Mesonychidae

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Mesonychidae
Temporal range: Early Paleocene–Early Oligocene
Harpagolestes immanis skull
Life reconstruction of Harpagolestes
macrocephalus
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Mesonychia
Family: Mesonychidae
Cope 1880[1]
Genera

See text

Mesonychidae (meaning "middle

artiodactyls.[2] Some researchers now consider the family a sister group either to whales or to artiodactyls, close relatives rather than direct ancestors. Other studies define Mesonychia as basal to all ungulates, occupying a position between Perissodactyla and Ferae. In this case, the resemblances to early whales would be due to convergent evolution among ungulate-like herbivores that developed adaptations related to hunting or eating meat.[3]

Description

The mesonychids were an unusual group of

condylarths with a specialized dentition featuring tri-cuspid upper molars and high-crowned lower molars with shearing surfaces. They had large heads with relatively long necks. Over time, the family evolved foot and leg adaptations for faster running, and jaw adaptations for greater bite force. Like the Paleocene family Arctocyonidae, mesonychids were once viewed as primitive carnivorans, and the diet of most genera probably included meat or fish. Various genera and species coexisted in some locations, as hunters and omnivores or scavengers.[4]
In contrast to arctocyonids, the mesonychids had only four digits furnished with hooves supported by narrow fissured end phalanges. [5]

Evolutionary history

They first appeared in the Early Paleocene, undergoing numerous speciation events during the Paleocene, and Eocene. Mesonychids fared very poorly at the close of the Eocene epoch, with only one genus, Mongolestes,[6] surviving into the Early Oligocene epoch.

Restoration of Mesonyx

Mesonychids probably originated in

Bering land bridge
.

Taxonomy

Mesonychidae was named by Cope (1880). Its type genus is Mesonyx. It was assigned to Creodonta by Cope (1880); to Creodonta by Cope (1889); to Carnivora by Peterson (1919); to Mesonychia by Carroll (1988) and Zhou et al. (1995); and to Cete by Archibald (1998);[7] and to Mesonychia by Carroll (1988), Zhou et al. (1995), Geisler and McKenna (2007) and Spaulding et al. (2009).[8]

Classification

Family Mesonychidae

  • Genus Ankalagon
    • A. saurognathus
  • Genus Dissacus
    • D. argenteus
    • D. europaeus
    • D. indigenus
    • D. magushanensis
    • D. navajovius
    • D. praenuntius
    • D. rotundus
    • D. serior
    • D. serratus
    • D. willwoodensis
    • D. zanabazari
    • D. zengi
  • Genus Guilestes
    • G. acares
  • Genus Harpagolestes
    • H. immanis
    • H. koreanicus
    • H. orientalis
  • Genus Hessolestes
    • H. ultimus
  • Genus Hukoutherium
    • H. ambigum
    • H. shimemensis
  • Genus
    Jiangxia
    • J. chaotoensis
  • Genus Mesonyx
    • M. nuhetingensis
    • M. obtusidens
    • M. uintensis
    • M. uqbulakensis
  • Genus Mongolestes
    • M. hadrodens
    • M. huangheensis
  • Genus Mongolonyx
    • M. dolichognathus
    • M. robustus
  • Genus Pachyaena
    • P. gigantea
    • P. intermedia
    • P. ossifraga
    • P. gracilis
  • Genus Sinonyx
    • S. jiashanensis
  • Genus Synoplotherium
    • S. vorax
  • Genus Yantanglestes
    • Y. conexus

References

Notes

  1. ^ E. D. Cope 1880 in the Paleobiology Database. Retrieved 11 August 2013.
  2. ^ Wong, Kate (2002). "The Mammals that Conquered the Seas; New Fossils and DNA Analyses Elucidate the Remarkable History of Whales" (PDF). Scientific American. May 2002: 71–79 – via mtsac.edu.
  3. PMID 19774069
    .
  4. .
  5. ^ Jordi & Anton 2002, p. 13
  6. ^ Jin 2005
  7. ^ J. D. Archibald. 1998. Archaic ungulates ("Condylarthra"). In C. M. Janis, K. M. Scott, and L. L. Jacobs (eds.), Evolution of Tertiary Mammals of North America 1:292-331
  8. ^ Mesonychidae (condylarth) in the Paleobiology Database. Retrieved 11 August 2013.

Cited sources