Pterodactyloidea

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Pterodactyloids
Temporal range:
Ma
Cast of a
Pterodactylus antiquus specimen, Carnegie Museum of Natural History
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Clade: Caelidracones
Suborder: Pterodactyloidea
Plieninger, 1901
Subgroups
Synonyms

Pterodactyloidea (derived from the

metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless.[2] Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tupandactylus. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs
and most marine reptiles.

"Pterodactyl" is also a common term for pterodactyloid pterosaurs, though it can also be used to refer to Pterodactylus specifically. Well-known examples of pterodactyloids include Pterodactylus, Pteranodon, and Quetzalcoatlus.

In 2014, fossils from the

teleosaurid stem-crocodilian instead of a pterosaur.[3] O'Sullivan and Martill (2018) described a partial synsacrum from the Stonesfield Slate identified as possibly pterodactyloid based on the number of incorporated sacrals although they commented that the morphology was perhaps closer to that of wukongopterids. If correctly identified, it would be the oldest pterodactyloid fossil known.[5]

Classification

Pterodactyloidea is traditionally considered to be the group of short-tailed pterosaurs with long wrists (metacarpus), compared with the relatively long tails and short wrist bones of basal pterosaurs ("rhamphorhynchoids"). In 2004, Kevin Padian formally defined Pterodactyloidea as an

apomorphy-based clade containing those species possessing a metacarpal at least 80% of the length of the humerus, homologous with that of Pterodactylus. This definition was adopted by the PhyloCode in 2020.[6]

A subgroup of pterodactyloids, called the Lophocratia, was named by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of Pterodaustro guinazui and Quetzalcoatlus northropi, and all its descendants.[7] This group was named for the presence of a head crest in most known species, though this feature has since been found in more primitive pterosaurs and was probably an ancestral feature for all pterodactyloids.[8]

There are competing theories of pterodactyloid phylogeny. Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set (Andres, 2021). This study found the two traditional groupings of ctenochasmatoids and kin as an early branching group, with all other pterodactyloids grouped into the Eupterodactyloidea.[9]

 Caelidracones 

Some studies based on a different type of analysis have found that this basic division into primitive (archaeopterodactyloid) and advanced (eupterodactyloid) species may not be correct. Beginning in 2014, Steven Vidovic and David Martill constructed an analysis in which several pterosaurs traditionally thought of as archaeopterodactyloids closely related to the ctenochasmatoids may have been more closely related to ornithocheiroids, or in some cases, fall outside both groups. The results of their updated 2017 analysis are shown below.[10]

Pterodactyloidea

Eosipterus yangi

Pterodactylus antiquus

Lophocratia
Euctenochasmatia

Diopecephalus kochi

Ctenochasmatoidea
Aurorazhdarchia

Aerodactylus scolopaciceps

Aurorazhdarchidae

Gallodactylidae

Ctenochasmatidae

Eupterodactyloidea

Altmuehlopterus ramphastinus

Dsungaripteroidea

Germanodactylus cristatus

Elanodactylus prolatus

Ornithocheiroidea
Dsungaripteromorpha

Lonchodectes compressirostris

Prejanopterus curvirostra

Kepodactylus insperatus

Dsungaripteridae

Pteranodontoidea

Ornithocheiromorpha

Hamipterus tianshanensis

Ikrandraco avatar

Azhdarchoidea

Sinopterus dongi

Tapejarinae

Eoazhdarcho liaoxiensis

Neoazhdarchia

Tapejaromorpha

Bennettazhia oregonensis

Huaxiapterus jii

Thalassodromeus sethi

Tupuxuara leonardii

Azhdarchidae

References

  1. ^ Colbert, Edwin H. (Edwin Harris); Knight, Charles Robert (1951). The dinosaur book: the ruling reptiles and their relatives. New York: McGraw-Hill. p. 153.
  2. ^ Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria) – ZooKeys
  3. ^
    PMID 24768054
    .
  4. ^ Michael O’Sullivan; David M. Martill (2018). "Pterosauria of the Great Oolite Group (Bathonian, Middle Jurassic) of Oxfordshire and Gloucestershire, England". Acta Palaeontologica Polonica. Online edition. doi:10.4202/app.00490.2018.
  5. ^ de Queiroz. K., Cantino. P. D., Gauthier. J. A. eds. (2020). Phylonyms: A Companion to the PhyloCode. CRC Press Boca Raton, FL
  6. ^ Unwin, D. M., (2003). "On the phylogeny and evolutionary history of pterosaurs." Pp. 139–190. in Buffetaut, E. & Mazin, J.-M., (eds.) (2003). Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217, London, 1–347.
  7. .
  8. ^ Andres, B. (2021) Phylogenetic systematics of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea). Journal of Vertebrate Paleontology, 41:sup1, 203-217. DOI: 10.1080/02724634.2020.1801703 https://www.tandfonline.com/doi/full/10.1080/02724634.2020.1801703