BC] and most probably during this time. The coincidence of this event with a major reorganization of the copper industry of the region—attributed to the results of the campaign of Pharaoh Shoshenq I—raises the possibility that the two were connected, and that camels were introduced as part of the efforts to improve efficiency by facilitating trade.[87]

Textiles

Desert tribes and Mongolian nomads use camel hair for tents, yurts, clothing, bedding and accessories. Camels have outer guard hairs and soft inner down, and the fibers are sorted[by whom?] by color and age of the animal. The guard hairs can be felted for use as waterproof coats for the herdsmen, while the softer hair is used for premium goods.[90] The fiber can be spun for use in weaving or made into yarns for hand knitting or crochet. Pure camel hair is recorded as being used for western garments from the 17th century onwards, and from the 19th century a mixture of wool and camel hair was used.[91]

Military uses

A special BSF camel contingent, Republic Day Parade, New Delhi (2004)
A painting of soldiers on camels
Camel Corps at Magdhaba, Egypt, 23 December 1916, by Harold Septimus Power (1925)

By at least 1200 BC the first camel saddles had appeared, and

Bactrian camels could be ridden. The first saddle was positioned to the back of the camel, and control of the Bactrian camel was exercised by means of a stick. However, between 500 and 100 BC, Bactrian camels came into military use. New saddles, which were inflexible and bent, were put over the humps and divided the rider's weight over the animal. In the seventh century BC the military Arabian saddle evolved, which again improved the saddle design slightly.[92][93]

Military forces have used camel cavalries in wars throughout Africa, the Middle East, and into the modern-day Border Security Force (BSF) of India (though as of July 2012, the BSF planned the replacement of camels with ATVs). The first documented use of camel cavalries occurred in the Battle of Qarqar in 853 BC.[94][95][96] Armies have also used camels as freight animals instead of horses and mules.[97][98]

The

Achaemenid Persians when fighting Lydia in the Battle of Thymbra (547 BC).[54][101][102]

19th and 20th centuries

, 1912

The

Secretary of War in 1858, recommended that funds be allocated towards obtaining a thousand more camels), the outbreak of the American Civil War in 1861 saw the end of the Camel Corps: Texas became part of the Confederacy, and most of the camels were left to wander away into the desert.[98]

France created a

Tuareg and Arab insurgents, as previous efforts to defeat them on foot had failed.[105] The Free French Camel Corps fought during World War II, and camel-mounted units remained in service until the end of French rule over Algeria in 1962.[106]

In 1916, the British created the

Sinai and Palestine Campaign in World War I. The Imperial Camel Corps comprised infantrymen mounted on camels for movement across desert, though they dismounted at battle sites and fought on foot. After July 1918, the Corps began to become run down, receiving no new reinforcements, and was formally disbanded in 1919.[107]

In World War I, the British Army also created the Egyptian Camel Transport Corps, which consisted of a group of Egyptian camel drivers and their camels. The Corps supported British war operations in Sinai, Palestine, and Syria by transporting supplies to the troops.[108][109][110]

The Somaliland Camel Corps was created by colonial authorities in British Somaliland in 1912; it was disbanded in 1944.[111]

Bactrian camels were used by Romanian forces during World War II in the Caucasian region.[112] At the same period the Soviet units operating around Astrakhan in 1942 adopted local camels as draft animals due to shortage of trucks and horses, and kept them even after moving out of the area. Despite severe losses, some of these camels ended up as far west as to Berlin itself.[113]

The

British India fought alongside the British Indian Army in World Wars I and II.[114]

The

Sahrawi tribesmen serving in the colonial army in Spanish Sahara (today Western Sahara). Operational from the 1930s until the end of the Spanish presence in the territory in 1975, the Tropas Nómadas were equipped with small arms and led by Spanish officers. The unit guarded outposts and sometimes conducted patrols on camelback.[115][116]

21st century competition

The annual King Abdulaziz Camel Festival is held in Saudi Arabia. In addition to camel racing and camel milk tasting, the festival holds a camel "

botox.[117] In a similar incident in 2021, over 40 camels were disqualified.[118]

Food uses

Camel meat and milk are foods that are found in many cuisines, typically in

Dairy

Camels at the Khan and old bridge, Lajjun, Palestine (now in Israel) - 1870s drawing
A camel calf nursing on camel milk

Camel milk is a staple food of desert nomad tribes and is sometimes considered a meal itself; a nomad can live on only camel milk for almost a month.[19][39][123][124]

Camel milk can readily be made into

École Nationale Supérieure d'Agronomie et des Industries Alimentaires, who was able to produce curdling by the addition of calcium phosphate and vegetable rennet in the 1990s.[126] The cheese produced from this process has low levels of cholesterol and is easy to digest, even for the lactose intolerant.[127][128]

Camel milk can also be made into ice cream.[129][130]

Meat

A Somali camel meat and rice dish
Camel meat pulao, from Pakistan

Camels provide food in the form of meat and milk.[131] Approximately 3.3 million camels and camelids are slaughtered each year for meat worldwide.[132] A camel carcass can provide a substantial amount of meat. The male dromedary carcass can weigh 300–400 kg (661–882 lb), while the carcass of a male Bactrian can weigh up to 650 kg (1,433 lb). The carcass of a female dromedary weighs less than the male, ranging between 250 and 350 kg (550 and 770 lb).[18] The brisket, ribs and loin are among the preferred parts, and the hump is considered a delicacy.[133] The hump contains "white and sickly fat", which can be used to make the khli (preserved meat) of mutton, beef, or camel.[134] On the other hand, camel milk and meat are rich in protein, vitamins, glycogen, and other nutrients making them essential in the diet of many people. From chemical composition to meat quality, the dromedary camel is the preferred breed for meat production. It does well even in arid areas due to its unusual physiological behaviors and characteristics, which include tolerance to extreme temperatures, radiation from the sun, water paucity, rugged landscape and low vegetation.[135] Camel meat is reported to taste like coarse beef, but older camels can prove to be very tough,[13][18] although camel meat becomes tenderer the more it is cooked.[136]

Camel is one of the animals that can be ritually slaughtered and divided into three portions (one for the home, one for extended family/social networks, and one for those who cannot afford to slaughter an animal themselves) for the qurban of Eid al-Adha.[137][138]

The

Karachi, Pakistan, some restaurants prepare nihari from camel meat.[140] Specialist camel butchers provide expert cuts, with the hump considered the most popular.[141]

Camel meat has been eaten for centuries. It has been recorded by

Persia, usually roasted whole.[142] The Roman emperor Heliogabalus enjoyed camel's heel.[39] Camel meat is mainly eaten in certain regions, including Eritrea, Somalia, Djibouti, Saudi Arabia, Egypt, Syria, Libya, Sudan, Ethiopia, Kazakhstan, and other arid regions where alternative forms of protein may be limited or where camel meat has had a long cultural history.[18][39][133] Camel blood is also consumable, as is the case among pastoralists in northern Kenya, where camel blood is drunk with milk and acts as a key source of iron, vitamin D, salts and minerals.[18][133][143]

A 2005 report issued jointly by the Saudi Ministry of Health and the United States Centers for Disease Control and Prevention details four cases of human bubonic plague resulting from the ingestion of raw camel liver.[144]

Australia

Camel meat is also occasionally found in

Somalis, and other Australians have also been buying it. The feral nature of the animals means they produce a different type of meat to farmed camels in other parts of the world,[146] and it is sought after because it is disease-free, and a unique genetic group. Demand is outstripping supply, and governments are being urged not to cull the camels, but redirect the cost of the cull into developing the market. Australia has seven camel dairies, which produce milk, cheese and skincare products in addition to meat.[147]

Religion

Islam

Abū Ḥanīfah, the drinking of camel urine is discouraged.[149]

Islamic texts contain several stories featuring camels. In the story of the people of

his she-camel to roam there; the location where the camel stopped to rest determined the location where he would build his house in Medina.[150]

Judaism

According to

kosher.[151] Camels possess only one of the two kosher criteria; although they chew their cud, they do not have cloven hooves: "But these you shall not eat among those that bring up the cud and those that have a cloven hoof: the camel, because it brings up its cud, but does not have a [completely] cloven hoof; it is unclean for you."[152]

The Palestinian Muslim Makhamara clan in Yatta, who claim descent from Jews, reportedly avoid eating camel meat, a practice cited as evidence of their Jewish origins.[153][154]

Cultural depictions

What may be the oldest carvings of camels were discovered in 2018 in Saudi Arabia. They were analysed by researchers from several scientific disciplines and, in 2021, were estimated to be 7,000 to 8,000 years old.[155] The dating of rock art is made difficult by the lack of organic material in the carvings that may be tested, so the researchers attempting to date them tested animal bones found associated with the carvings, assessed erosion patterns, and analysed tool marks in order to determine a correct date for the creation of the sculptures. This Neolithic dating would make the carvings significantly older than Stonehenge (5,000 years old) and the Egyptian pyramids at Giza (4,500 years old) and it predates estimates for the domestication of camels.

  • Shadda (cover,detail), Karabagh region, southwest Caucasus, early 19th century
    Shadda (cover,detail), Karabagh region, southwest Caucasus, early 19th century
  • Vessel in the form of a recumbent camel with jugs, 250 BC – 224 AD, Brooklyn Museum
    Vessel in the form of a recumbent camel with jugs, 250 BC – 224 AD, Brooklyn Museum
  • Maru Ragini (Dhola and Maru Riding on a Camel), c. 1750, Brooklyn Museum
    Maru Ragini (Dhola and Maru Riding on a Camel), c. 1750, Brooklyn Museum
  • The Magi Journeying (Les rois mages en voyage)—James Tissot, c. 1886, Brooklyn Museum
    The Magi Journeying (Les rois mages en voyage)—James Tissot, c. 1886, Brooklyn Museum
  • How the Camel Got His Hump (From Rudyard Kipling's Just So Stories)
    How the Camel Got His Hump (From Rudyard Kipling's Just So Stories)

Distribution and numbers

A view into a canyon: many camels gathering around a watering hole
Camels in the Guelta d'Archei, in northeastern Chad

There are approximately 14 million camels alive as of 2010, with 90% being dromedaries.[156] Dromedaries alive today are domesticated animals (mostly living in the Horn of Africa, the Sahel, Maghreb, Middle East and South Asia). The Horn region alone has the largest concentration of camels in the world,[22] where the dromedaries constitute an important part of local nomadic life. They provide nomadic people in Somalia[18] and Ethiopia with milk, food, and transportation.[124][157][158][159]

A world map with large camel populations marked
Commercial camel market headcount in 2003

Over one million dromedary camels are estimated to be feral in Australia, descended from those introduced as a method of transport in the 19th and early 20th centuries.[160] This population is growing about 8% per year;[161] it was estimated at around 700,000 in 2008.[143][156][162] Representatives of the Australian government have culled more than 100,000 of the animals in part because the camels use too much of the limited resources needed by sheep farmers.[163]

A small population of introduced camels, dromedaries and Bactrians, wandered through

U.S. Camel Corps experiment. When the project ended, they were used as draft animals in mines and escaped or were released. Twenty-five U.S. camels were bought and exported to Canada during the Cariboo Gold Rush.[98]

The Bactrian camel is, as of 2010[update], reduced to an estimated 1.4 million animals, most of which are domesticated.[42][156][164] The Wild Bactrian camel is a separate species and is the only truly wild (as opposed to feral) camel in the world. The wild camels are critically endangered and number approximately 1400, inhabiting the Gobi and Taklamakan Deserts in China and Mongolia.[12][165]

See also

References

Notes

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Bibliography

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