Vitis vinifera
Vitis vinifera | |
---|---|
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Rosids |
Order: | Vitales |
Family: | Vitaceae |
Genus: | Vitis |
Species: | V. vinifera
|
Binomial name | |
Vitis vinifera |
Vitis vinifera, the common grape vine, is a species of flowering plant, native to the Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran.[2] There are currently between 5,000 and 10,000 varieties of Vitis vinifera grapes though only a few are of commercial significance for wine and table grape production.[3]
The wild grape is often classified as Vitis vinifera sylvestris (in some classifications considered Vitis sylvestris), with Vitis vinifera vinifera restricted to cultivated forms. Domesticated vines have
flowers on separate plants) and pollination is required for fruit to develop.Grapes can be eaten fresh or dried to produce
History
Prehistory
This section needs additional citations for verification. (June 2014) |
Wild grapes were harvested by neolithic foragers and early farmers. For thousands of years, the fruit has been harvested for both medicinal and nutritional value; its history is intimately entwined with the history of wine.
Changes in pip (seed) shape (narrower in domesticated forms) and distribution point to domestication occurring about 4100–3000 BC,[4] in southwest Asia, South Caucasus (Armenia[5][6] and Georgia), or the Western Black Sea shore region (Bulgaria, Romania). The earliest evidence of domesticated grapes has been found at Gadachrili Gora, near the village of Imiri, Marneuli Municipality, in southeastern Georgia; carbon-dating points to the date of about 6000 BC. The oldest winery in the world (dating to 4100BCE) was found in the Areni-1 cave, which lies in Areni, Armenia.[7][6] Grape pips dating back to the 5th–4th millennium BC were also found in Shulaveri; others dating back to the 4th millennium BC were also found in Khizanaant Gora. Wild grapes were harvested by neolithic foragers and early farmers. For thousands of years, the fruit has been harvested for both medicinal and nutritional value; its history is intimately entwined with the history of wine.[8]
Antiquity
Cultivation of the domesticated grape spread to other parts of the Old World in pre-historic or early historic times.[9] The first written accounts of grapes and wine can be found in the Epic of Gilgamesh, an ancient Sumerian text from the 3rd millennium BC. There are also numerous hieroglyphic references from ancient Egypt, according to which wine was reserved exclusively for priests, state functionaries and the pharaoh.[10]
Hesiod in his Works and Days gives detailed descriptions of grape harvests and wine making techniques, and there are also many references in Homer. Greek colonists then introduced these practices in their colonies, especially in southern Italy (Magna Grecia), which was even known as Enotria due to its propitious climate.
The
Medieval era
Between the 5th and 10th centuries, viticulture was sustained almost exclusively by the different religious orders in monasteries. The Benedictines and others extended the grape growing limit northwards and also planted new vineyards at higher altitudes than was customary before. Apart from 'ecclesiastical' viticulture, there also developed, especially in France, a 'noble' viticulture, practiced by the aristocracy as a symbol of prestige.
Early modern period
Between the Low Middle Ages and the Renaissance, viticulture began to flourish again. Demographic pressure, population concentration in towns and cities, and the increased spending power of artisans and merchants gave rise to increased investment in viticulture, which became economically feasible once more.[citation needed] Much was written during the Renaissance on grape growing and wine production, favouring a more scientific approach. This literature can be considered the origin of modern ampelography.[citation needed]
Grapes followed European colonies around the world, coming to North America around the 17th century, and to Africa, South America and Australia. In North America it formed hybrids with native species from the genus Vitis; some of these were intentional hybrids created to combat phylloxera, an insect pest which affected the European grapevine to a much greater extent than North American ones and in fact managed to devastate European wine production in a matter of years. Later, North American rootstocks became widely used to graft V. vinifera cultivars so as to withstand the presence of phylloxera.[15]
Contemporary period
NCBI genome ID | 401 |
---|---|
Ploidy | diploid |
Genome size | about 500 Mb |
Number of chromosomes | 19 pairs |
Year of completion | 2008 |
Sequenced organelle | plastid |
In the second half of the 20th century there was a shift in attitude in viticulture from traditional techniques to the scientific method based on fields such as microbiology, chemistry and ampelography. This change came about also due to changes in economic and cultural aspects and in the way of life and in the consumption habits of wide sectors of the population starting to demand quality products.[citation needed]
In 2007, Vitis vinifera was the fourth angiosperm species whose genome was completely sequenced. These data contributed significantly to understanding the evolution of plants and also how the aromatic characteristics of wine are determined in part by the plant's genes.
Also in 2007, scientists from
Description
It is a liana growing 12–15 m (39–49 ft) tall at a fast rate.[19][20] Having a flaky bark, its leaves are alternate, palmately lobed, deciduous, with 3 to 5 pointed lobes, coarsely prickly-toothed leaf margins and a heart-shaped foot, 5–20 cm (2.0–7.9 in) long and broad. They are glossy dark green on top, light green below, usually hairless.
The vine attaches to supports by
The species typically occurs in humid forests and streamsides.
Inflorescences
Their flowers, small and greenish to white, are grouped in
The fruit is a berry, known as a grape that is ovoid or globular, dark blue or greenish, usually 2-locular with 5 seeds; in the wild species it is 6 mm (0.24 in) diameter and ripens dark purple to blackish with a pale wax bloom; in cultivated plants it is usually much larger, up to 3 cm (1.2 in) long, and can be green, red, or purple (black).
Distribution
V. vinifera accounts for the majority of world wine production; all of the most familiar grape varieties used for wine production belong to V. vinifera.[21]
In
Cultivation
Use of
In the Mediterranean Basin, leaves and young stems are traditionally used to feed sheep and goats after grapevine pruning.[26]
Using the sap of grapevines, European folk healers sought to cure skin and eye diseases. Other historical uses include the
Seedless grape varieties were developed to appeal to consumers, but researchers are now discovering that many of the healthful properties of grapes may actually come from the seeds themselves, thanks to their enriched phytochemical content.[27][28]
Grapevine leaves are filled with minced meat (such as lamb, pork or beef), rice and onions in the making of Balkan traditional dolma.
A popular cultivar in Australia, Vitis 'Ornamental Grape', derived from Vitis vinifera x Vitis rupestris, is used in gardens for its impressive foliage that turn brilliant red, scarlet, purple and/or orange in autumn. Originally bred in France, it thrives in a range of climates from hot and dry, to cool moist and subtropical, with different soil types benefitting the plant.[29]
Climate change
Grapevines are very responsive to their surrounding environment with a seasonal variation in yield of 32.5%.
Of all environmental factors, temperature seems to have the most profound effect on viticulture as the temperature during the winter dormancy affects the budding for the following growing season.[35] Prolonged high temperature can have a negative impact on the quality of the grapes as well as the wine as it affects the development of grape components that give colour, aroma, accumulation of sugar, the loss of acids through respiration as well as the presence of other flavour compounds that give grapes their distinctive traits. Sustained intermediate temperatures and minimal day-to-day variability during the growth and ripening periods are favourable. Grapevine annual growth cycles begin in spring with bud break initiated by consistent day time temperatures of 10 degrees Celsius.[36] The unpredictable nature of climate change may also bring occurrences of frosts which may occur outside of the usual winter periods. Frosts cause lower yields and effects grape quality due to reduction of bud fruitfulness and therefore grapevine production benefits from frost free periods.
Organic acids are essential in wine quality. The
Altered precipitation patterns are also anticipated (both annually and seasonally) with rainfall occurrences varying in amount and frequency. Increases in the amount of rainfall have will likely cause an increase in soil erosion; while occasional lack of rainfall, in times when it usually occurs, may result in drought conditions causing stress on grapevines.[42] Rainfall is critical at the beginning of the growing season for the budburst and inflorescence development while consistent dry periods are important for the flowering and ripening periods.[43]
Increased CO2 levels will likely have an effect on the photosynthetic activity in grapevines as photosynthesis is stimulated by a rise in CO2 and has been known to also lead to an increase leaf area and vegetative dry weight.[44] Raised atmospheric CO2 is also believed to result in partial stomatal closure which indirectly leads to increased leaf temperatures. A rise in leaf temperatures may alter ribulose 1,5-bisphosphate carboxylase/oxygenase (RuBisCo) relationship with carbon dioxide and oxygen which will also affect the plants' photosynthesis capabilities.[42] Raised atmospheric carbon dioxide is also known to decrease the stomatal density of some grapevine varieties.[45]
Cultivation variations
The gradually increasing temperatures will lead to a shift in suitable growing regions.[46] It is estimated that the northern boundary of European viticulture will shift north 10 to 30 kilometres (6.2 to 18.6 mi) per decade up to 2020 with a doubling of this rate predicted between 2020 and 2050.[47][needs update] This has positive and negative effects, as it opens doors to new cultivars being grown in certain regions but a loss of suitability of other cultivars and may also risk production quality and quantity in general.[48][46]
Adapting wine production
Systems have been developed to manipulate the temperatures of vines. These include a chamber free system where air can be heated or cooled and then blown across grape bunches to get a 10 °C (50 °F) differential.
Chemistry
Phenolics
V. vinifera contains many phenolic compounds.[52] Anthocyanins can be found in the skin of the berries, hydroxycinnamic acids in the pulp and condensed tannins of the proanthocyanidins type in the seeds. Stilbenoids can be found in the skin and in wood.
Stilbenoids
Trans-
Anthocyanins
Vitis vinifera red cultivars are rich in anthocyanins that impart their colour to the berries (generally in the skin). The 5 most basic anthocyanins found in grape are:
- Cyanidin-3-O-glucoside
- Delphinidin-3-O-glucoside
- Malvidin-3-O-glucoside
- Petunidin-3-O-glucoside
- Peonidin-3-O-glucoside
Cultivars like Graciano[55][56] may also contain :
- acetylated anthocyanins
- Cyanidin-3-(6-acetyl)-glucoside
- Delphinidin-3-(6-acetyl)-glucoside
- Malvidin-3-(6-acetyl)-glucoside
- Petunidin-3-(6-acetyl)-glucoside
- Peonidin-3-(6-acetyl)-glucoside
- coumaroylated anthocyanins
- Cyanidin-3-(6-p-coumaroyl)-glucoside
- Delphinidin-3-(6-p-coumaroyl)-glucoside
- Malvidin-3-(6-p-coumaroyl)-glucoside cis
- Malvidin-3-(6-p-coumaroyl)-glucoside trans
- Petunidin-3-(6-p-coumaroyl)-glucoside
- Peonidin-3-(6-p-coumaroyl)-glucoside
- caffeoylated anthocyanins
Other chemicals
Isoprenoid monoterpenes are present in grape, above all acyclic
See also
References
- ^ Participants of the FFI/IUCN SSC Central Asian regional tree Red Listing workshop, Bishkek, Kyrgyzstan (11-13 July 2006) (2007). "Vitis vinifera". IUCN Red List of Threatened Species. 2007: e.T63537A12687723. Retrieved 8 February 2024.
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- ^ Hotz, Robert Lee (11 January 2011). "Perhaps a Red, 4,100 B.C." Wall Street Journal. Retrieved 21 August 2016.
- ^ "6,000-year-old winery found in Armenian cave (Wired UK)". Archived from the original on 8 December 2015. Retrieved 1 November 2015.
- ^ a b James Owen (12 January 2011). "Earliest Known Winery Found in Armenian Cave". National Geographic News. Archived from the original on 12 January 2011. Retrieved 2 April 2019.
- ^ "Areni-1 Cave, Armenia: A Chalcolithic–Early Bronze Age settlement and ritual site in the southern Caucasus". Research Gate. 1 March 2012. Retrieved 10 April 2017.
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- ^ Eijkhoff, P. (2000). Wine in China; its history and contemporary developments.
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- ^ Heuzé V., Thiollet H., Tran G., 2017. Grape leaves and vine shoots. Feedipedia, a programme by INRA, CIRAD, AFZ and FAO. https://www.feedipedia.org/node/512.
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Further reading
- Francesco Emanuelli; Silvia Lorenzi; Lukasz Grzeskowiak; Valentina Catalano; Marco Stefanini; Michela Troggio; Sean Myles; José M. Martinez-Zapater; Eva Zyprian; Flavia M. Moreira & M. Stella Grando (2013). "Genetic diversity and population structure assessed by SSR and SNP markers in a large germplasm collection of grape". BMC Plant Biology. 13: 39. PMID 23497049.
- Manzi Luigi, La viticoltura e l'enologia al tempo dei romani, Er. Botta, Roma 1883.
- Marescalchi Arturo, Dalmasso Giovanni, Storia della vite e del vino in Italia, 3 voll., Unione Italiana Vini, Milano 1931-33-37.
- Daniel Zohary; Maria Hopf (2000). Domestication of plants in the Old World. Oxford: Oxford University Press. ISBN 978-0-19-850356-9.