Coptoclavidae

Coptoclavidae Temporal range: Late Triassic–Early Cretaceous PreꞒ Ꞓ O S D C P T J K Pg N
Holcoptera giebeli (specimen NMHUK PI II.3101)
Larva of Coptoclava
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Adephaga
Family:	†Coptoclavidae Ponomarenko, 1961
Subfamilies
Hispanoclavininae Timarchopsinae Coptoclavidae (incertae sedis )

Coptoclavidae is an extinct family of aquatic

• †Agrascapha Lin, 1992 – Huangshanjie Formation, China, Carnian
• †Amblycephalonius Bode, 1953 – Posidonia Shale, Germany, Toarcian
• subfamily †Charonoscaphinae Ponomarenko, 1977
  • †Charonoscapha Ponomarenko, 1977 – Karabastau Formation, Kazakhstan, Callovian
  • †Charonoscaphidia Ponomarenko, 1977 – Karabastau Formation, Kazakhstan, Callovian
• †Coptoclavia Ping, 1928 – Cassange Group, Upper Triassic
• subfamily †Coptoclavinae Ponomarenko, 1961
  • †Bolbonectes Ponomarenko, 1987
    • †Bolbonectes intermedius Ponomarenko, 1987 – Byankino Formation, Russia, Tithonian; Leskovskaya Formation, Russia, Barremian
    • †Bolbonectes occidentalis Ponomarenko, 1993 – Ichetuy Formation, Russia, Oxfordian
    • †Bolbonectus lithographicus Ponomarenko and Martínez-Delclòs, 2000 – La Pedrera de Rúbies Formation, Spain, Barremian
  • †Coptoclava Ping, 1928 – Ustkarskaya Formation, Russia, Hauterivian; Dabeigou Formation, China, Hauterivian; Kutinskaya Formation, Russia, Barremian; Dalazi Formation, Laiyang Formation, Jianchang Formation, Shouchang Formation, Jiufotang Formation, Chijinbao Formation, Aptian; Zaza Formation, Gidarinskaya Formation, Mangutskaya Formation, Turga Formation, Russia, Aptian
  • †Hoyaclava Soriano et al., 2007 – Las Hoyas, Spain, Barremian
  • †Megacoptoclava Ponomarenko and Martínez-Delclòs, 2000 – Las Hoyas, Spain, Barremian
• subfamily †Coptoclaviscinae Soriano et al., 2007
  • †Coptoclavella Ponomarenko, 1980
    • †Coptoclavella elegans Ponomarenko, 1980 – Mogotuin Formation, Mongolia, Aptian
    • †Coptoclavella inexpecta Soriano et al., 2007 – La Pedrera de Rúbies Formation, Spain, Barremian
    • †Coptoclavella jurassica Ponomarenko, 2014 – Sharteg, Mongolia, Tithonian
    • †Coptoclavella minor Ponomarenko, 1980 – Takshin Formation, Kalgan Formation Russia, Callovian; Daya Formation, Russia, Hauterivian; Mogotuin Formation, Mongolia, Aptian; Turga Formation, Russia, Aptian
    • †Coptoclavella purbeckensis Ponomarenko et al., 2005 – Durlston Formation, United Kingdom, Berriasian
    • †Coptoclavella vittata Ponomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Aptian
    • †Coptoclavella striata Ponomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Aptian
  • †Coptoclavisca Ponomarenko, 1987 – Laiyang Formation, China, Aptian; Tsagaantsav Formation, Mongolia, Early Cretaceous
  • †Stargelytron Ponomarenko, 2015 – Röt Formation, Germany, Anisian; Hassberge Formation, Germany, Carnian
• †Euroscapha Lin, 1992 – Huangshanjie Formation, China, Carnian
• †Holcoptera Handlirsch, 1906
  • †Holcoptera alisonae Kelly et al., 2017 – Charmouth Mudstone Formation, United Kingdom, Sinemurian
  • †Holcoptera giebeli Handlirsch, 1906 – Lilstock Formation, United Kingdom, Rhaetian; Charmouth Mudstone Formation, United Kingdom, Sinemurian
  • †Holcoptera pigmentatus Kelly et al., 2017 – Lilstock Formation, United Kingdom, Rhaetian; Portland Formation, Connecticut, Hettangian; Mount Toby Formation, Massachusetts, Hettangian
  • †Holcoptera schlotheimi (Giebel, 1856) (=Holcoptera confluens Cockerell, 1915) – Argilliti di Riva di Solto Formation, Italy, Norian; Lilstock Formation, United Kingdom, Rhaetian; Blue Lias, United Kingdom, Hettangian; Portland Formation, Connecticut, Hettangian
  • †Holcoptera solitensis Kelly et al., 2017 – Cow Branch Formation, North Carolina, Norian
• subfamily †Hispanoclavinae Soriano et al., 2007
  • †Hispanoclavina Soriano et al., 2007 – Las Hoyas, Spain, Barremian
• subfamily †Timarchopsinae Wang et al., 2010
  • †Actea Germar, 1842 – Solnhofen, Germany, Tithonian
  • †
    Daohugou
    , China, Callovian
  • †Ditomoptera Germar, 1839 – Solnhofen, Germany, Tithonian
  • †Exedia Ponomarenko, 1977 – Karabastau Formation, Kazakhstan, Callovian
  • †Ovonectes Soriano et al., 2007 – Las Hoyas, Spain, Barremian
  • †Protonectes Prokin and Ponomarenko, 2013 – Hassberge Formation, Germany, Carnian
  • †Pseudohydrophilus Deichmüller, 1886 – Solnhofen, Germany, Tithonian
  • †Stygeonectes Ponomarenko, 1977 – Cheremkhovskaya Formation, Russia, Toarcian; Udinskaya Formation, Russia, Callovian; Uda Formation, Ichetuy Formation, Russia, Oxfordian; Glushkovo Formation, Russia, Tithonian; Leskovskaya Formation, Bukachacha Formation, Russia, Barremian; Godymboyskaya Formation, Turga Formation, Russia, Aptian
  • †Timarchopsis Brauer et al., 1889
    • †Timarchopsis aquaticus Ponomarenko, 1977 – Cheremkhovskaya Formation, Russia, Toarcian; Ichetuy Formation, Russia, Oxfordian
    • †Timarchopsis cyrenaicus Ponomarenko, 1977 – Algeria, Hauterivian
    • †Timarchopsis czekanowskii Brauer et al., 1889 – Cheremkhovskaya Formation, Russia, Toarcian
    • †Timarchopsis gigas Ponomarenko, 1977 – Karabastau Formation, Kazakhstan, Callovian
    • †Timarchopsis gobiensis Ponomarenko, 1987 – Ulaan-Ereg Formation, Mongolia, Tithonian
    • †Timarchopsis latus Ponomarenko, 1977 – Sagul Formation, Kyrgyzstan, Toarcian
    • †Timarchopsis longus Ponomarenko, 2014 – Sharteg, Mongolia, Tithonian
    • †Timarchopsis mongolicus Ponomarenko, 1985 – Zhargalant Formation, Mongolia, Bathonian
    • †Timarchopsis sainshandensis Ponomarenko, 1987 – Khamarkhoburinskaya Formation, Mongolia, Aalenian
  • †Tuhanectes Hong, 1995 – Sanjianfang Formation, China, Bathonian