Diversity of fish

Source: Wikipedia, the free encyclopedia.

angler fish that "angles
" for its prey with a lure attached to a line from its head.

Fish are very diverse animals and can be categorised in many ways. Although most fish species have probably been discovered and described, about 250 new ones are still discovered every year. According to FishBase about 34,800 species of fish had been described as of February 2022,[5] which is more than the combined total of all other vertebrate species: mammals, amphibians, reptiles and birds.

Fish species diversity is roughly divided equally between

Neotropical fishes represent about 10% of all vertebrate species on the Earth. Exceptionally rich sites in the Amazon basin, such as Cantão State Park, can contain more freshwater fish species than occur in all of Europe.[6]

By taxonomy

Further information: Evolution of fish

Fish

taxa into systems. It is complex and still evolving. Controversies over "arcane, but important, details of classification are still quietly raging".[7]

The term "fish" describes any non-

paraphyletic, since the tetrapod clade is within the clade of lobe-finned fishes.[9][10]

Jawless fish

anadromous
, moving between both fresh and salt water habitats.

Extant jawless fish are either lamprey or hagfish. Juvenile lamprey feed by sucking up mud containing micro-organisms and organic debris. The lamprey has well-developed eyes, while the hagfish has only primitive eyespots. The hagfish coats itself and carcasses it finds with noxious slime to deter predators, and periodically ties itself into a knot to scrape the slime off. It is the only invertebrate fish and the only animal which has a skull but no vertebral column.[11]

Cartilaginous fish

placoid scales (dermal denticles) that are as rough as sandpaper. Because cartilaginous fish do not have bone marrow, the spleen and special tissue around the gonads produces red blood cells. Their tails can be asymmetric, with the upper lobe longer than the lower lobe. Some cartilaginous fishes possess an organ called a Leydig's organ
which also produces red blood cells.

There are over 980 species of cartilaginous fish. They include sharks, rays and chimaera.

Bony fish

lepidotrichia
or "fin rays", their fins being webs of skin supported by bony or horny spines ("rays").

There are three types of ray finned fishes: the

teleosts
. The chondrosteans and holosteans are among the earlier fish to evolve, and share characteristics with both teleosts and sharks. In comparison with the other chondrosteans, the holosteans are closer to the teleosts and further from sharks.

Teleosts

recreational fishes.[13]

Teleosts have a movable

caudal fin.[14]

By habitat

See also:
Marine habitats

There is 10,000 times as much saltwater in the oceans as there is freshwater in the lakes and rivers. However, only 58 per cent of

anadromous).[16] This diversity in freshwater species is perhaps not surprising, since the thousands of separate lake habitats promote speciation.[17]

Habitat Area Volume Depth Species Fish biomass
million km2 million cu km (mean) count per cent million tonnes
Saltwater 361[18] 1370.8[19] 3.8 km 18,000 58[16] 800–2,000[20]
Freshwater 1.5[21] 0.13[22] 87 m 13,000 41[16]

Fish can also be

bathypelagic fish
inhabiting the cold and pitch black depths below.

Most oceanic species (78 per cent, or 44 per cent of all fish species), live near the

epipelagic, 5 per cent are pelagic, and 7 per cent are deep water.[16]

Fish are found in nearly all natural aquatic environments.[23] Most fish, whether by species count or abundance, live in warmer environments with relatively stable temperatures.[17] However, some species survive temperatures up to 44.6 °C (112.3 °F), while others cope with colder waters; there are over 200 finfish species south of the Antarctic Convergence.[24] Some fish species tolerate salinities over 10 per cent.[23]

Habitat Abyssobrotula galatheae The world's deepest living fish,
cusk eel, lives in the Puerto Rico Trench at a depth of 8,372 meters (27,467 ft).[23][25] Due to the extreme pressure, this appears to be around the theoretical maximum depth possible for fish.[26][27]
Stone loach
 At the other extreme, the Tibetan
stone loach lives at altitudes over 5,200 meters (17,100 ft) in the Himalayas.[23][28]
Blue shark Some marine pelagic fish range over vast areas, such as the blue shark that lives in all oceans.
Blind cave fish
 Other fish are confined to single, small living spaces, such as the
blind cave fish in North America.[29]
Death Valley
pupfish 		Equally isolated desert pupfish, like the Death Valley pupfish (pictured), live in small desert spring systems in Mexico and the southwest United States.
Thermichthys
hollisi 		The
thermal vents 2,400 metres (1,300 fathoms) deep.[23][30]
Sargassum frogfish
 The highly camouflaged
sargassum frogfish lives in drifting sargassum seaweed. It has adapted fins which can grab strands of sargassum, enabling it to climb through the seaweed.[31] It avoids threats from larger predator fish by climbing out of water onto the surface of a seaweed mat, where it can survive for some time.[32]

By life span

Some of the shortest-lived species are gobies, which are small coral reef–dwelling fish. Some of the longest-lived are rockfish.

Life
span
Seven-figure pygmy goby
External image
image icon World's shortest lived fish

seven-figure pygmy goby is the shortest lived of all fish species. It lives at most for 59 days, which is the shortest lifespan for any vertebrate.[33]

Ram cichlid Short lived fish have particular value in genetic studies on aging. In particular, the ram cichlid is used in laboratory studies because of its ease of breeding and predictable aging pattern.[34][35]
Rougheye rockfish Some of the longest living fishes are
North Pacific at 25–900 metres (14–492 fathoms) and exhibits negligible senescence.[36][37][38]
Orange
roughy 		The orange roughy may be the longest lived commercial fish, with a maximum reported age of 149 years.[39]
Koi There are stories about Japanese koi goldfish passed from generation to generation for 300 years. Scientists are sceptical. Counting growth lines on the scales of fish confined to ponds or bowls is unreliable, since they lay down extra lines.[40][41] The maximum reliably reported age for a goldfish is 41 years.[42]
Atlantic tarpon One of the longest living
sport fish is the Atlantic tarpon, with a reported age of 55 years.[43][44][45]
Green sturgeon Some of the longest living fish are living fossils, such as the green sturgeon. This species is among the longest-living species found in freshwater, with a reported age of 60 years. They are also among the largest fish species found in freshwater, with a maximum reported length of 2.5 meters (8.2 ft) and a maximum reported weight of 159 kg (351 lb).[46][47][48]
Australian lungfish Another living fossil is the Australian lungfish. One individual has lived in an aquarium for at least 75 years, and is the oldest fish in captivity. According to fossil records, the Australian lungfish has hardly changed for 380 million years.[49][50][51]
Greenland shark The Greenland shark has a lifespan of 392 ± 120 years. This is the longest known lifespan of all vertebrate species.[52]

By size

Size Paedocypris progenetica
External image
image icon World's smallest fish

peat swamps of the Indonesian island of Sumatra. The females of this species have a standard length of 7.9 mm (0.31 in) at maturity.[53][54][55] Until recently, this was the smallest of all known vertebrates. However, in 2012 a minute Papua New Guinea frog, Paedophryne amauensis, with a standard length of 7.7 mm (0.30 in) was discovered.[56]
The slender Indonesian fish may still be the smallest vertebrate by weight.

Photocorynus spiniceps
 		Male individuals of the
sexual parasitism on the larger female.[57][58][59][60]
Stout infantfish
 The
stout infantfish, a type of goby, is the second smallest known fish.[61]
Females grow to a length of 8.4 millimetres (0.33 in) and males are mature at 7 millimetres (0.28 in).
Sinarapan According to the
Guinness Book of World Records, the sinarapan, also a goby, is the world's smallest commercially harvested fish.[62] Found in the Philippines, they have an average length of 12.5 mm (0.49 in), and are threatened by overfishing.[55]
Whale shark The largest fish is the whale shark. It is a slow-moving, filter-feeding shark with a maximum published length of 20 m (66 ft) and a maximum weight of 34 tonnes (33 long tons; 37 short tons). Whale sharks can live up to 70 years[63] and are a vulnerable fish.
Ocean sunfish The ocean sunfish is the heaviest bony fish. It can weigh up to 2,300 kg (5,100 lb). It is found in all warm and temperate oceans.[64]
King of herrings
 The
king of herrings is the longest bony fish. Its total length can reach 11 m (36 ft), and it can weigh up to 272 kilograms (600 lb). It is a rarely seen oarfish, found in all the world's oceans at depths of between 20 m (66 ft) and 1,000 m (3,300 ft).[65]
Mekong giant catfish The largest recorded freshwater fish is a
critically endangered
.

By breeding behavior

See also:
Spawning and Ichthyoplankton

In very deep waters, it is not easy for a fish to find a mate. There is no light, so some species depend on bioluminescence. Others are hermaphrodites, which doubles their chances of producing both eggs and sperm when an encounter does occur.[68]

Breeding Grouper Female
protogynous hermaphrodites, who school in harems
of three to fifteen females. When no male is available, the most aggressive and largest females change their sex to male.
Toadfish Male toadfish "sing" at up to 100 decibels with their swim bladders to attract mates.[69][70][71]
Anglerfish Female
gonads. This extreme sexual dimorphism ensures that, when the female is ready to spawn, she has a mate immediately available.[73]
Hammerheads Some sharks such as
fertilization
.

By brooding behavior

See also:
Spawning

Fish adopt a variety of strategies for nurturing their brood.

oviparous
, laying their eggs to hatch in the water.

Some animals, predominantly fish such as

mouthbrooding
, caring for their offspring by holding them in the mouth of a parent for extended periods of time. Mouthbrooding has evolved independently in several different families of fish.

Brooding Chain catshark The
oviparous
, laying its eggs to hatch in the water.
Great white shark The
ovoviviparous
, gestating eggs in the uterus for 11 months before giving birth.
Scalloped hammerhead The
viviparous
, bearing its young after nourishing hatchlings internally.
Cyphotilapia frontosa The female Cyphotilapia frontosa mouthbroods its fry. The fry can be seen looking out of her mouth.
Seahorses Seahorse males practice pouch-brooding similar to kangaroos. When seahorses mate, the female deposits her eggs into a special pouch on the male's belly. The pouch seals shut while he nurtures the developing eggs. Once the eggs hatch, the pouch opens and the male goes into labour.[78]

By feeding behaviour

External images
image icon Video of a slingjaw wrasse catching prey by protruding its jaw
image icon Video of a red bay snook catching prey by suction feeding
Further information: Aquatic feeding mechanisms

There are three basic methods by which food is gathered into the mouths of fish: by

ram feeding, and by manipulation or biting.[79] Nearly all fish species use one of these styles, and most use two.[80]

Early fish lineages had inflexible jaws limited to little more than opening and closing. Modern teleosts have evolved protusible jaws that can reach out to engulf prey.

slingjaw wrasse. Its mouth extends into a tube half as long as its body, which creates a strong suction to catch prey. The extended mouth tucks away under its body when not in use.[83][84]

In practice, feeding modes lie on a spectrum, with suction and ram feeding at the extremes. Many fish capture their prey using both suction pressure combined with a forward motion of the body or jaw.[85]

Most fish are food opportunists, or generalists. They eat whatever is most easily available.

ascidians, or crustaceans.[87][88] Ocean sunfish prefer jellyfish.[64]

Feeding Anglerfish
bioluminescent lure.[89]
Archerfish Archerfish prey on land-based insects and other small animals by shooting them down with water droplets from their specialized mouths. Archerfish are remarkably accurate; adults almost always hit the target on the first shot. They can bring down arthropods such as grasshoppers,[90] spiders and butterflies on a branch of an overhanging tree[91] 3 m (9.8 ft) above the water's surface.[92] This is partially due to good eyesight, but also due to their ability to compensate for light refraction when aiming.[93]
Triggerfish Triggerfish also use jets of water to uncover sand dollars buried in sand or overturn sea urchins.[94]
Silver arowana Other fish have developed extreme specializations. Silver arowana, also called monkey fish, can leap two meters out of the water to capture prey. They usually swim near the surface of the water waiting for potential prey. Their main diet consists of crustaceans, insects, smaller fishes and other animals that float on the water surface, for which its draw-bridge-like mouth is exclusively adapted for feeding. The remains of small birds, bats, and snakes have also been found in their stomachs.[95]
Cookiecutter shark The
cetaceans and larger fish, including other sharks. The cookiecutter attaches to its larger prey with its suctorial lips, and then protrudes its teeth to remove a symmetrical scoop of flesh.[96] Pictured is a pomfret
with bite wounds from a cookiecutter shark.
Striped bass Striped bass eat smaller fish.
Chinese algae eater
Chinese algae eaters are kept in aquaria
to control algae.
Emperor angelfish The Emperor angelfish feeds on coral sponges.
Herring Schooling herrings ram feed on copepods.
Mangrove jack
 The
mangrove jack eats crustaceans
.
Puffer fish
 Many
molluscs
.
Bucktoothed tetra
 The
molluscs
.
Cleaner fish These two small
wrasses are cleaner fish
, which eat parasites off other fish.
Cleaning station A reef manta ray at a cleaning station, maintaining a near stationary position atop a coral patch for several minutes while being cleaned by cleaner fishes.[97]
Doctor fish
Doctor fish nibbling on the diseased skin of patients. Doctor fish (nibble fish) live and breed in the outdoor pools of some Turkish spas, where they feed on the skin of patients with psoriasis. The fish are like cleaner fish
in that they only consume the affected and dead areas of the skin, leaving the healthy skin to recover.

By vision

Main article:
Vision in fishes

Many species of fish can see the

visible light.[98]

Mesopelagic fishes live in the deeper waters of the twilight zone, down to depths of 1000 metres, where the amount of sunlight available is not sufficient to support photosynthesis
. These fish are adapted for an active life under low light conditions.

Vision Four-eyed fish The four-eyed fish feeds at the surface of the water with eyes that allow it to see above and below the surface at the same time. Four-eyed fish have two specially-adapted eyes which are raised above the top of their head. The eyes are divided in two different parts, and the fish floats at the water surface with only the lower half of each eye underwater. The two halves are divided by a band of tissue and the eye has two pupils, connected by part of the iris. The upper half of the eye is adapted for vision in air, while the lower half is adapted for vision in water.[99] The lens of the eye also changes in thickness top to bottom to account for the difference in the refractive indices of air versus water. Their diet mostly consists of the terrestrial insects which are available at the surface, where they spend most of their time.[100]
Two stripe damselfish The
Predatory species cannot see this if their vision is not sensitive to ultraviolet. There is further evidence for this view that some fish use ultraviolet as a "high-fidelity secret communication channel hidden from predators", while yet other species use ultraviolet to make social or sexual signals.[102][103]
Barreleye
siphonophores from which it is believed the barreleye steals food.[104][105][106]
Flashlight fish Flashlight fish use a retroreflector behind the retina and photophores to detect eyeshine in other fish.[107][108][109]

By shape

Boxfishes have heavily armoured plate-like scales fused into a solid, triangular, boxlike carapace, from which the fins, tail, eyes and mouth protrude. Because of this heavy armour, boxfish move slowly, but few other fish are able to eat the adults.[110]

  • The humpback turretfish is a boxfish with an armoured triangular shaped body
    The
    boxfish
    with an armoured triangular shaped body
  • The leafy sea dragon is camouflaged to look like floating seaweed
    The
    leafy sea dragon
    is camouflaged to look like floating seaweed

By locomotion

See also: Fish locomotion

A number of species jump while swimming near the surface, skimming the water. Other species walk along the bottom on their fins.

Loco-
motion 		Dwarf seahorse The slowest-moving fishes are the
sea horses. The slowest of these, the tiny dwarf seahorse, has a sprint speed of one inch per minute.[111]
Atlantic bluefin tuna The Atlantic bluefin tuna is capable of sustained high speed cruising, and maintains high muscle temperatures so it can cruise in relatively cold waters.
Indo-Pacific sailfish

Among the fastesr sprinters are the Indo-Pacific sailfish (left) and the black marlin (right). Both have been recorded in a burst at over 110 kilometres per hour (68 mph). For the sailfish, that is equivalent to 12 to 15 times their own length per second.

Shortfin mako
External videos
video icon What makes mako sharks fast? National Geographic

The

exothermic constitution partly accounts for its relatively great speed.[113]

Wahoo The wahoo is perhaps the fastest fish for its size, attaining a speed of 19 lengths per second, reaching 78 kilometres per hour (48 mph).
Flying fish

Flying fish have unusually large pectoral fins, which enable the fish to take short gliding flights above the surface of the water in order to escape from predators. Their glides are typically around 50 meters (160 ft), but they can use updrafts at the leading edge of waves to cover distances of at least 400 meters (1,300 ft).[114] In May 2008, a flying fish was filmed off the coast of Japan (see video). The fish spent 45 seconds aloft, and was able to stay aloft by occasionally beating the surface of the water with its caudal (tail) fin.[115] The previous record was 42 seconds.[115]

Climbing perches
labyrinth organ
, a structure in the fish's head which allows it to breathe atmospheric oxygen. Their method of terrestrial locomotion uses the gill plates as supports, and the fish pushes itself using its fins and tail.
Mudskipper
External videos
video icon Mudskippers: The Fish That Walk on Land BBC Earth

The

lateral undulation, and tripod-like walking. The mudskipper is able to spend days moving about out of water and can even climb mangroves, although to only modest heights.[116] There are some species of fish that can "walk" along the sea floor but not on land. One such animal is the flying gurnard
.

Handfish
External videos
video icon Fish that walk CSIRO

The

pectoral fins
, which look like hands.

Tripod fish
External videos
video icon Fish that perch Okeanos Explorer ROV

caudal fin
, which act like "stilts", to perch and walk on the seafloor.

By toxicity

Toxic fish produce strong poisons in their bodies. Both poisonous fish and venomous fish contain toxins, but deliver them differently.
  • Venomous fish bite, sting, or stab, causing an envenomation. Venomous fish don't necessarily cause poisoning if they are eaten, since the digestive system often destroys the venom.[117]
  • By contrast, poisonous fish do not bite, sting, or stab to deliver their toxins, but they are poisonous to eat because they contain toxins in their body that the digestive system does not destroy.[117]

A 2006 study found that there are at least 1200 species of venomous fish.[118] There are more venomous fish than venomous snakes. In fact, there are more venomous fish than the combined total of all other venomous vertebrates.[118] Venomous fish are found in almost all habitats around the world, but mostly in tropical waters. They wound over 50,000 people every year.[119]

Venomous fish carry their venom in venom glands and use various delivery systems, such as spines, sharp fins, barbs, spikes or fangs. Venomous fish tend to be either very visible, using flamboyant colors to warn enemies, or skilfully camouflaged and may be buried in the sand. Apart from the defense or hunting value, venom helps bottom-dwelling fish by killing the bacteria that tries to invade their skin. Few of these venoms have been studied. They are a yet-to-be-tapped resource for bioprospecting to find drugs with medical uses.[120]

Treatment for venom stings usually includes the application of heat, using water at temperatures of about 45 °C (113 °F), since heat breaks down most complex venom proteins.

Toxicity
Puffer fish
 The
blowfish, to create a delicacy called "fugu", including just enough toxin for that "special flavour".[citation needed
]
Spotted trunkfish The
nurse sharks can die from eating a trunkfish.[121]
Giant moray The
harmful algal blooms, could be a reason why Polynesians migrated to Easter Island, New Zealand, and possibly Hawaii.[124][125]
Reef stonefish
 The most venomous known fish is the
reef stonefish.[126][127] It has a remarkable ability to camouflage itself amongst rocks. It is an ambush predator that sits on the bottom waiting for prey to approach. Instead of swimming away if disturbed, it erects the 13 venomous spines along its back. For defense, it can shoot venom from each or all of these spines. Each spine is like a hypodermic needle, delivering the venom from two sacs attached to the spine. The stonefish has control over whether to shoot its venom, and does so when provoked or frightened.[120] The venom results in severe pain, paralysis and tissue death, and can be fatal if not treated. Despite its formidable defenses, stonefish have predators. Some bottom feeding rays and sharks with crushing teeth feed on them, as does the Stokes's sea snake.[128]
Lionfish
 Head on view of the
lionfish can release venom only if something strikes its spines. Although not native to the U.S. coast, lionfish have appeared around Florida and have spread up the coast to New York. They are attractive aquarium fish, sometimes used to stock ponds, and may have been washed into the sea during a hurricane. Lionfish can aggressively dart at scuba divers and attempt to puncture their facemask with their venomous spines.[120]
Stargazer The stargazer, Uranoscopus sulphureus.[130] The stargazer buries itself and can deliver electric shocks as well as venom. It is a delicacy in some cultures (cooking destroys the venom), and can be found for sale in some fish markets with the electric organ removed. They have been called "the meanest things in creation".[120]
Stingray Stingrays can sting with their stinger (pictured). Such envenomations can occur to people who wade in shallow water and tread on them. This can be avoided by shuffling through the sand or stamping on the bottom, as the rays detect this and swim away. The stinger usually breaks off in the wound. It is barbed, so it can easily penetrate but cannot be easily removed. The stinger causes local trauma from the cut itself, pain and swelling from the venom, and possible later infection from bacteria. Occasionally, severed arteries or death can result.[131]

By human use

Predator fish size up schooling forage fish

Fish are sought after by humans for their value as commercial

scuba divers
.

Throughout human history, important

North Atlantic and the North Sea. Likewise, important traditional for anchovy and sardine fisheries have operated in the Pacific, the Mediterranean, and the southeast Atlantic.[133]
The world annual catch of forage fish in recent years has been around 25 million tonnes, or one quarter of the world's total catch.

Higher in the food chain,

Grand Banks.[134] Declining numbers led to international "cod wars" and eventually the virtual abandonment of these fisheries. In modern times, the Alaska pollock supports an important fishery in the Bering Sea and the north Pacific, yielding about 6 million tonnes, while cod amounts to about 9 million tonnes.[133]

porgie, shad, mahi-mahi, smelt whiting, swordfish, and walleye
.

aquarium fish
.

Snorkeling and scuba diving attracts millions of people to beaches, coral reefs, lakes, and other bodies of water to view fish and other marine life.

Human
use 		Yellowfin tuna Yellowfin tuna are now being fished as a replacement for the depleted southern bluefin tuna.
Anchovy These schooling anchovy are forage fish.
Atlantic cod Atlantic cod fisheries have collapsed.
Alaska pollock The Alaska pollock has been described as "the largest remaining source of palatable fish in the world".[137]
Koi Koi (and goldfish) have been kept in decorative ponds for centuries in China and Japan.

By vulnerability

  • resilience

Other

Other Bony-eared assfish Fish hold the records for the relative brain weights of vertebrates. Most vertebrate species have similar brain-to-body weight ratios. The deep sea bathypelagic bony-eared assfish[138] has the smallest ratio of all known vertebrates.[139]
Elephantnose fish
 At the other extreme, the
elephantnose fish, an African freshwater fish, has an exceptionally large brain-to-body weight ratio. These fish have the largest brain-to-body oxygen consumption ratio of all known vertebrates.[140]
Hallucinogenic fish The
Reunion Island, and Mulloidichthys samoensis,[143] called "the chief of ghosts" in Hawaii.[144]
Nopoli rockclimbing goby The Nopoli rockclimbing goby uses its mouth to climb waterfalls by inching up rocks like a caterpillar, using its mouth as a sucker together with another sucker on its stomach. When the fish is young, it undergoes a radical transformation when it moves from saltwater to a freshwater stream. The mouth migrates over a period of two days from the front of its head to its chin. This allows the fish to feed by scraping algae from rocks. Pictured is the goby before and after the transformation.[145][146]
Vampire fish Smaller species of vampire fish, native to the Amazon River, have an alleged tendency to burrow into and parasitise the human urethra. However, despite ethnological reports dating back to the late 19th century, the first documented case of the removal of a vampire fish from a human urethra did not occur until 1997, and even that incident has remained a matter of controversy.[147]

See also

Notes

  1. ^ Froese, Rainer; Pauly, Daniel (eds.) (2009). "Phycodurus eques" in FishBase. July 2009 version.
  2. ^ Leafy and Weedy Sea Dragon National Geographic Profile. Retrieved 20 July 2009.
  3. doi:10.2305/IUCN.UK.2017-2.RLTS.T17096A67622420.en
    . Retrieved 12 November 2021.
  4. S2CID 84572467
    .
  5. ^ FishBase
  6. ^ Estudo das Espécies Ícticas do Parque Estadual do Cantão, fish species survey of Cantão (in Portuguese)
  7. ^ Moyle & Cech 2003, p. Chapter 1
  8. ISBN 978-0-471-25031-9
    .
  9. ^ Helfman et al. 2009, p. 3
  10. ^ Tree of life web project - Chordates.
  11. N. A. Campbell and J. B. Reece
    (2005). Biology Seventh Edition. Benjamin Cummings, San Francisco CA.
  12. ^ Clack, J. A. (2002) Gaining Ground. Indiana University
  13. ^ Teleost Encyclopædia Britannica Online. 15 July 2009
  14. ^
    ISBN 978-0-412-54010-3
    .
  15. ^ Ben Waggoner (17 July 1995). "Telostei". Museum of Paleontology, University of California, Berkeley. Retrieved 8 June 2006.
  16. ^ a b c d Cohen, DM (1970). "How many recent fishes are there?". Proceedings of the California Academy of Sciences. 38 (17): 341–346.
  17. ^ a b Bone & Moore 2008, p. 3
  18. ^ CIA Factbook: World.
  19. ^ Elert, Glenn Volume of Earth's Oceans. The Physics Factbook. Retrieved 19 April 2008.
  20. ^ Wilson RW, Millero FJ, Taylor JR, Walsh PJ, Christensen V, Jennings S and Grosell M (2009) "Contribution of Fish to the Marine Inorganic Carbon Cycle" Science, 323 (5912) 359-362. (This article contains the first ever estimate of global fish biomass)
  21. ^ Shiklomanov, I A, (1993) World fresh water resources in Glick, P H, ed., Water in Crisis: Oxford University Press, p 13-24.
  22. ^ Horn, MH (1972). "The amount of space available for marine and freshwater fishes" (PDF). NOAA: Fishery Bulletin. 70: 1295–1297.
  23. ^ a b c d e Bone & Moore 2008, p. 35
  24. ^ C.Michael Hogan. 2011. Ross Sea. Eds. P.Saundry & C.J.Cleveland. Encyclopedia of Earth. National Council for Science and the Environment. Washington DC
  25. ^ Froese, Rainer; Pauly, Daniel (eds.) (2009). "Abyssobrotula galatheae" in FishBase. July 2009 version.
  26. ^ Jamieson, A.J., and Yancey, P. H. (2012). On the Validity of the Trieste Flatfish: Dispelling the Myth. The Biological Bulletin 222(3): 171-175
  27. ^ Yanceya, P.H.; Gerringera, E.M.; Drazen, J.C.; Rowden, A.A.; and Jamieson, A. (2014). Marine fish may be biochemically constrained from inhabiting the deepest ocean depths. PNAS 111(12): 4461–4465
  28. ^ Froese, Rainer; Pauly, Daniel (eds.) (2009). "Triplophysa stoliczkai" in FishBase. July 2009 version.
  29. ^ Froese, Rainer; Pauly, Daniel (eds.) (2014). "Astyanax mexicanus" in FishBase. 2014 version.
  30. ^ Froese, Rainer; Pauly, Daniel (eds.) (2009). "Thermichthys hollisi" in FishBase. July 2009 version.
  31. ^ Antennariidae: Frogfishes Tree of Life Web Project
  32. ^ Biological profiles: Sargassumfish Florida Museum of Natural History. Retrieved 2012-01-04.
  33. S2CID 22684907
    .
  34. PMID 14999022
    .
  35. ^ Froese, Rainer; Pauly, Daniel (eds.) (2009). "Cynolebias nigripinnis" in FishBase. July 2009 version.
  36. ^ Munk, K. (2001) "Maximum Ages of Groundfishes in Waters off Alaska and British Columbia and Considerations of Age Determination". Alaska Fishery Research Bulletin 8 :1.
  37. ^ Cailliet, G.M., Andrews, A.H., Burton, E.J., Watters, D.L., Kline, D.E., Ferry-Graham, L.A. (2001) "Age determination and validation studies of marine fishes: do deep-dwellers live longer?" Exp. Gerontol. 36 : 739–764.
  38. ^ Froese, Rainer; Pauly, Daniel (eds.) (2009). "Sebastes aleutianus" in FishBase. July 2009 version.
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References

External links
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