Extinction
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Extinct |
Threatened |
Lower Risk |
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Evolutionary biology |
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Extinction is the termination of a
More than 99% of all
Through evolution, species arise through the process of speciation—where new varieties of organisms arise and thrive when they are able to find and exploit an ecological niche—and species become extinct when they are no longer able to survive in changing conditions or against superior competition. The relationship between animals and their ecological niches has been firmly established.[8] A typical species becomes extinct within 10 million years of its first appearance,[5] although some species, called living fossils, survive with little to no morphological change for hundreds of millions of years.
Mass extinctions are relatively rare events; however, isolated extinctions of species and clades are quite common, and are a natural part of the evolutionary process.[9] Only recently have extinctions been recorded and scientists have become alarmed at the current high rate of extinctions.[10][11][12][13][14] Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100.[15] A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during the human era since the Late Pleistocene would require 5 to 7 million years to recover.[16]
According to the 2019
In June 2019, one million species of plants and animals were at risk of extinction. At least 571 plant species have been lost since 1750, but likely many more. The main cause of the extinctions is the destruction of natural habitats by human activities, such as cutting down forests and converting land into fields for farming.[21]
A dagger symbol (†) placed next to the name of a species or other taxon normally indicates its status as extinct.
Examples
Examples of species and subspecies that are extinct include:
- Steller's sea cow (the last known member died circa 1768)
- Dodo (the last confirmed sighting was in 1662)
- Chinese paddlefish (last seen in 2003; declared extinct in 2022)
- Great auk (last confirmed pair was killed in the 1840s)
- Thylacine (the last thylacine killed in the wild was shot in 1930; the last captive tiger lived in Hobart Zoo until 1936)
- Kauai O'o (last known member was heard in 1987; the entire Mohoidae family became extinct with it)
- Spectacled cormorant (last known members were said to live in the 1850s)
- Incasdied in captivity in 1918; declared extinct in 1939)
- Passenger pigeon (last known member named Martha died in captivity in 1914)
- Tasmanian emu (the last claimed sighting of the emu was in 1839)
- Japanese Sea Lion(the last confirmed record was a juvenile specimen captured in 1974)
- Schomburgk's deer (became extinct in the wild in 1932; the last captive deer was killed in 1938)
- Natura Artis Magistrain 1883)
Definition
A species is extinct when the last existing member dies. Extinction therefore becomes a certainty when there are no surviving individuals that can reproduce and create a new generation. A species may become functionally extinct when only a handful of individuals survive, which cannot reproduce due to poor health, age, sparse distribution over a large range, a lack of individuals of both sexes (in sexually reproducing species), or other reasons.
Pinpointing the extinction (or
In
Currently, an important aspect of extinction is human attempts to preserve critically endangered species. These are reflected by the creation of the
The extinction of one species' wild population can have knock-on effects, causing further extinctions. These are also called "chains of extinction".[27] This is especially common with extinction of keystone species.
A 2018 study indicated that the
Pseudoextinction
Extinction of a parent species where daughter species or subspecies are still extant is called pseudoextinction or phyletic extinction. Effectively, the old taxon vanishes, transformed (anagenesis) into a successor,[29] or split into more than one (cladogenesis).[30]
Pseudoextinction is difficult to demonstrate unless one has a strong chain of evidence linking a living species to members of a pre-existing species. For example, it is sometimes claimed that the extinct Hyracotherium, which was an early horse that shares a common ancestor with the modern horse, is pseudoextinct, rather than extinct, because there are several extant species of Equus, including zebra and donkey; however, as fossil species typically leave no genetic material behind, one cannot say whether Hyracotherium evolved into more modern horse species or merely evolved from a common ancestor with modern horses. Pseudoextinction is much easier to demonstrate for larger taxonomic groups.
Lazarus taxa
A Lazarus taxon or Lazarus species refers to instances where a species or taxon was thought to be extinct, but was later rediscovered. It can also refer to instances where large gaps in the fossil record of a taxon result in fossils reappearing much later, although the taxon may have ultimately become extinct at a later point.
The
Attenborough's long-beaked echidna (Zaglossus attenboroughi) is an example of a Lazarus species from Papua New Guinea that had last been sighted in 1962 and believed to be possibly extinct, until it was recorded again in November 2023.[33]
Some species currently thought to be extinct have had continued speculation that they may still exist, and in the event of rediscovery would be considered Lazarus species. Examples include the
Causes
As long as species have been evolving, species have been going extinct. It is estimated that over 99.9% of all species that ever lived are extinct. The average lifespan of a species is 1–10 million years,[35] although this varies widely between taxa. A variety of causes can contribute directly or indirectly to the extinction of a species or group of species. "Just as each species is unique", write Beverly and
Assessing the relative importance of genetic factors compared to environmental ones as the causes of extinction has been compared to the debate on
Human-driven extinction started as humans migrated out of Africa more than 60,000 years ago.
Genetics and demographic phenomena
If adaptation increasing population fitness is slower than environmental degradation plus the accumulation of slightly deleterious mutations, then a population will go extinct.[44] Smaller populations have fewer beneficial mutations entering the population each generation, slowing adaptation. It is also easier for slightly deleterious mutations to fix in small populations; the resulting positive feedback loop between small population size and low fitness can cause mutational meltdown.
Limited geographic range is the most important determinant of genus extinction at background rates but becomes increasingly irrelevant as mass extinction arises.[45] Limited geographic range is a cause both of small population size and of greater vulnerability to local environmental catastrophes.
Extinction rates can be affected not just by population size, but by any factor that affects
Genetic pollution
Extinction sometimes results for species evolved to specific ecologies
The gene pool of a
Habitat degradation
Habitat degradation is currently the main anthropogenic cause of species extinctions. The main cause of habitat degradation worldwide is agriculture, with
Habitat destruction, particularly the removal of vegetation that stabilizes soil, enhances erosion and diminishes nutrient availability in terrestrial ecosystems. This degradation can lead to a reduction in agricultural productivity. Furthermore, increased erosion contributes to poorer water quality by elevating the levels of sediment and pollutants in rivers and streams.[53]
Habitat degradation through toxicity can kill off a species very rapidly, by killing all living members through contamination or sterilizing them. It can also occur over longer periods at lower toxicity levels by affecting life span, reproductive capacity, or competitiveness.
Habitat degradation can also take the form of a physical destruction of niche habitats. The widespread destruction of tropical rainforests and replacement with open pastureland is widely cited as an example of this;[15] elimination of the dense forest eliminated the infrastructure needed by many species to survive. For example, a fern that depends on dense shade for protection from direct sunlight can no longer survive without forest to shelter it. Another example is the destruction of ocean floors by bottom trawling.[54]
Diminished resources or introduction of new competitor species also often accompany habitat degradation.
Predation, competition, and disease
In the natural course of events, species become extinct for a number of reasons, including but not limited to: extinction of a necessary host, prey or pollinator,
Coextinction
Coextinction refers to the loss of a species due to the extinction of another; for example, the extinction of
Climate change
Extinction as a result of
Sexual selection and male investment
Studies of fossils following species from the time they evolved to their extinction show that species with high sexual dimorphism, especially characteristics in males that are used to compete for mating, are at a higher risk of extinction and die out faster than less sexually dimorphic species, the least sexually dimorphic species surviving for millions of years while the most sexually dimorphic species die out within mere thousands of years. Earlier studies based on counting the number of currently living species in modern taxa have shown a higher number of species in more sexually dimorphic taxa which have been interpreted as higher survival in taxa with more sexual selection, but such studies of modern species only measure indirect effects of extinction and are subject to error sources such as dying and doomed taxa speciating more due to splitting of habitat ranges into more small isolated groups during the habitat retreat of taxa approaching extinction. Possible causes of the higher extinction risk in species with more sexual selection shown by the comprehensive fossil studies that rule out such error sources include expensive sexually selected ornaments having negative effects on the ability to survive natural selection, as well as sexual selection removing a diversity of genes that under current ecological conditions are neutral for natural selection but some of which may be important for surviving climate change.[63]
Mass extinctions
There have been at least five mass extinctions in the history of life on earth, and four in the last 350 million years in which many species have disappeared in a relatively short period of geological time. A massive eruptive event that released large quantities of tephra particles into the atmosphere is considered to be one likely cause of the "Permian–Triassic extinction event" about 250 million years ago,[64] which is estimated to have killed 90% of species then existing.[65] There is also evidence to suggest that this event was preceded by another mass extinction, known as Olson's Extinction.[64] The Cretaceous–Paleogene extinction event (K–Pg) occurred 66 million years ago, at the end of the Cretaceous period; it is best known for having wiped out non-avian dinosaurs, among many other species.
Modern extinctions
According to a 1998 survey of 400 biologists conducted by
Biologist
In January 2020, the UN's Convention on Biological Diversity drafted a plan to mitigate the contemporary extinction crisis by establishing a deadline of 2030 to protect 30% of the Earth's land and oceans and reduce pollution by 50%, with the goal of allowing for the restoration of ecosystems by 2050.[78][79] The 2020 United Nations' Global Biodiversity Outlook report stated that of the 20 biodiversity goals laid out by the Aichi Biodiversity Targets in 2010, only 6 were "partially achieved" by the deadline of 2020.[80] The report warned that biodiversity will continue to decline if the status quo is not changed, in particular the "currently unsustainable patterns of production and consumption, population growth and technological developments".[81] In a 2021 report published in the journal Frontiers in Conservation Science, some top scientists asserted that even if the Aichi Biodiversity Targets set for 2020 had been achieved, it would not have resulted in a significant mitigation of biodiversity loss. They added that failure of the global community to reach these targets is hardly surprising given that biodiversity loss is "nowhere close to the top of any country's priorities, trailing far behind other concerns such as employment, healthcare, economic growth, or currency stability."[82][83]
History of scientific understanding
For much of history, the modern understanding of extinction as the end of a species was incompatible with the prevailing worldview. Prior to the 19th century, much of Western society adhered to the belief that the world was created by God and as such was complete and perfect.[86] This concept reached its heyday in the 1700s with the peak popularity of a theological concept called the great chain of being, in which all life on earth, from the tiniest microorganism to God, is linked in a continuous chain.[87] The extinction of a species was impossible under this model, as it would create gaps or missing links in the chain and destroy the natural order.[86][87] Thomas Jefferson was a firm supporter of the great chain of being and an opponent of extinction,[86][88] famously denying the extinction of the woolly mammoth on the grounds that nature never allows a race of animals to become extinct.[89]
A series of fossils were discovered in the late 17th century that appeared unlike any living species. As a result, the scientific community embarked on a voyage of creative rationalization, seeking to understand what had happened to these species within a framework that did not account for total extinction. In October 1686,
Cuvier's fossil evidence showed that very different life forms existed in the past than those that exist today, a fact that was accepted by most scientists.[86] The primary debate focused on whether this turnover caused by extinction was gradual or abrupt in nature.[93] Cuvier understood extinction to be the result of cataclysmic events that wipe out huge numbers of species, as opposed to the gradual decline of a species over time.[94] His catastrophic view of the nature of extinction garnered him many opponents in the newly emerging school of uniformitarianism.[94]
Jean-Baptiste Lamarck, a gradualist and colleague of Cuvier, saw the fossils of different life forms as evidence of the mutable character of species.[93] While Lamarck did not deny the possibility of extinction, he believed that it was exceptional and rare and that most of the change in species over time was due to gradual change.[93] Unlike Cuvier, Lamarck was skeptical that catastrophic events of a scale large enough to cause total extinction were possible. In his geological history of the earth titled Hydrogeologie, Lamarck instead argued that the surface of the earth was shaped by gradual erosion and deposition by water, and that species changed over time in response to the changing environment.[93][95]
The concept of extinction was integral to Charles Darwin's On the Origin of Species, with less fit lineages disappearing over time. For Darwin, extinction was a constant side effect of competition.[96] Because of the wide reach of On the Origin of Species, it was widely accepted that extinction occurred gradually and evenly (a concept now referred to as background extinction).[89] It was not until 1982, when David Raup and Jack Sepkoski published their seminal paper on mass extinctions, that Cuvier was vindicated and catastrophic extinction was accepted as an important mechanism[citation needed]. The current understanding of extinction is a synthesis of the cataclysmic extinction events proposed by Cuvier, and the background extinction events proposed by Lyell and Darwin.
Human attitudes and interests
Extinction is an important research topic in the field of
Biologist Bruce Walsh states three reasons for scientific interest in the preservation of species: genetic resources, ecosystem stability, and ethics; and today the scientific community "stress[es] the importance" of maintaining biodiversity.[100][101]
In modern times, commercial and industrial interests often have to contend with the effects of production on plant and animal life. However, some technologies with minimal, or no, proven harmful effects on
Governments sometimes see the loss of native species as a loss to
People who live close to nature can be dependent on the survival of all the species in their environment, leaving them highly exposed to extinction risks. However, people prioritize day-to-day survival over species conservation; with human overpopulation in tropical developing countries, there has been enormous pressure on forests due to subsistence agriculture, including slash-and-burn agricultural techniques that can reduce endangered species's habitats.[108]
Planned extinction
Completed
- The smallpox virus is now extinct in the wild,[111] although samples are retained in laboratory settings.
- The infected domestic cattle, is now extinct in the wild.[112]
Proposed
Disease agents
The poliovirus is now confined to small parts of the world due to extermination efforts.[113]
Dracunculus medinensis, or Guinea worm, a parasitic worm which causes the disease dracunculiasis, is now close to eradication thanks to efforts led by the Carter Center.[114]
Treponema pallidum pertenue, a bacterium which causes the disease yaws, is in the process of being eradicated.
Disease vectors
Biologist
Biologist E. O. Wilson has advocated the eradication of several species of mosquito, including malaria vector Anopheles gambiae. Wilson stated, "I'm talking about a very small number of species that have co-evolved with us and are preying on humans, so it would certainly be acceptable to remove them. I believe it's just common sense."[116]
There have been many campaigns – some successful – to locally eradicate tsetse flies and their trypanosomes in areas, countries, and islands of Africa (including Príncipe).[117][118] There are currently serious efforts to do away with them all across Africa, and this is generally viewed as beneficial and morally necessary,[119] although not always.[120]
Cloning
Some, such as Harvard geneticist
In 2003, scientists tried to clone the extinct Pyrenean ibex (C. p. pyrenaica).
See also
- Bioevent
- Empty forest
- Endling
- Extinction: The Facts (2020 documentary)
- Genocide
- Habitat fragmentation
- Lists of extinct animals
- List of extinct birds
- Living Planet Index
- Our Final Hour
- Refugium (population biology)
- Sepilok Orang Utan Rehabilitation Centre
- The Sixth Extinction (2014 book)
- Voluntary Human Extinction Movement
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It is noteworthy that human concern about human extinction takes a different form from human concern (where there is any) about the extinction of non-human species. Most humans who are concerned about the extinction of non-human species are not concerned about the individual animals whose lives are cut short in the passage to extinction, even though that is one of the best reasons to be concerned about extinction (at least in its killing form). The popular concern about animal extinction is usually concern for humans—that we shall live in a world impoverished by the loss of one aspect of faunal diversity, that we shall no longer be able to behold or use that species of animal. In other words, none of the typical concerns about human extinction are applied to non-human species extinction.
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It's no wonder that we react so nonchalantly to the ever-mounting statistics about the crisis of mass extinction. We have a habit of taking this information with surprising calm. We don't weep. We don't get worked up. Why? Because we see humans as fundamentally separate from the rest of the living community. Those species are out there, in the environment. They aren't in here; they aren't part of us. It is not surprising that we behave this way. After all, this is the core principle of capitalism: that the world is not really alive, and it is certainly not our kin, but rather just stuff to be extracted and discarded – and that includes most of the human beings living here too. From its very first principles, capitalism has set itself at war against life itself.
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- S2CID 17608648.
- S2CID 67788418.
- ^ A. Zitner (24 December 2000). "Cloned Goat Would Revive Extinct Line". Los Angeles Times. Archived from the original on 25 August 2011. Retrieved 17 May 2010.
- ^ Nicholas Wade (19 November 2008). "Regenerating a Mammoth for $10 Million". The New York Times. Archived from the original on 12 March 2017. Retrieved 17 May 2010. The cell could be converted into an embryo and brought to term by an elephant, a project he estimated would cost some $10 million. "This is something that could work, though it will be tedious and expensive,"
- PMID 19167744.
- ^ Steve Connor (2 February 2009). "Cloned goat dies after attempt to bring species back from extinction". The Independent. London. Archived from the original on 13 October 2017. Retrieved 17 May 2010.
- ^ Gray, Richard; Dobson, Roger (31 January 2009). "Extinct ibex is resurrected by cloning". Telegraph. London. Archived from the original on 11 January 2022.
Further reading
- Day, David (1981). The Doomsday Book of Animals: A Natural History of Vanished Species. New York: Viking Press. ISBN 978-0670279876.
- PMID 35757873.
- Elizabeth Pennisi (9 March 2022). "Bringing back the woolly mammoth and other extinct creatures may be impossible". Science.
- Pelley, Scott (1 January 2023). "Scientists say planet in midst of sixth mass extinction, Earth's wildlife running out of places to live". CBS News.