Eudromaeosauria
Eudromaeosaurians | |
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Eudromaeosauria diversity, featuring from top left to lower right: Utahraptor, Deinonychus, Velociraptor and Bambiraptor | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | †Dromaeosauridae |
Clade: | †Eudromaeosauria Longrich & Currie, 2009 |
Subgroups | |
Eudromaeosauria ("true dromaeosaurs") is a subgroup of terrestrial
Eudromaeosaur fossils are known almost exclusively from the northern hemisphere. They first appeared in the early Cretaceous Period (early
Description
While other dromaeosaurids filled a variety of specialized ecological niches, mainly those of small predators or larger fish-eating forms, eudromaeosaurs functioned as large-bodied predators of often medium- to large-sized prey. Aside from their generally larger size, eudromaeosaurs are characterized by several features of the foot. First, differences existed in the positions of the grooves that anchored blood vessels and keratin sheathes of the toe claws. In primitive dromaeosaurids like Hesperonychus, these grooves ran parallel to each other on either side of the claw along its length. In eudromaeosaurs, the grooves were asymmetrical, with the inner one split into two distinct grooves and elevated toward the top of the claw, while the single outer groove remained positioned at the midline.[4]
The second distinguishing characteristic of eudromaeosaurs is an expanded and enlarged "heel" on the last bone in the second toe (phalanx), which bore the enlarged, sickle-like toe claw. Finally, the first bone of the second toe also possessed an enlarged expansion at the joint, another adaptation relating to the unusually enlarged claw, and which helped the animal hold the claw high off the ground. Also unlike their more basal relatives, the sickle claw of eudromaeosaurs was sharper and more blade-like. In
Classification
Eudromaeosauria was first defined as a node-based clade by Nick Longrich and Philip J. Currie in 2009, as the most inclusive natural group containing Dromaeosaurus, Velociraptor, Deinonychus, and Saurornitholestes, their most recent common ancestor and all of its other descendants. The various "subfamilies" have also been redefined as clades, usually defined as all species closer to the group's namesake than to Dromaeosaurus or any namesakes of other subclades.[4]
The subgroups of Eudromaeosauria frequently shift in content based on new analysis, but typically consist of the following groups. For example, the subfamily Velociraptorinae has traditionally included
Relationships
The below cladogram follows an analysis by Evans et al. in 2013. Their analysis used an updated version of the dataset originally compiled by Nick Longrich and Phil Currie to study dromaeosaurid relationships, and found a relatively traditional arrangement of eudromaeosaurian relationships.[6]
Eudromaeosauria |
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See also
References
- PMID 21124919.
- .
- .
- ^ PMID 19289829.
- S2CID 31096081. Archived from the original(PDF) on 2009-03-26. Retrieved 2007-03-29.
- S2CID 14978813.