Euctenochasmatia

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Euctenochasmatians
Temporal range:
Ma
Cast of the sub-adult
Pterodactylus antiquus
Cast of fossil specimen of the
Pterodaustro guinazui
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Infraorder: Archaeopterodactyloidea
Clade: Euctenochasmatia
Unwin, 2003
Subgroups

Euctenochasmatia is an extinct group of

pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.[1]

Euctenochasmatians were specialized pterosaurs that had elongated necks as well as specialized teeth. A peculiar family within this group is the Ctenochasmatidae, which most of the members had very distinguishing teeth that were lined within their elongated snouts.[2] A genus called Pterofiltrus only had 112 teeth, but these teeth cover about 55.8% of the total skull, and the skull itself measured about 208 millimeters (8.2 in) in length.[3]

Description

Restored skull of Pterodaustro

Euctenochasmatians had very distinctive features in comparison to other pterosaurs, including the shape of their jaws, as well as their highly specialized teeth. These teeth are thought to have been used for

crustaceans, plankton, algae, and other small creatures from the water. The teeth of Pterodaustro are unique within pterosaurs, and no other discovered genera had this type of teeth.[4][5]

Other members of this group, such as the

ctenochasmatid Feilongus also had its teeth confined within its jaw tips, as well as having crests on the rear portion of the skull and jaws, but differed Feilongus from the gallodactylids by having a possible pronounced overbite, and 76 teeth, which were needle-like.[7]

One of the largest toothed pterosaurs was Moganopterus, it was, yet again, a ctenochasmatid, and was similar in build to Feilongus. What made Moganopterus distinct was its size; while Feilongus had a wingspan of about 2.4 meters (7.9 ft), Moganopterus had an impressive wingspan of more than 7 meters (23 ft), making it more than three times larger than Feilongus.[8]

Classification

Researchers such as David Unwin, have traditionally defined the dubious family Pterodactylidae in such a way to ensure it is nested within the clade Ctenochasmatoidea. In 2003, Unwin defined the same clade (Pterodactylus + Pterodaustro), but erected the name Euctenochasmatia instead of Pterodactylidae for his conclusion.[9] Unwin had considered Euctenochasmatia to be a subgroup within Ctenochasmatoidea, similar to his former conclusion of Pterodactylidae, but most analyses have since found the genus Pterodactylus to be more primitive than previously thought, making the clade Euctenochasmatia the more inclusive group containing both Pterodactylus and Ctenochasmatoidea.[10]

Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set (Andres, 2021). Andres' analysis found Pterodactylus to be a close relative of the ctenochasmatoids.[11]

 Pterodactyloidea 

Kryptodrakon

Painten pro-pterodactyloid

Lophocratia

Eupterodactyloidea

Archaeopterodactyloidea
Germanodactylidae

Cuspicephalus scarfi

Germanodactylus cristatus

Altmuehlopterus rhamphastinus

Euctenochasmatia

Pterodactylus antiquus

Ctenochasmatoidea
Gallodactylidae

Cycnorhamphus suevicus

Normannognathus wellnhoferi

Ctenochasmatidae

Cycnorhamphus suevicus

Earlier, in 2017, Steven Vidovic and David Martill had recovered a significantly different set of relationships for early

Lophocratia.[1]

 Pterodactyloidea 

Eosipterus yangi

Archaeopterodactyloidea

Pterodactylus antiquus

Lophocratia

Eupterodactyloidea

Euctenochasmatia

Diopecephalus kochi

Ctenochasmatoidea

Aurorazhdarchia

Ctenochasmatidae

References

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  4. PMID 18308672.{{cite journal}}: CS1 maint: multiple names: authors list (link
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  11. ^ Andres, B. (2021) Phylogenetic systematics of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea). Journal of Vertebrate Paleontology, 41:sup1, 203-217. DOI: 10.1080/02724634.2020.1801703 https://www.tandfonline.com/doi/full/10.1080/02724634.2020.1801703