Galanthus
Snowdrop | |
---|---|
Galanthus nivalis common snowdrop | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Order: | Asparagales |
Family: | Amaryllidaceae |
Subfamily: | Amaryllidoideae |
Tribe: | Galantheae |
Genus: | Galanthus L. |
Type species | |
Galanthus nivalis | |
Synonyms[2] | |
|
Galanthus (from
Snowdrops have been known since the earliest times under various names, but were named Galanthus in 1753. As the number of recognised species increased, various attempts were made to divide the species into subgroups, usually on the basis of the pattern of the emerging leaves (
Most species flower in winter, before the vernal equinox (20 or 21 March in the Northern Hemisphere), but some flower in early spring and late autumn. Sometimes snowdrops are confused with the two related genera within the tribe Galantheae, snowflakes Leucojum and Acis.
Description
General
All
Vegetative
- Leaves
These are basal, emerging from the bulb initially enclosed in a tubular membranous sheath of
The scape (flowering stalk) is erect, leafless, terete, or compressed.[7]
Reproductive
- Inflorescence
At the top of the
- Androecium
The six
- Gynoecium, fruit and seeds
The
The
Distribution and habitat
The genus Galanthus is native to Europe and the Middle East, from the Spanish and French
Galanthus nivalis is the best-known and most widespread representative of the genus Galanthus. It is native to a large area of Europe, stretching from the Pyrenees in the west, through France and Germany to Poland in the north, Italy, northern Greece, Bulgaria, Romania, Ukraine, and
Most Galanthus species grow best in
or mountain species.Taxonomy
History
Early
Snowdrops have been known since early times, being described by the classical Greek author Theophrastus, in the fourth century BCE, in his Περὶ φυτῶν ἱστορία (Latin: Historia plantarum, Enquiry into plants). He gave it, and similar plants, the name λευκόἲον (λευκος, leukos "white" and ἰόν, ion "violet") from which the later name Leucojum was derived. He described the plant as "ἑπεἰ τοῖς γε χρώμασι λευκἂ καἱ οὐ λεπυριώδη" (in colour white and bulbs without scales)[16] and of their habits "Ἰῶν δ' ἁνθῶν τὀ μἑν πρῶτον ἑκφαἱνεται τὁ λευκόἲον, ὅπου μἑν ό ἀἠρ μαλακώτερος εὐθὑς τοῦ χειμῶνος, ὅπου δἐ σκληρότερος ὕστερον, ἑνιαχοῡ τοῡ ἣρος" (Of the flowers, the first to appear is the white violet. Where the climate is mild, it appears with the first sign of winter, but in more severe climates, later in spring)[17]
Modern
In 1763
Phylogeny
Cladogram of evolutionary lines in Galanthus sensu Ronsted et al.[3] | |||||||||||||||
|
Galanthus is one of three closely related genera making up the tribe Galantheae within subfamily Amaryllidoideae (family Amaryllidaceae). Sometimes snowdrops are confused with the other two genera, Leucojum and Acis (both called snowflakes). Leucojum species are much larger and flower in spring (or early summer, depending on the species), with all six tepals in the flower being the same size, although some "poculiform" (goblet- or cup-shaped) Galanthus species may have inner segments similar in shape and length to the outer ones. Galantheae are likely to have arisen in the Caucusus.[39]
Subdivision
Galanthus has approximately 20 species, but new species continue to be described.
Many species are difficult to identify, however, and traditional infrageneric classification based on
- section Nivales Beck (flat leaves)
- section Plicati Beck (plicate leaves)
- section Latifolii Stern (convolute leaves)
Stern further utilised characteristics such as the markings of the inner segments, length of the pedicels in relation to the spathe, and the colour and shape of the leaves in identifying and classifying species
Traub considered them as
- subgenus Galanthus
- subgenus Plicatanthus Traub & Moldk.
- subgenus Platyphyllanthe Traub
By contrast Davis, with much more information and specimens, included biogeography in addition to vernation, forming two series. He used somewhat different terminology for vernation, namely applanate (flat), explicative (plicate), and supervolute (convolute). He merged Nivalis and Plicati into series Galanthus, and divided Latifolii into two subseries, Glaucaefolii (Kem.-Nath) A.P.Davis and Viridifolii (Kem.-Nath) A.P.Davis.[50]
Early
By contrast, another study performed at the same time, using both nuclear and chloroplast DNA, but limited to the 14 species found in Turkey, largely confirmed Davis' series and subseries, and with biogeographical correlation. Series Galanthus in this study corresponded to clade nivalis, subseries Glaucaefolii with clade Elwesii and subseries Viridifolii with clades Woronowii and Alpinus. However, the model did not provide complete resolution.[55]
Clades
Cladogram of evolutionary lines in Galanthus sensu Ronsted et al.[3] | |||||||||||||||||||||||||||||||||||||||
|
sensu Ronsted et al. 2013[3]
- Platyphyllus clade (Transcaucasus, NE Turkey)
- Galanthus krasnovii Khokhr. 1963
- Galanthus platyphyllus Traub & Moldenke 1948
- Galanthus panjutinii Zubov & A.P.Davis 2012
- Trojanus clade (NW Turkey)
- Galanthus trojanus A.P.Davis & Özhatay 2001
- Ikariae clade (Aegean Islands)
- Galanthus ikariae Baker 1893
- Elwesii clade (Turkey, Aegean Islands, SE Europe)
- Galanthus cilicicus Baker 1897
- Galanthus elwesii Hook.f. 1875 (2 variants)
- Galanthus gracilis Celak. 1891
- Galanthus peshmenii A.P.Davis & C.D.Brickell 1994
- Nivalis clade (Europe, NW Turkey)
- Galanthus nivalis L. 1753
- Galanthus plicatus M.Bieb. 1819 (2 subspecies)
- Galanthus reginae-olgae Orph. 1874 (2 subspecies)
- Woronowii clade (Caucasus, E. and NE Turkey, N. Iran)
- Galanthus fosteri Baker 1889
- Galanthus lagodechianus Kem.-Nath. 1947
- Galanthus rizehensis Stern 1956
- Galanthus woronowii Losinsk. 1935
- Alpinus clade (Caucasus, NE Turkey, N.Iran)
- Galanthus × allenii Baker 1891
- Galanthus angustifolius Koss 1951
- Galanthus alpinus Sosn. (2 variants) 1911
- Galanthus koenenianus Lobin 1993
- Galanthus transcaucasicus Fomin 1909
- Unplaced
- Galanthus bursanus Zubov, Konca & A.P.Davis 2019 (NW Turkey)
- Galanthus samothracicus Kit Tan & Biel 2014 (Greece)
Cladogram of evolutionary lines in Galanthus sensu Margoz et al.[55] | |||||||||||||||
|
- Selected species
- Common snowdrop, Galanthus nivalis, grows to around 7–15 cm tall, flowering between January and April in the northern temperate zone (January–May in the wild). Applanate vernation[12] Grown as ornamental.
- Crimean snowdrop, Galanthus plicatus, 30 cm tall, flowering January/March, white flowers, with broad leaves folded back at the edges (explicative vernation)
- Giant snowdrop, Galanthus elwesii, a native of the Levant, 23 cm tall, flowering January/February, with large flowers, the three inner segments of which often have a much larger and more conspicuous green blotch (or blotches) than the more common kinds; supervolute vernation. Grown as ornamental.[56]
- Galanthus reginae-olgae, from Greece and Sicily, is quite similar in appearance to G. nivalis, but flowers in autumn before the leaves appear. The leaves, which appear in the spring, have a characteristic white stripe on their upper side; applanate vernation
- G. reginae-olgae subsp. vernalis, from Sicily, northern Greece and the southern part of former Yugoslavia, blooms at the end of the winter with developed young leaves and is thus easily confused with G. nivalis.
Etymology
Galanthus is derived from the
Ecology
Snowdrops are
Conservation
Some snowdrop species are threatened in their wild
Cultivation
Galanthus species and cultivars are extremely popular as symbols of spring and are traded more than any other wild-source ornamental bulb genus. Millions of bulbs are exported annually from Turkey and Georgia.[3] For instance export quotas for 2016 for G. elwesii were 7 million for Turkey.[65] Quotas for G. worononowii were 5 million for Turkey and 15 million for Georgia.[66] These figures include both wild-taken and artificially propagated bulbs.
Snowdrop gardens
Celebrated as a sign of spring, snowdrops may form impressive carpets of white in areas where they are native or have been naturalised. These displays may attract large numbers of sightseers.
Cultivars
Numerous single- and double-flowered
Awards
As of July 2017[update], the following have gained the Royal Horticultural Society's Award of Garden Merit:[73]
- Galanthus 'Ailwyn' [74]
- Galanthus 'Atkinsii' [75]
- Galanthus 'Bertram Anderson' [76]
- Galanthus elwesii[77]
- Galanthus elwesii 'Comet' [78]
- Galanthus elwesii 'Godfrey Owen' [79]
- Galanthus elwesii 'Mrs Macnamara' [80]
- Galanthus elwesii var. monostictus[81]
- Galanthus 'John Gray' [82]
- Galanthus 'Lady Beatrix Stanley' [83]
- Galanthus 'Magnet' [84]
- Galanthus 'Merlin' [85]
- Galanthus nivalis[86]
- Galanthus nivalis f. pleniflorus 'Flore Pleno' [87]
- Galanthus nivalis 'Viridapice' [88]
- Galanthus plicatus[89]
- Galanthus plicatus 'Augustus' [90]
- Galanthus plicatus 'Diggory' [91]
- Galanthus plicatus 'Three Ships' [92]
- Galanthus reginae-olgae subsp. reginae-olgae [93]
- Galanthus 'S. Arnott' [94]
- Galanthus 'Spindlestone Surprise' [95]
- Galanthus 'Straffan' [96]
- Galanthus 'Trumps' [97]
- Galanthus woronowii[98]
Propagation
Propagation is by offset bulbs, either by careful division of clumps in full growth ("in the green"), or removed when the plants are dormant, immediately after the leaves have withered; or by seeds sown either when ripe, or in spring. Professional growers and keen amateurs also use such methods as "twin-scaling" to increase the stock of choice cultivars quickly.
Toxicity
Snowdrops contain an active lectin or agglutinin named GNA for Galanthus nivalis agglutinin.[99]
Medicinal use
In 1983, Andreas Plaitakis and Roger Duvoisin suggested that the mysterious magical herb,
In popular culture
Snowdrops figure prominently in art and literature,
- In the fairy-tale play The Twelve Months by Russian writer Samuil Marshak, a greedy queen decrees that a basket of gold coins shall be rewarded to anyone who can bring her galanthus flowers in the dead of winter. A young orphan girl is sent out during a snow storm by her cruel stepmother to find the spirits of the 12 months of the year, who take pity on her and not only save her from freezing to death, but also make it possible for her to gather the flowers even in winter. The Soviet traditionally animated film The Twelve Months (1956), Lenfilm film The Twelve Months (1972), and the anime film Twelve Months (1980) (Sekai meisaku dowa mori wa ikiteiru in Japan), are based on this fairy-tale play.
- "Snowdrops" was the nickname that the British people gave during the Second World War to the military police of the United States Army (who were stationed in the UK preparatory to the invasion of the continent) because they wore a white helmet, gloves, gaiters, and Sam Browne belt against their olive drab uniforms.
- In the German fairy tale, Snow White and the Seven Dwarfs, "Snowdrop" is used as an alternate name for the Princess Snow White.
- The short story The Snowdrop by Hans Christian Andersen follows the fate of a snowdrop from a bulb striving toward the light to a picked flower placed in a book of poetry.
- Russian composer Tchaikovsky wrote a series of 12 piano pieces, each one named after a month of the year with a second name suggesting something associated with that month. His "April" piece is subnamed "Snow Drop". The Russian climate having a later spring, and winter ending a bit later than in other places.
- Johann Strauss II named his very successful waltz Schneeglöckchen (Snowdrops) op. 143 after this flower. The inspiration is especially evident in the cello introduction and in the slow unfurling of the opening waltz. Strauss composed this piece for a Russian Embassy dinner given at the Sperl ballroom in Vienna on 2 December 1853, but did not perform it publicly until the year 1854. The Sperl banquet was given in honour of her Excellency Frau Maria von Kalergis, daughter of the Russian diplomat and foreign minister Count Karl Nesselrode, and Strauss also dedicated his waltz to her.[107]
Symbolism
Early names refer to the association with the
In the
In more recent times, the snowdrop was adopted as a symbol of sorrow and of hope following the Dunblane massacre in Scotland, and lent its name to the subsequent campaign to restrict the legal ownership of handguns in the UK.[108][109]
See also
References
- ^ "Appendices | CITES". cites.org. Retrieved 17 January 2024.
- ^ a b c d WCLSPF 2016, Galanthus
- ^ a b c d e f g h i Ronsted et al. 2013.
- ^ Bishop et al. 2001, p. 1–2.
- ^ Stearn vernation 1992.
- ^ Davis vernation 1999.
- ^ a b c d e f g Brickell 2011.
- ^ a b Meerow & Snijman 1998
- ^ a b Dahlgren, Clifford & Yeo 1985, p. 206
- ^ Bishop et al. 2001, p. 7.
- ^ Davis 1999, G. nivalis pp. 95–96
- ^ a b Bishop, Davis & Grimshaw (2002), p. 17.
- ^ HPS 2016, V Oakes. Snowdrop mania. 2009
- ^ Bishop et al. 2001, p. 17–57.
- ^ Bishop et al. 2001, p. 40.
- ^ Theophrastus 1916, 7.13.9 p. 134
- ^ Theophrastus 1916, 6.8.1 p. 49
- ^ Gerard 1597, i cap. 78 p. 120
- ^ Linnaeus 1753, Galanthus nivalis i p. 288
- ^ Stevens 2016, Galantheae
- ^ a b c d Church 1908, Galanthus nivalis p. 17
- ^ Adanson 1763, Galanthus p. 560
- ^ Adanson 1763, Narcissi p. 57
- ^ Lamarck 1783–1808, Galanthus ii 1786 p. 590
- ^ Lamarck 1793, Galanthus pp. 359, 376
- ^ Clos 1862, Perce-neige p. 658.
- ^ Jussieu 1789, Galanthus p. 55
- ^ Jaume Saint-Hilaire 1805, Galanthus p. 139
- ^ Brown 1810, Prodromus. Amaryllideae p. 296
- ^ de Candolle 1813, Esquisse. D'une Série linéaire et par conséquent artificielle, pour la disposition des familles naturelles du règne végetal p. 219
- ^ de Lamarck & de Candolle 1815, Galanthus p. 234
- ^ Meerow et al. 1999.
- ^ Lindley 1830, Amaryllideae p. 259
- ^ Lindley 1853, Amarylleae p. 158
- ^ Bentham & Hooker 1883.
- ^ Bentham & Hooker 1883, Amaryllideae p. 711
- ^ Bentham & Hooker 1883, Amarylleae p. 718
- ^ Bentham & Hooker 1883, Galanthus p. 719
- ^ Meerow et al. 2006a.
- ^ WCLSPF 2016, G. trojanus
- ^ Davis & Özhatay 2001.
- ^ WCLSPF 2016, G. panjutinii
- ^ a b RBG 2016, Galanthus panjutinii
- ^ Zubov & Davis 2012.
- ^ WCLSPF 2016, G. samothracicus
- ^ Tan et al. 2014.
- ^ a b Stern 1956.
- ^ Traub 1963, Galanthus p. 59–60
- ^ Davis 2016.
- ^ a b Davis 1999, Taxonomy pp. 77–192
- ^ Davis 2001.
- ^ Kamenetsky 2012.
- ^ a b Lledo et al. 2004.
- ^ a b Larsen et al. 2010.
- ^ a b Margoz et al. 2013.
- ^ MBG 2016, Galanthus elwesii
- ^ Hyam & Pankhurst 1995.
- ^ Hollinger.
- ^ a b c d e f Harland 2016.
- ^ Davis 1999.
- ^ CITES Convention.
- ^ Bishop et al. 2001, pp. 341–343.
- ^ "The IUCN Red List of Threatened Species". IUCN Red List of Threatened Species. Retrieved 28 October 2018.
- ^ "The IUCN Red List of Threatened Species". IUCN Red List of Threatened Species. Retrieved 28 October 2018.
- ^ "Export quotas Galanthus elwesii". CITES. Retrieved 20 June 2021.
- ^ "Export quotas Galanthus woronowii". CITES. Retrieved 20 June 2021.
- ^ "Snowdrops and Snowdrop Gardens 2007". Great British Gardens. Archived from the original on 1 April 2007. Retrieved 11 March 2007.
- ^ "Scottish Snowdrop Festival". VisitScotland.com. Archived from the original on 3 March 2007. Retrieved 11 March 2007.
- ^ "Henry John Elwes". Colesbourne Gardens.
- ^ Ellis 2010.
- ^ Bishop et al. 2001, p. 329.
- ^ "Galanthus plicatus 'Wendy's Gold'". Plants. RHS. Retrieved 18 March 2021.
- ^ "AGM Plants – Ornamental" (PDF). Royal Horticultural Society. July 2017. p. 39. Archived (PDF) from the original on 9 October 2022. Retrieved 27 February 2018.
- ^ "Galanthus 'Ailwyn'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'Atkinsii'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'Bertram Anderson'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus elwesii". RHS Plant Selector. RHS. Retrieved 7 July 2020.
- ^ "Galanthus elwesii 'Comet'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus elwesii 'Godfrey Owen'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus elwesii 'Mrs Macnamara'". RHS. Retrieved 8 July 2020.
- ^ "RHS Plant Selector – Galanthus elwesii var. monostictus". Retrieved 7 July 2020.
- ^ "Galanthus 'John Gray'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus 'Lady Beatrix Stanley'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'Magnet'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus × hybridus 'Merlin'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus nivalis". RHS Plant Selector. RHS. Retrieved 7 July 2020.
- ^ "Galanthus nivalis f. pleniflorus 'Flore Pleno'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus nivalis 'Viridapice'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus plicatus". RHS Plant Selector. RHS. Retrieved 7 July 2020.
- ^ "Galanthus plicatus 'Augustus'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus plicatus 'Diggory'". RHS. Retrieved 8 July 2020.
- ^ "Galanthus plicatus 'Three Ships'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus reginae-olgae subsp. reginae-olgae". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'S. Arnott'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'Spindestone Surprise'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'Straffan'". RHS. Retrieved 7 July 2020.
- ^ "Galanthus 'Trumps'". RHS. Retrieved 8 July 2020.
- ^ "RHS Plant Selector – Galanthus woronowii". Retrieved 5 July 2020.
- ^ Hester et al. 1995.
- ^ Plaitakis & Duvoisin 1983.
- ^ Kaplan 2015, p. [page needed].
- ^ Loy & Schneider 2006.
- ^ "NNFCC Project Factsheet: Sustainable Production of the Natural Product Galanthamine (Defra), NF0612".
- ^ a b de la Mare 1950.
- ^ Harland 2016, Introduction
- ^ de la Mare 1929.
- ^ Kemp 1989.
- ^ McGivern 2016.
- ^ "The "Snowdrop" Campaign". www.wcc-coe.org. Retrieved 10 April 2020.
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- ISBN 9781313223874.
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- ISBN 978-3-642-64903-5. Retrieved 10 February 2014.
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Articles
- Clos, M.D. (June 1862). "Discussion de quelques points de glossologie botanique: revue critique des dénominations françaises des plantes". .
- Davis, Aaron P; Özhatay, Neriman (2001). "Galanthus trojanus: a new species of Galanthus (Amaryllidaceae) from north-western Turkey" (PDF). (PDF) from the original on 9 October 2022.
- Ewen, Stanley W. B. & S2CID 17252112.
- Fishchuk, Oksana & Odintsova, A. (August 2020). "Micromorphology and anatomy of the flowers of Galanthus nivalis and Leucojum vernum (Amaryllidaceae)". Regulatory Mechanisms in Biosystems. 11 (3): 463–468. S2CID 229370844.
- Hester, Gerko; Kaku, Hanae; Goldstein, Irwin J. & Schubert Wright, Christine (1995). "Structure of mannose-specific snowdrop (Galanthus nivalis) lectin is representative of a new plant lectin family". S2CID 45031999.
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- Lledó, Ma D.; Davis, A. P.; Crespo, M. B.; S2CID 43826738.
- Loy, C; Schneider, L (25 January 2006). "Galantamine for Alzheimer's disease and mild cognitive impairment". Cochrane Database of Systematic Reviews. 2009 (1). CD001747. PMID 16437436.
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- S2CID 85953035. Retrieved 29 April 2018.
- Mi, Xiaoxiao; Liu, Xue; Yan, Haolu; Liang, Lina; Zhou, Xiangyan; Yang, Jiangwei; Si, Huaijun; Zhang, Ning (January 2017). "Expression of the Galanthus nivalis agglutinin (GNA) gene in transgenic potato plants confers resistance to aphids". Comptes Rendus Biologies. 340 (1): 7–12. PMID 27938939.
- Plaitakis, Andreas & Duvoisin, Roger C. (1983). "Homer's moly identified as Galanthus nivalis L.: physiologic antidote to stramonium poisoning". S2CID 19839512.
- Rønsted, Nina; Zubov, Dimitri; Bruun-Lund, Sam; Davis, Aaron P. (October 2013). "Snowdrops falling slowly into place: An improved phylogeny for Galanthus (Amaryllidaceae)". PMID 23747523.
- Semerdjieva, Ivanka; Sidjimova, Boryana; Yankova-Tsvetkova, Elina; Kostova, Milena; Zheljazkov, Valtcho (December 2019). "Study on Galanthus species in the Bulgarian flora". PMID 32373724.
- Taşci Margoz, Nivart; Yüzbaşioğlu, İbrahim Sırrı; Çelen, Zeynep; Ekim, Tuna; Bilgin, Ayşe Neşe (2013). "Molecular phylogeny of Galanthus (Amaryllidaceae) of Anatolia inferred from multiple nuclear and chloroplast DNA regions". Turkish Journal of Botany. 37: 993–1007. .
- Tan, Kit; Siljak-Yakovlev, Sonja; Biel, Burkhard (2014). "Galanthus samothracicus (Amaryllidaceae) from the island of Samothraki, northeastern Greece". Phytologica Balcanica. 20 (1): 6570.
- Zubov, Dmitriy A.; Davis, Aaron P. (20 April 2012). "Galanthus panjutinii sp. nov.: a new name for an invalidly published species of Galanthus (Amaryllidaceae) from the northern Colchis area of Western Transcaucasia" (PDF). (PDF) from the original on 9 October 2022.
Websites
- de la Mare, Walter (1929). "The Snowdrop". Retrieved 14 October 2016 – via Poetry Nook.
- Ellis, Siân (1 May 2010). "Snowdrops and Strange Galanthophiles". British Heritage Travel. British Heritage Society.
- Hollinger, Jason. "Plant Latin". Archived from the original on 13 October 2016. Retrieved 13 October 2016.
- Kemp, Peter (1989). "STRAUSS II, J.: Edition - Vol. 7". Naxos. Retrieved 29 June 2020.
- McGivern, Mark (12 March 2016). "Dunblane 20 years on: Scotland unites to pay tribute". Daily Record. Retrieved 10 April 2020.
- Randerson, James (15 January 2008). "Árpád Pusztai: Biological Divide". The Guardian. London. Retrieved 25 April 2010.
Images
- "Digital Collections: Galanthus" (Botanical illustrations). New York Public Library. 2016. Retrieved 26 October 2016.
- "Vernation patterns in Galanthus" (Figure). hort.net. Retrieved 20 October 2016., in Stearn (1992)
- "Leaf vernation" (Figure). Kehan Harman: An Interactive Key to the Genus Galanthus L. Retrieved 20 October 2016., in Davis (1999)
Organisations
- Convention on International Trade in Endangered Species of Wild Fauna and Flora (Report). 3 March 1973 [Amended on 22 June 1979, Amended on 30 April 1983]. Retrieved 20 June 2021. Convention (PDF) (Report). Archived (PDF) from the original on 9 October 2022.
- "Appendices". 1973 [valid from 14 February 2021]. Retrieved 20 June 2021. "Appendices" (PDF). Archived (PDF) from the original on 9 October 2022.
- MBG. "Missouri Botanical Garden". Retrieved 7 October 2016.
- Stevens, P.F. (2016) [2001], Angiosperm Phylogeny Website, Missouri Botanical Garden, retrieved 10 October 2016
- . Board of Trustees of the Royal Botanic Gardens, Kew. Retrieved 11 October 2016.
- "World Checklist of Selected Plant Families". The Board of Trustees of the Royal Botanic Gardens, Kew. Retrieved 6 October 2016.
- "Davis, Aaron P". Royal Botanical Gardens Kew. Retrieved 12 October 2016.
- "Galanthus nivalis (common snowdrop)". Royal Botanic Garden Kew.
- HPS. "Hardy Plant Society". Kent. Retrieved 12 October 2016.
- "Shepton Mallet Horticultural Society". Shepton Mallet, Somerset. 2011.
Further reading
- Articles
- Yüzbaşıoğlu, Sırrı (2012). "Morphological variations of Galanthus elwesii in Turkey and difficulties on identification" (PDF). Bocconea. 24: 335–339. Archived (PDF) from the original on 9 October 2022.
- Websites
- "Galanthamomanie – von der Liebe zu Galanthus Schneeglöckchen, Snowdrops, Sneeuwklokjes, Perce-Neiges" (in German). Retrieved 16 October 2016.
- "Snowdrop Week: Galanthus 'John Gray'". The Frustrated Gardener. 9 February 2015. Retrieved 12 October 2016.
- "Snowdrops: An introduction". 23 March 2015.
- Bourne, Val (2 February 2015). "Wonderful snowdrop varieties". Daily Telegraph. Retrieved 15 October 2016.
- Bourne, Val. "Signs of spring". Val Bourne. Archived from the original on 18 October 2016. Retrieved 15 October 2016.
- Winterman, Denise (2 February 2012). "Snowdrop fanciers and their mania". BBC News. BBC. Retrieved 12 October 2016.
External links
- "Convention on International Trade in Endangered Species of Wild Fauna and Flora". UNEP.
- "The IUCN Red List of Threatened Species". International Union for Conservation of Nature and Natural Resources. Archived from the originalon 27 June 2014. Retrieved 18 November 2014.
- Antheunisse, Max. "Galanthus L." plantillustrations.org. Retrieved 19 October 2016.
- Vigneron, Pascal. "Amaryllidaceae". Amaryllidaceae.org (in French). Archived from the original on 4 January 2015. Retrieved 23 October 2014.