Permocupedidae
Permocupedidae Temporal range:
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Fossil and life restoration of Permocupes sojanensis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Suborder: | †Protocoleoptera |
Superfamily: | †Permocupedoidea |
Family: | †Permocupedidae Martynov, 1933[1] |
Genera | |
See text | |
Synonyms | |
Kaltanocoleidae Rohdendorf, 1961 |
Permocupedidae is a family of
xylophagous (wood eating), which is presumed to be the ancestral ecology of beetles.[3]
Taxonomy
Kirejtshuk (2020) included the following genera in an expanded (sensu stricto) Permocupedidae and related Taldycupedidae as subfamilies Permocupedinae and Taldycupedinae respectively.[4] However, other studies have recovered Talycupedidae as more closely related to crown-group beetles than to Permocupedidae.[5]
- Afrocupes Geertsema & van den Heever, 1996 – South Africa: Western Cape (Kenmoore Farm), Whitehill Formation, Permian, Cisuralian, Sakmarian/Artinskian, −290.1–279.5 Ma.
- Afrocupes firmae Geertsema & van den Heever, 1996 (Type species)
- syn Palaeocupes Rohdendorf, 1961) – Russia, Kemerovo Region(Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma
- Archicupes jacobsoni Rohdendorf, 1961 (Type species)
- "Archicupes" reichardti Rohdendorf, 1961 – considered indeterminate to genus by Kirejtshuk (2020)
- Cytocupoides Ponomarenko, 1969 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
- Cytocupoides longatus Ponomarenko, 1969 (Type species)
- Cytocupes Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
- Cytocupes angustus Rohdendorf, 1961 (Type species)
- Eocupes Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma).
- Eocupes cellulosus Ponomarenko, 1969
- Eocupes lukjanovitshi Rohdendorf, 1961 (Type species)
- Frankencupes Ponomarenko & Bashkuev, 2018 – Germany: Gambach am Main (Lower Franconia), Röt Formation Middle Triassic, Olenekian/Anisian, −247.2–242.0 Ma.
- Frankencupes ultimus Ponomarenko & Bashkuev, 2018 (Type species)
- Ichthyocupes Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Kemerovo Region (Tykhta River, Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma.
- Ichthyocupes kuznetskiensis Ponomarenko, 1969
- Ichthyocupes skoki (Rohdendorf, 1961)
- Ichthyocupes tyzhnovi Rohdendorf, 1961 (Type species)
- Kaltanocoleus Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
- Kaltanocoleus pospelovi Rohdendorf, 1961 (Type species)
- Orenburg Region (Kargala), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Sweverodvinian), −265.0–259.0 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. One undescribed species of this genus was recorded from Indian Barakar Formationof the Lower Permian.
- Kaltanicupes acutus Ponomarenko, 1963
- Kaltanicupes kargalensis Ponomarenko, 1963
- Kaltanicupes kitjakensis Ponomarenko, 1963
- Kaltanicupes richteri Rohdendorf, 1961 (Type species)
- Linicupes Ponomarenko, Yan & Huang, 2014[6] – China: Anhui (Houdong, Yinping Mountain, SW Chaohu City), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian/Vyatkian), −265.0–259.0 Ma.
- Linicupes yinpinensis Ponomarenko, Yan & Huang, 2014 (Type species)
- Maricoleus Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 – Russia, Primorsky Krai (Cape, Novosilsky, Russky Island), Lower Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
- Maricoleus valentinae Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 (Type species)
- Permocupes Martynov, 1933 (syn Permocupoides Martynov, 1933) – Russia: Arkhangel’sk Region (Letopala River and Soyana), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District, Tikhie Gory), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
- Permocupes distinctus (Martynov, 1933)
- Permocupes grandis Ponomarenko, 1963
- Permocupes latus Ponomarenko, 2013
- Permocupes semenovi Martynov, 1933 (Type species)
- Permocupes sojanensis Ponomarenko, 1963
- Pintolla Kirejtshuk, 2020 – Brazil: Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
- Pintolla ponomarenkoi (Pinto, 1987)[7] (Type species)
- Vologda Region(Isady locality, Mutovino), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinskian/Vyatkian), −259.0–254.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma; Kemerovo Region (Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
- Protocupoides acer (Rohdendorf, 1961)
- Protocupoides carinatus (Rohdendorf, 1961)
- Protocupoides elongatus Ponomarenko, 2013
- Protocupoides esini Ponomarenko, 2013
- Protocupoides plavilstshikovi Rohdendorf, 1961
- Protocupoides sharovi (Rohdendorf, 1961)
- "Protocupoides" rohdendorfi Pinto, 1987[7] – Brazil (Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Kungurian, −279.5–272.5 Ma; considered indeterminate to genus by Kirejtshuk (2020)
- Tatarocupes Ponomarenko, 2004 – Russia: Orenburg Region (Novo-Aleksandrovka), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian), −265.0–259.0 Ma.
- Tatarocupes granulatus Ponomarenko, 2004 (Type species)
- Uralocupes Ponomarenko, 1969 – Russia: Belebeevo Formation Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
- Uralocupes major (Ponomarenko, 1963) (Type species)
References
- ^ Martynov, A.V. (1933). Пермские ископаемые насекомые Архангельского края. Часть II. Сетчатокрылые, вислокрылые и жуки, с приложением описания двух новых жуков из Тихих Гор [Permian fossil insects from the Arkhangelsk District. Part II. Neuroptera, Megaloptera and Coleoptera with the description of two new beetles from Tikhie Gory] (DjVu). Trudy Paleozoologicheskogo Instituta Akademii Nauk SSSR (in Russian and English). 2: 63–96.
- S2CID 133665417.
- PMID 34747694.
- ISSN 2076-3263.
- S2CID 251171914.
- .
- ^ .