Permocupedidae

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Permocupedidae
Temporal range: Cisuralian–Anisian
Fossil and life restoration of Permocupes sojanensis
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: †Protocoleoptera
Superfamily: †Permocupedoidea
Family: †Permocupedidae
Martynov, 1933[1]
Genera

See text

Synonyms

Kaltanocoleidae Rohdendorf, 1961

Permocupedidae is a family of

xylophagous (wood eating), which is presumed to be the ancestral ecology of beetles.[3]

Taxonomy

Kirejtshuk (2020) included the following genera in an expanded (

sensu stricto) Permocupedidae and related Taldycupedidae as subfamilies Permocupedinae and Taldycupedinae respectively.[4] However, other studies have recovered Talycupedidae as more closely related to crown-group beetles than to Permocupedidae.[5]

  • Afrocupes Geertsema & van den Heever, 1996 – South Africa: Western Cape (Kenmoore Farm), Whitehill Formation, Permian, Cisuralian, Sakmarian/Artinskian, −290.1–279.5 Ma.
    • Afrocupes firmae Geertsema & van den Heever, 1996 (Type species)
  • syn Palaeocupes Rohdendorf, 1961) – Russia, Kemerovo Region
    (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma
    • Archicupes jacobsoni Rohdendorf, 1961 (Type species)
    • "Archicupes" reichardti Rohdendorf, 1961 – considered indeterminate to genus by Kirejtshuk (2020)
  • Cytocupoides Ponomarenko, 1969 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Cytocupoides longatus Ponomarenko, 1969 (Type species)
  • Cytocupes Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Cytocupes angustus Rohdendorf, 1961 (Type species)
  • Eocupes Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma).
    • Eocupes cellulosus Ponomarenko, 1969
    • Eocupes lukjanovitshi Rohdendorf, 1961 (Type species)
  • Frankencupes Ponomarenko & Bashkuev, 2018 – Germany: Gambach am Main (Lower Franconia), Röt Formation Middle Triassic, Olenekian/Anisian, −247.2–242.0 Ma.
    • Frankencupes ultimus Ponomarenko & Bashkuev, 2018 (Type species)
  • Ichthyocupes Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Kemerovo Region (Tykhta River, Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma.
    • Ichthyocupes kuznetskiensis Ponomarenko, 1969
    • Ichthyocupes skoki (Rohdendorf, 1961)
    • Ichthyocupes tyzhnovi Rohdendorf, 1961 (Type species)
  • Kaltanocoleus Rohdendorf, 1961 – Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Kaltanocoleus pospelovi Rohdendorf, 1961 (Type species)
  • Orenburg Region (Kargala), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Sweverodvinian), −265.0–259.0 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. One undescribed species of this genus was recorded from Indian Barakar Formation
    of the Lower Permian.
    • Kaltanicupes acutus Ponomarenko, 1963
    • Kaltanicupes kargalensis Ponomarenko, 1963
    • Kaltanicupes kitjakensis Ponomarenko, 1963
    • Kaltanicupes richteri Rohdendorf, 1961 (Type species)
  • Linicupes Ponomarenko, Yan & Huang, 2014[6] – China: Anhui (Houdong, Yinping Mountain, SW Chaohu City), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian/Vyatkian), −265.0–259.0 Ma.
    • Linicupes yinpinensis Ponomarenko, Yan & Huang, 2014 (Type species)
  • Maricoleus Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 – Russia, Primorsky Krai (Cape, Novosilsky, Russky Island), Lower Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
    • Maricoleus valentinae Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 (Type species)
  • Permocupes Martynov, 1933 (syn Permocupoides Martynov, 1933) – Russia: Arkhangel’sk Region (Letopala River and Soyana), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District, Tikhie Gory), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Permocupes distinctus (Martynov, 1933)
    • Permocupes grandis Ponomarenko, 1963
    • Permocupes latus Ponomarenko, 2013
    • Permocupes semenovi Martynov, 1933 (Type species)
    • Permocupes sojanensis Ponomarenko, 1963
  • Pintolla Kirejtshuk, 2020 – Brazil: Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
    • Pintolla ponomarenkoi (Pinto, 1987)[7] (Type species)
  • Vologda Region
    (Isady locality, Mutovino), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinskian/Vyatkian), −259.0–254.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma; Kemerovo Region (Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Protocupoides acer (Rohdendorf, 1961)
    • Protocupoides carinatus (Rohdendorf, 1961)
    • Protocupoides elongatus Ponomarenko, 2013
    • Protocupoides esini Ponomarenko, 2013
    • Protocupoides plavilstshikovi Rohdendorf, 1961
    • Protocupoides sharovi (Rohdendorf, 1961)
    • "Protocupoides" rohdendorfi Pinto, 1987[7] – Brazil (Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Kungurian, −279.5–272.5 Ma; considered indeterminate to genus by Kirejtshuk (2020)
  • Tatarocupes Ponomarenko, 2004 – Russia: Orenburg Region (Novo-Aleksandrovka), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian), −265.0–259.0 Ma.
    • Tatarocupes granulatus Ponomarenko, 2004 (Type species)
  • Uralocupes Ponomarenko, 1969 – Russia: Belebeevo Formation Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Uralocupes major (Ponomarenko, 1963) (Type species)

References

  1. ^ Martynov, A.V. (1933). Пермские ископаемые насекомые Архангельского края. Часть II. Сетчатокрылые, вислокрылые и жуки, с приложением описания двух новых жуков из Тихих Гор [Permian fossil insects from the Arkhangelsk District. Part II. Neuroptera, Megaloptera and Coleoptera with the description of two new beetles from Tikhie Gory] (DjVu). Trudy Paleozoologicheskogo Instituta Akademii Nauk SSSR (in Russian and English). 2: 63–96.
  2. S2CID 133665417
    .
  3. PMID 34747694
    .
  4. ISSN 2076-3263
    .
  5. S2CID 251171914
    .
  6. doi:10.1134/S0031030114010109
    .
  7. ^
    doi:10.22456/1807-9806.21678
    .
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