Marine habitat

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(Redirected from
Marine habitats
)
This article is about a specific type of environment within the marine ecosystem. For the broader concept with all interactions and characteristics of the marine environment, see marine ecosystem.

Coral reefs provide marine habitats for tube sponges, which in turn become marine habitats for fishes
Marine habitats
Coastal habitats
Ocean surface
Open ocean
Sea floor

A marine habitat is a

ecological or environmental area inhabited by one or more living species.[1]
The marine environment supports many kinds of these habitats.

Marine habitats can be divided into

shoreline out to the edge of the continental shelf
. Most marine life is found in coastal habitats, even though the shelf area occupies only seven percent of the total ocean area. Open ocean habitats are found in the deep ocean beyond the edge of the continental shelf.

Alternatively, marine habitats can be divided into pelagic and demersal zones. Pelagic habitats are found near the surface or in the open water column, away from the bottom of the ocean. Demersal habitats are near or on the bottom of the ocean. An organism living in a pelagic habitat is said to be a pelagic organism, as in pelagic fish. Similarly, an organism living in a demersal habitat is said to be a demersal organism, as in demersal fish. Pelagic habitats are intrinsically shifting and ephemeral, depending on what ocean currents are doing.

Marine habitats can be modified by their inhabitants. Some marine organisms, like corals, kelp, mangroves and seagrasses, are ecosystem engineers which reshape the marine environment to the point where they create further habitat for other organisms. By volume the ocean provides most of the habitable space on the planet.[2]

Overview

Part of a series of overviews on
Marine life

In contrast to terrestrial habitats, marine habitats are shifting and ephemeral. Swimming organisms find areas by the edge of a continental shelf a good habitat, but only while upwellings bring nutrient rich water to the surface. Shellfish find habitat on sandy beaches, but storms, tides and currents mean their habitat continually reinvents itself.

The presence of seawater is common to all marine habitats. Beyond that many other things determine whether a marine area makes a good habitat and the type of habitat it makes. For example:

Two views of the ocean from space
Only 29 percent of the world surface is land. The rest is ocean, home to the marine habitats. The oceans are nearly four kilometres deep on average and are fringed with coastlines that run for nearly 380,000 kilometres.

There are five major oceans, of which the Pacific Ocean is nearly as large as the rest put together. Coastlines fringe the land for nearly 380,000 kilometres.

Ocean Area
million km2 		% Volume[3]
million cu km 		% Mean depth
km 		Max depth
km 		Coastline
km 		% Ref
Pacific Ocean 155.6 46.4 679.6 49.6 4.37 10.924 135,663 [4]
Atlantic Ocean 76.8 22.9 313.4 22.5 4.08 8.605 111,866 [5]
Indian Ocean 68.6 20.4 269.3 19.6 3.93 7.258 66,526 [6]
Southern Ocean 20.3 6.1 91.5 6.7 4.51 7.235 17,968 [7]
Arctic Ocean 14.1 4.2 17.0 1.2 1.21 4.665 45,389 [8]
Overall 335.3 1370.8[9] 4.09 10.924 377,412
Land runoff
, pouring into the sea, can contain nutrients

Altogether, the ocean occupies 71 percent of the world surface, averaging nearly four kilometres in depth. By volume, the ocean contains more than 99 percent of the Earth's liquid water.[10][11][12] The science fiction writer Arthur C. Clarke has pointed out it would be more appropriate to refer to the planet Earth as the planet Sea or the planet Ocean.[13][14]

Marine habitats can be broadly divided into

demersal habitats. Pelagic habitats are the habitats of the open water column, away from the bottom of the ocean. Demersal habitats are the habitats that are near or on the bottom of the ocean. An organism living in a pelagic habitat is said to be a pelagic organism, as in pelagic fish. Similarly, an organism living in a demersal habitat is said to be a demersal organism, as in demersal fish. Pelagic habitats are intrinsically ephemeral, depending on what ocean currents
are doing.

The land-based ecosystem depends on topsoil and fresh water, while the marine ecosystem depends on dissolved nutrients washed down from the land.[15]

Ocean deoxygenation poses a threat to marine habitats, due to the growth of low oxygen zones.[16]

Ocean currents

Main article: Ocean current
land runoff or upwellings
at the edge of the continental shelf.
North Atlantic
gyre
North Atlantic
gyre
North Atlantic
gyre
Indian
Ocean
gyre
North
Pacific
gyre
South
Pacific
gyre
South Atlantic
        gyre
Map showing 5 circles. The first is between western Australia and eastern Africa. The second is between eastern Australia and western South America. The third is between Japan and western North America. Of the two in the Atlantic, one is in hemisphere.
Ocean gyres rotate clockwise in the north and counterclockwise in the south

In marine systems,

marine invertebrates. If sufficient zooplankton establish themselves, the current becomes a candidate habitat for the forage fish that feed on them. And then if sufficient forage fish move to the area, it becomes a candidate habitat for larger predatory fish
and other marine animals that feed on the forage fish. In this dynamic way, the current itself can, over time, become a moving habitat for multiple types of marine life.

Ocean currents can be generated by differences in the density of the water. How dense water is depends on how

waves and seiches. Ocean currents are also generated by the gravitational pull of the sun and moon (tides), and seismic activity (tsunami).[17]

The rotation of the Earth affects the direction ocean currents take, and explains which way the large circular

Coriolis effect. It is weakest at the equator and strongest at the poles. The effect is opposite south of the equator, where currents curve left.[17]

Topography

This section is an excerpt from Seabed § Topography.[edit]
World map with ocean topography

Seabed topography (ocean topography or marine topography) refers to the shape of the land (topography) when it interfaces with the ocean. These shapes are obvious along coastlines, but they occur also in significant ways underwater. The effectiveness of marine habitats is partially defined by these shapes, including the way they interact with and shape ocean currents, and the way sunlight diminishes when these landforms occupy increasing depths. Tidal networks depend on the balance between sedimentary processes and hydrodynamics however, anthropogenic influences can impact the natural system more than any physical driver.[18]

Marine topographies include

shorelines to continental shelves and coral reefs. Further out in the open ocean, they include underwater and deep sea features such as ocean rises and seamounts. The submerged surface has mountainous features, including a globe-spanning mid-ocean ridge system, as well as undersea volcanoes,[19] oceanic trenches, submarine canyons, oceanic plateaus and abyssal plains
.

The mass of the oceans is approximately 1.35×1018 
metric tons, or about 1/4400 of the total mass of the Earth. The oceans cover an area of 3.618×108 km2 with a mean depth of 3,682 m, resulting in an estimated volume of 1.332×109 km3.[20]

Biomass

See also:
Marine biomass

One measure of the relative importance of different marine habitats is the rate at which they produce biomass.

Producer Biomass productivity
(gC/m2/yr) 		Ref Total area
(million km2) 		Ref Total production
(billion tonnes C/yr) 		Comment
swamps and marshes 2,500 [21] Includes freshwater
coral reefs 2,000 [21] 0.28 [22] 0.56
algal beds 2,000 [21]
river estuaries 1,800 [21]
open ocean 125 [21][23] 311 39

Coastal

Coastlines can be volatile habitats
Main article:
Marine coastal ecosystems

drowned river valleys
.

The main agents responsible for deposition and erosion along coastlines are waves, tides and currents. The formation of coasts also depends on the nature of the rocks they are made of – the harder the rocks the less likely they are to erode, so variations in rock hardness result in coastlines with different shapes.

Tides often determine the range over which sediment is deposited or eroded. Areas with high tidal ranges allow waves to reach farther up the shore, and areas with lower tidal ranges produce deposition at a smaller elevation interval. The tidal range is influenced by the size and shape of the coastline. Tides do not typically cause erosion by themselves; however, tidal bores can erode as the waves surge up river estuaries from the ocean.[24]

Shores that look permanent through the short perceptive of a human lifetime are in fact among the most temporary of all marine structures.[25]

Waves erode coastline as they break on shore releasing their energy; the larger the wave the more energy it releases and the more sediment it moves. Sediment deposited by waves comes from eroded cliff faces and is moved along the coastline by the waves. Sediment deposited by rivers is the dominant influence on the amount of sediment located on a coastline.[26]

The

Francis Shepard classified coasts as primary or secondary.[27]
  The global continental shelf, highlighted in light green, defines the extent of marine coastal habitats, and occupies 5% of the total world area

Continental coastlines usually have a

land runoff into coastal waters. Further, periodic upwellings
from the deep ocean can provide cool and nutrient rich currents along the edge of the continental shelf.

As a result, coastal marine life is the most abundant in the world. It is found in

sea grass meadows in between: the reefs protect the mangroves and seagrass from strong currents and waves that would damage them or erode the sediments in which they are rooted, while the mangroves and seagrass protect the coral from large influxes of silt, fresh water and pollutants. This additional level of variety in the environment is beneficial to many types of coral reef animals, which for example may feed in the sea grass and use the reefs for protection or breeding.[32]

Coastal habitats are the most visible marine habitats, but they are not the only important marine habitats. Coastlines run for 380,000 kilometres, and the total volume of the ocean is 1,370 million cu km. This means that for each metre of coast, there is 3.6 cu km of ocean space available somewhere for marine habitats.

Waves and currents shape the intertidal shoreline, eroding the softer rocks and transporting and grading loose particles into shingles, sand or mud

Intertidal

Main article: Intertidal zone

tides
. A huge array of life lives within this zone.

Shore habitats range from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of bioerosion.

Sandy shores

Sandy shores provide shifting homes to many species
See also: Beach

Sandy shores, also called

offshore bars, which give beaches some stability by reducing erosion.[33]

Sandy shores are full of life, The grains of sand host diatoms, bacteria and other microscopic creatures. Some fish and turtles return to certain beaches and spawn eggs in the sand. Birds habitat beaches, like gulls, loons, sandpipers, terns and pelicans. Aquatic mammals, such sea lions, recuperate on them. Clams, periwinkles, crabs, shrimp, starfish and sea urchins are found on most beaches.[34]

Sand is a

sand dunes. Beyond the high tide mark, if the beach is low-lying, the wind can form rolling hills of sand dunes. Small dunes shift and reshape under the influence of the wind while larger dunes stabilise the sand with vegetation.[33]

Ocean processes grade loose sediments to

particle sizes other than sand, such as gravel or cobbles. Waves breaking on a beach can leave a berm, which is a raised ridge of coarser pebbles or sand, at the high tide mark. Shingle beaches are made of particles larger than sand, such as cobbles, or small stones. These beaches make poor habitats. Little life survives because the stones are churned and pounded together by waves and currents.[33]

Rocky shores

Tidepools
on rocky shores make turbulent habitats for many forms of marine life
See also:
Tidal pool

The relative solidity of rocky shores seems to give them a permanence compared to the shifting nature of sandy shores. This apparent stability is not real over even quite short geological time scales, but it is real enough over the short life of an organism. In contrast to sandy shores, plants and animals can anchor themselves to the rocks.[36]

Competition can develop for the rocky spaces. For example, barnacles can compete successfully on open intertidal rock faces to the point where the rock surface is covered with them. Barnacles resist desiccation and grip well to exposed rock faces. However, in the crevices of the same rocks, the inhabitants are different. Here mussels can be the successful species, secured to the rock with their byssal threads.[36]

Rocky and sandy coasts are vulnerable because humans find them attractive and want to live near them. An increasing proportion of the humans live by the coast, putting pressure on coastal habitats.[36]

Mudflats

Mudflats become temporary habitats for migrating birds
See also: Bay mud

estuarine silts, clays and marine animal detritus. Most of the sediment within a mudflat is within the intertidal zone
, and thus the flat is submerged and exposed approximately twice daily.

Mangrove forests and salt marshes

Mangroves provide nurseries for fish
See also: Mangrove forest, Ecological values of mangroves, and Salt marsh

Mangrove swamps and salt marshes
form important coastal habitats in tropical and temperate areas respectively.

Mangroves are species of shrubs and medium size trees that grow in saline coastal sediment habitats in the tropics and subtropics – mainly between latitudes 25° N and 25° S. The saline conditions tolerated by various species range from brackish water, through pure seawater (30 to 40 ppt), to water concentrated by evaporation to over twice the salinity of ocean seawater (up to 90 ppt).[37][38] There are many mangrove species, not all closely related. The term "mangrove" is used generally to cover all of these species, and it can be used narrowly to cover just mangrove trees of the genus Rhizophora.

Mangroves form a distinct characteristic saline

depositional coastal environments, where fine sediments (often with high organic content) collect in areas protected from high-energy wave action. Mangroves dominate three quarters of tropical coastlines.[38]

Estuaries

Estuaries
occur when rivers flow into a coastal bay or inlet. They are nutrient rich and have a transition zone which moves from freshwater to saltwater.
Main article:
Estuaries

An

coastal body of water with one or more rivers or streams flowing into it, and with a free connection to the open sea.[40] Estuaries form a transition zone between river environments and ocean environments and are subject to both marine influences, such as tides, waves, and the influx of saline water; and riverine influences, such as flows of fresh water and sediment. The inflow of both seawater and freshwater provide high levels of nutrients in both the water column and sediment, making estuaries among the most productive natural habitats in the world.[41]

Most estuaries were formed by the flooding of river-eroded or glacially scoured valleys when sea level began to rise about 10,000-12,000 years ago.[42] They are amongst the most heavily populated areas throughout the world, with about 60% of the world's population living along estuaries and the coast. As a result, estuaries are suffering degradation by many factors, including sedimentation from soil erosion from deforestation; overgrazing and other poor farming practices; overfishing; drainage and filling of wetlands; eutrophication due to excessive nutrients from sewage and animal wastes; pollutants including heavy metals, PCBs, radionuclides and hydrocarbons from sewage inputs; and diking or damming for flood control or water diversion.[42]

Estuaries provide habitats for a large number of organisms and support very high productivity. Estuaries provide habitats for

dinoflagellates
which are abundant in the sediment.

Kelp forests

Further information: Kelp forest
Kelp forests provide habitat for many marine organisms

polar coastal oceans.[45]

Kelp forests provide a unique three-dimensional habitat for marine organisms and are a source for understanding many ecological processes. Over the last century, they have been the focus of extensive research, particularly in

ecosystem services.[47]

However, humans have contributed to kelp

herbivores from their normal population regulation and result in the over-grazing of kelp and other algae.[48] This can rapidly result in transitions to barren landscapes where relatively few species persist.[49]

Frequently considered an

Plantae). The morphological structure of a kelp thallus is defined by three basic structural units:[49]

  • The holdfast is a root-like mass that anchors the thallus to the sea floor, though unlike true roots it is not responsible for absorbing and delivering nutrients to the rest of the thallus;
  • The stipe is analogous to a plant stalk, extending vertically from the holdfast and providing a support framework for other morphological features;
  • The
    fronds
    are leaf- or blade-like attachments extending from the stipe, sometimes along its full length, and are the sites of nutrient uptake and photosynthetic activity.

In addition, many kelp species have pneumatocysts, or gas-filled bladders, usually located at the base of fronds near the stipe. These structures provide the necessary buoyancy for kelp to maintain an upright position in the water column.

The environmental factors necessary for kelp to survive include hard substrate (usually rock), high nutrients (e.g., nitrogen, phosphorus), and light (minimum annual

perennial, living for more than 20 years.[53] In perennial kelp forests, maximum growth rates occur during upwelling months (typically spring and summer) and die-backs correspond to reduced nutrient availability, shorter photoperiods and increased storm frequency.[49]

Seagrass meadows

Fan mussel in a Mediterranean seagrass meadow
Main article:
Seagrass meadows

Seagrasses are flowering plants from one of four plant families which grow in marine environments. They are called seagrasses because the leaves are long and narrow and are very often green, and because the plants often grow in large meadows which look like grassland. Since seagrasses photosynthesize and are submerged, they must grow submerged in the photic zone, where there is enough sunlight. For this reason, most occur in shallow and sheltered coastal waters anchored in sand or mud bottoms.

Seagrasses form extensive

green turtles, dugongs, manatees, fish, geese, swans, sea urchins and crabs
.

Seagrasses are

fishing grounds, wave protection, oxygen production and protection against coastal erosion. Seagrass meadows account for 15% of the ocean's total carbon storage.[54]

Reefs

Main article: Reef

A reef is a ridge or

biotic processes dominated by corals and coralline algae. Artificial reefs such as shipwrecks and other anthropogenic underwater structures may occur intentionally or as the result of an accident, and sometimes have a designed role in enhancing the physical complexity of featureless sand bottoms, thereby attracting a more diverse assemblage of organisms. Reefs are often quite near to the surface, but not all definitions require this.[55] Fringing reefs, the most common type of reef, are found close to shorelines and surrounding islands.[56]

Rocky reefs

Coral reefs

Main article: Coral reef
See also: Coral reef fish

tropical coral reefs which exist in most tropical waters; however, coral reefs can also exist in cold water. Reefs are built up by corals and other calcium-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create artificial reefs. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic zooxanthellae
, tropical fish and many other organisms.

Much attention in marine biology is focused on coral reefs and the

global warming could exacerbate this trend.[59][60][61][62]

Surface waters

epipelagic waters get enough light for photosynthesis, but there are often not enough nutrients. As a result, large areas contain little life apart from migrating animals.[63]
See also: Ocean surface ecosystem

Surface microlayer

The surface microlayer of the ocean serves as the transitional area between the atmosphere and the ocean. It covers around 70% of the Earth's surface as it covers most of the ocean waters on the planet.[64] The microlayer is known for its unique biological and chemical properties which give it a small ecosystem of its own and serves as a distinct habitat from the deeper ocean waters.

The surface microlayer is not in fact entirely aqueous like the rest of the ocean, but is closer to a kind of hydrated gel composed of concentrated nutrients forming a biological film over the water it covers. This film is rich in microbes which mediate the interactions between the sun, the atmosphere, and the waters below.

Although thin, the surface microlayer is critical for life beneath it. Because of the environment rich in microbes and nutrients, larvae of fish and other aquatic animals are often laid in the microlayer to incubate. The plankton in the microlayer are distinctly adapted to withstand high levels of radiation, and serve as buffers to prevent this potentially harmful radiation from reaching the deeper water. Environmental changes such as aerosols or dust storms can cause these surface plankton to become overproductive, leading to blooms.[64]

Halfbeak as larvae are one of the organisms adapted to the unique properties of the microlayer

Because of the unique properties of the microlayer, pollutants often accumulate within and use it to reach other parts of the ocean. Hydrophobic compounds, such as petroleum, flame retardants, and heavy metals, have a particular affinity for the surface microlayer. Recently, the abundance of aerosols and microplastics has also had an impact on the SML and their accumulation has led to many problems, such as animal ingestion of these compounds leading to widespread disruption of balance and spread of these compounds among marine communities.

The surface microlayer is also critical to gas exchange between the atmosphere and the ocean. Because the microlayer is filled with microbes, it is widely theorized that it plays a critical role in gas exchange and uptake of nutrients, but relatively little data on this has been collected. The central feature of the microlayer is the temperature, as it is an indicator of how pollutants and human activity affects the ocean.[64]

Main article: Sea surface microlayer

Epipelagic zone

The surface waters are sunlit. The waters down to about 200 metres are said to be in the

epipelagic zone. Enough sunlight enters the epipelagic zone to allow photosynthesis by phytoplankton. The epipelagic zone is usually low in nutrients. This partially because the organic debris produced in the zone, such as excrement and dead animals, sink to the depths and are lost to the upper zone. Photosynthesis can happen only if both sunlight and nutrients are present.[63]

In some places, like at the edge of continental shelves, nutrients can

land runoff can be distributed by storms and ocean currents. In these areas, given that both sunlight and nutrients are now present, phytoplankton can rapidly establish itself, multiplying so fast that the water turns green from the chlorophyll, resulting in an algal bloom. These nutrient rich surface waters are among the most biologically productive in the world, supporting billions of tonnes of biomass.[63]

"Phytoplankton are eaten by

foodchain, the small forage fish are in turn eaten by larger predators, such as tuna, marlin, sharks, large squid, seabirds, dolphins, and toothed whales.[63]

Open ocean

Elevation-area graph showing the proportion of land area at given heights and the proportion of ocean area at given depths
See also: Pelagic fish

The open ocean is relatively unproductive because of a lack of nutrients, yet because it is so vast, it has more overall primary production than any other marine habitat. Only about 10 percent of marine species live in the open ocean. But among them are the largest and fastest of all marine animals, as well as the animals that dive the deepest and migrate the longest. In the depths lurk animal that, to our eyes, appear hugely alien.[65]

Deep sea

Some representative ocean animal life (not drawn to scale) within their approximate depth-defined ecological habitats. Marine microorganisms exist on the surfaces and within the tissues and organs of the diverse life inhabiting the ocean, across all ocean habitats.[66]
Scale diagram of the layers of the pelagic zone
See also:
Deep sea communities

The

bio-luminescence.[citation needed
]

In the deep ocean, the waters extend far below the epipelagic zone, and support very different types of pelagic life forms adapted to living in these deeper zones.[67]

Much of the

benthic ecosystems: As sunlight cannot reach them, deep-sea organisms rely heavily on marine snow as an energy source.[68]

Some deep-sea pelagic groups, such as the

continental slopes. The phenomenon is explained by the likewise abundance of prey species which are also attracted to the structures.[citation needed
]

The fish in the different pelagic and deep water benthic zones are physically structured, and behave in ways, that differ markedly from each other. Groups of coexisting species within each zone all seem to operate in similar ways, such as the small mesopelagic

rattails. "[69]

The umbrella mouth gulper eel can swallow a fish much larger than itself

Ray finned species, with spiny fins, are rare among deep sea fishes, which suggests that deep sea fish are ancient and so well adapted to their environment that invasions by more modern fishes have been unsuccessful.[70] The few ray fins that do exist are mainly in the Beryciformes and Lampriformes, which are also ancient forms. Most deep sea pelagic fishes belong to their own orders, suggesting a long evolution in deep sea environments. In contrast, deep water benthic species, are in orders that include many related shallow water fishes.[71]

The umbrella mouth gulper is a deep sea eel with an enormous loosely hinged mouth. It can open its mouth wide enough to swallow a fish much larger than itself, and then expand its stomach to accommodate its catch.[72]

Sea floor

Part of a series related to
Benthic life
Benthopelagic (coupling)        Seabed
By size
By type
By location
    • endobenthos
    • epibenthos
    • hyperbenthos
By habitat
    Shallow:
bivalve reef
                   mangrove forest • coral reef
                   seagrass meadow
        Deep: seamount • cold seep
                   hydrothermal vent
Communities
Related
See also: Benthic zone, benthos, and Marine sediment

Vents and seeps

marine microorganisms
and other lifeforms have been discovered at these locations.

Trenches

The deepest recorded

Trieste when it dove to the bottom in 1960.[73]

Seamounts

Marine life also flourishes around seamounts that rise from the depths, where fish and other sea life congregate to spawn and feed.

Anthropogenic impacts

Mudflat pollution
See also: Human impact on marine life

Mudflats are typically important regions for wildlife, supporting a large population, although levels of biodiversity are not particularly high. They are of particular importance to

Biodiversity Action Plan priority habitat. European countries such as France have also found it beneficial to use the Marine Influence Index (MII) to be able to monitor the responses to pollution the local plant and animal species may have as well as monitor any type of deviation from the natural patterns displayed previously.[76]

Although many parts of the seafloor have yet to be explored, researchers have found that parts of it have been greatly affected by human activity. Bottom trawling, microplastic pollution, and industrial metals have slowly changed and altered the composition of the sea floor. Bottom trawling refers to a commercial deep sea fishing technique in which the equipment drags across the sea floor.[77] This has had an adverse effect on the seafloor as it changes the surface structure and composition. In addition, microplastic pollution has become an increasing problem to the seafloor as plastics and other debris are found in many of the sediments.[78] Due to the build up of litter, the habitats and environments of organisms on the seafloor are being impacted and changed. This includes industrial facilities dumping new metals and minerals, such as cadmium, onto the seafloor that change the chemical composition of the water and poison the inhabitants.[79]

amphipod Eurythenes plasticus
, named after microplastics found in its body, demonstrating plastic pollution affects marine habitats even 6000m below sea level.
  • Microplastics found in sediments on the seafloor
    Microplastics found in sediments on the seafloor
  • 640 µm microplastic found in the deep sea amphipod Eurythenes plasticus
    640 µm microplastic found in the deep sea amphipod Eurythenes plasticus

There are also negative anthropogenic impacts on deep sea habitats, including trash pollution and chemical pollution.

Gill nets for example, have been recorded tangled around deep sea corals and continue ghost fishing for extended periods of time.[82]

Gallery

See also

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Sources

External links

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