Phorusrhacos

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Phorusrhacos
Temporal range: Early - Mid
Ma
Reconstructed skull,
Natural History Museum, Karlsruhe
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Cariamiformes
Family: Phorusrhacidae
Subfamily:
Phorusrhacinae
Genus: Phorusrhacos
Ameghino, 1887
Type species
Phorusrhacos longissimus
Ameghino, 1887
Synonyms
Genus synonymy
  • Callornis Ameghino, 1895
  • Darwinornis Moreno & Mercerat, 1891
  • Eucallornis Ameghino, 1901
  • Liornis Ameghino, 1895
  • Owenornis Moreno & Mercerat, 1891
  • Phororhacos Ameghino, 1889
  • Stereornis Moreno & Mercerat, 1891
  • Titanornis Mercerat, 1893
Species synonymy
  • Callornis giganteus Ameghino, 1895
  • Darwinornis copei Moreno & Mercerat, 1891
  • Darwinornis socialis Moreno & Mercerat, 1891
  • Darwinornis zittelli Moreno & Mercerat, 1891
  • Eucallornis giganteus Ameghino, 1901
  • Liornis floweri Ameghino, 1895
  • Liornis minor Dolgopol de Saez, 1927
  • Mesembriornis quatrefragesi Moreno & Mercerat, 1891
  • Mesembriornis studeri Moreno & Mercerat, 1891
  • Owenornis affinis Moreno & Mercerat, 1891
  • Owenornis lydekkeri Moreno & Mercerat, 1891
  • Phororhacos longissimus Ameghino 1889
  • Phororhacos platygnathus Ameghino 1891
  • Phororhacos sehuensis Ameghino, 1891
  • Phororhacos shenensis Ameghino, 1891
  • Stereornis gaundryi Moreno & Mercerat, 1891
  • Stereornis rollieri Moreno & Mercerat, 1891
  • Titanornis mirabilis Mercerat, 1893

Phorusrhacos (

grasslands
.

Discovery and naming

Holotype mandible

Remains are known from several localities in the

edentate mammal
which he named Phorusrhacos longissimus.

The generic name is derived from Greek -φόρος, (-phoros), an element meaning "bearer" in word combinations, and ῥάκος, (rhakos), "rag" or "wrinkle", probably in reference to the wrinkled jaw surface.[3] When the original derivation was no longer understood, other translations were given, such as the literal translation of "Rag-Thief",[4] and "branch-holder" from the mistaken assumption the name had been intended to be derived from a Greek rhakis, "branch".[5] The specific name means "very long" in Latin, again in reference to the lower jaws. The holotype is the mandible, specimen MLP-118 (Museo de La Plata). In 1889 Ameghino emended the name to a more grammatically correct Phororhacos but the earlier name has priority. In 1891, it was by him recognized to be a bird.[6]

Description

Life restoration

Phorusrhacos had a skull nearly 65 centimetres (26 in) long, stood nearly 2.4 meters (7 ft 10 in) tall, and probably weighed nearly 130 kilograms (290 lb), as much as a male ostrich.[7][8] It had very strong legs, capable of running at high speed, stubby, flightless wings, a long neck, and a proportionately large head. This ended in a huge, hooked beak that could tear through flesh easily, or stab into prey. The lower jaw was smaller than the upper jaw. There were three toes on each of the feet, all of which were armed with sharp claws.[1]

Classification

Fossil skull
Life restoration by Charles R. Knight, 1901

Phorusrhacos was part of the group called the

Great American Biotic Interchange (~3 Ma). Some remains from Africa and Europe and the Paleocene of Brazil have been referred to this clade or identified as phylogenetically related to the extant South American seriemas, but these assignments remain controversial.[9]

The following cladogram follows the analysis of Degrange and colleagues, 2015:[10]

Phorusrhacidae 
 
Mesembriornithinae
 

 

Mesembriornis incertus

 

Mesembriornis milneedwardsi

 

Llallawavis scagliai

 

Procariama simplex

 
Psilopterinae
 

 

Psilopterus affinis

 

Psilopterus bachmanni

 

Psilopterus colzecus

 

Psilopterus lemoinei

 

Kelenken guillermoi

 

Devincenzia pozzi

 

Titanis walleri

 Phorusrhacos longissimus

 

Andalgalornis steulleti

 

Andrewsornis abbotti

 

Patagornis marshi

 

Physornis fortis

 

Paraphysornis brasiliensis

References

  1. ^ .
  2. ^ "Phorusrhacos". Fossilworks. Retrieved 17 December 2021.
  3. ^ Creisler, Ben. "Phorusrhacos "wrinkle bearer (jaw)": Etymology and Meaning". Dinosaur Mailing List.
  4. ^ "Phororhacos". Century Dictionary.
  5. ^ Lydekker, R. (1893). "On the extinct giant birds of Argentina". Ibis Series. 6 (5): 40–47.
  6. ^ Ameghino, F. (1891). "Mamíferos y aves fósiles argentinas. Especies nuevas, adiciones y correcciones". Revista Argentina de Historia Natural. 1: 240–259.
  7. . Retrieved 30 March 2020.
  8. doi:10.1590/S0031-10492003000400001.{{cite journal}}: CS1 maint: multiple names: authors list (link
    )
  9. S2CID 199094122. Retrieved 30 March 2020.{{cite journal}}: CS1 maint: multiple names: authors list (link
    )
  10. .