Gargoyleosaurus

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Gargoyleosaurus
Temporal range:
Ma
Fossil skeleton, Denver Museum of Nature and Science
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Thyreophora
Clade: Ankylosauria
Family: Nodosauridae
Genus: Gargoyleosaurus
Carpenter et al. 1998
Species:
G. parkpinorum
Binomial name
Gargoyleosaurus parkpinorum
Carpenter et al. 1998

Gargoyleosaurus (meaning "

The type species, G. parkpinorum (originally G. parkpini) was described by Ken Carpenter et al. in 1998. A mounted skeletal reconstruction of Gargoyleosaurus parkpinorum can be seen at the Denver Museum of Nature and Science and, alongside a couple skeletons of baby Stegosaurus, has been on display there since around 2002. Gargoyleosaurus was present in stratigraphic zone 2 of the Morrison Formation.[2]

Discovery

Size comparison

The holotype specimen of Gargoyleosaurus parkpinorum was collected by Western Paleontology Labs in 1995 and is currently held in the collections of the Denver Museum of Nature and Science, Denver, Colorado. Besides the holotype, two other partial skeletons are known (although not yet described). The holotype consists of most of the skull and a partial postcranial skeleton. The specimen was originally described as Gargoyleosaurus parkpini by Carpenter, Miles and Cloward in 1998, then renamed G. parkpinorum by Carpenter et al. in 2001, in accordance with ICZN art. 31.1.2A.

Description

Gargoyleosaurus was a relatively small ankylosaur, reaching 3–3.5 m (9.8–11.5 ft) in length and 300–754 kg (661–1,662 lb) in body mass.

osseous nasal septum, a linearly arranged nasal cavity, the absence of an osseus secondary palate, and, as regards osteoderms, two sets of co-ossified cervical plates and a number of elongate conical spines.[1] A very unusual feature is the sagittal (midline) osteoderm on the first set of cervical plates; in most other ankylosaurs, these osteoderms are bilateral, i.e. paired with one on each side of the midline.[3]

Classification

Gargoyleosaurus parkpinorum (DMNH 27726) Skull and cervical armor in side and front view. Insert shows a cheek tooth. The skull is slightly crushed changing the shape of the orbit.
Life restoration

Vickaryous et al. (2004) place Gargoyleosaurus parkpinorum within the family

phylogenetic hypotheses, which place the genus as the sister group
to all other ankylosaurids (i.e., members of the Ankylosauridae). These studies however, only utilized the skull, whereas many of the distinctive features of the family Polacanthidae are in the postcranial skeleton.

Below is a reproduced phylogenetic analysis from Soto-Acuña et al. (2021):[6]

Thyreophora

Scelidosaurus

Stegosauria

Ankylosauria
Parankylosauria

Kunbarrasaurus

Antarctopelta

Stegouros

Euankylosauria
Nodosauridae

Cedarpelta

Dongyangopelta

Gargoyleosaurus

Gastonia

Hylaeosaurus

Mymoorapelta

Panoplosaurus

Peloroplites

Polacanthus

Hoplitosaurus

Sauroplites

Taohelong

Nodosaurinae

Ankylosauridae

Aletopelta

Chuanqilong

Liaoningosaurus

Gobisaurus

Shamosaurus

Ankylosaurinae

See also

References

  • Carpenter, K., Miles, C. and Cloward, K. (1998). "Skull of a Jurassic ankylosaur (Dinosauria)." Nature 393: 782–783.
  • Carpenter, K. (ed.) The Armored Dinosaurs. pp. 454–483. Indiana University Press, Bloomington.
  • Vickaryous, Maryanska, and Weishampel (2004). "Ankylosauria". in The Dinosauria (2nd edition), Weishampel, D. B., Dodson, P., and Osmólska, H., editors. University of California Press.
  • Killbourne, B. and Carpenter, K. (2005). "Redescription of Gargoyleosaurus parkpinorum, a polacanthid ankylosaur from the Upper Jurassic of Albany County, Wyoming". Neues Jahrbuch für Geologie und Paläontologie, 237, 111–160.
  1. ^ .
  2. ^ Foster, J. (2007). "Appendix." Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. pp. 327–329.
  3. ^ a b Kilbourne, B. and Carpenter, K. 2005. "Redescription of Gargoyleosaurus parkpinorum, a polacanthid ankylosaur from the Upper Jurassic of Albany County, Wyoming". N. Jb. Geol. Palaont. Abh. 237: 111–160
  4. S2CID 250422526
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  5. .
  6. .

External links