Snow leopard

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This article is about the cat. For other uses, see Snow Leopard (disambiguation).

Snow leopard
Irbis4.JPG
CITES Appendix I (CITES)[1]
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Pantherinae
Genus: Panthera
Species:
P. uncia
Binomial name
Panthera uncia
(Schreber, 1775)
SnowLeopard distribution.jpg
Distribution of the snow leopard, 2017[1]
Synonyms

The snow leopard (Panthera uncia),

subalpine zones at elevations of 3,000–4,500 m (9,800–14,800 ft), ranging from eastern Afghanistan, the Himalayas and the Tibetan Plateau to southern Siberia, Mongolia and western China
. In the northern part of its range, it also lives at lower elevations.

monotypic species. The species is widely depicted in Kyrgyz culture
.

Naming and etymology

Illustration of an 'Ounce'

The Old French word once, which was intended to be used for the Eurasian lynx (Lynx lynx), is where the Latin name uncia and the English word ounce both originate. Once is believed to have originated from a previous form of the word lynx through a process known as false splitting. The word once was originally considered to be pronounced as l'once, where l' stands for the elided form of the word la ('the') in French. Once was then understood to be the name of the animal.[2] The word panther derives from the classical Latin panthēra, itself from the ancient Greek πάνθηρ pánthēr, which was used for spotted cats.[3]

Taxonomy and evolution

Snow leopard skull in the collection of the Museum Wiesbaden

Felis uncia was the

genus name Uncia was proposed by John Edward Gray in 1854 for Asian cats with a long and thick tail.[5] Felis irbis, proposed by Christian Gottfried Ehrenberg in 1830, was a skin of a female snow leopard collected in the Altai Mountains. He also clarified that several leopard (P. pardus) skins were previously misidentified as snow leopard skins.[6] Felis uncioides proposed by Thomas Horsfield in 1855 was a snow leopard skin from Nepal in the collection of the Museum of the East India Company.[7]

Uncia uncia was used by Reginald Innes Pocock in 1930 when he reviewed skins and skulls of Panthera species from Asia. He also described morphological differences between snow leopard and leopard skins.[8] Panthera baikalensis-romanii proposed by a Russian scientist in 2000 was a dark brown snow leopard skin from the Petrovsk-Zabaykalsky District in southern Transbaikal.[9]

The snow leopard was long classified in the

monotypic genus Uncia.[10]
They were subordinated to the genus Panthera based on results of
phylogenetic studies.[11][12][13][14]

Until spring 2017, there was no evidence available for the recognition of subspecies. Results of a phylogeographic analysis indicate that three subspecies should be recognised:[15]

  • P. u. uncia in the range countries of the Pamir Mountains
  • P. u. irbis in Mongolia, and
  • P. u. uncioides in the Himalayas and Qinghai.

This view has been both contested and supported by different researchers.[16][17][18][19]

Additionally, an extinct subspecies Panthera uncia pyrenaica was described in 2022 based on material found in France.[20]

Evolution

Two cladograms proposed for Panthera. The upper cladogram is based on two studies published in 2006 and 2009,[11][21] the lower one is based on studies published in 2010 and 2011.[12][22]

Based on the phylogenetic analysis of the

excavated in western Tibet's Ngari Prefecture is the oldest known Panthera species and exhibits skull characteristics similar to the snow leopard.[23]

The

hybridised at some point in their evolution.[24]

Characteristics

Thickly furred tail

The snow leopard's fur is whitish to grey with black spots on the head and neck, with larger

muzzle is short, its forehead domed, and its nasal cavities are large. The fur is thick with hairs measuring 5 to 12 cm (2.0 to 4.7 in) in length, and its underbelly is whitish. They are stocky, short-legged, and slightly smaller than other cats of the genus Panthera, reaching a shoulder height of 56 cm (22 in), and ranging in head to body size from 75 to 150 cm (30 to 59 in). Its tail is 80 to 105 cm (31 to 41 in) long.[25] Males average 45 to 55 kg (99 to 121 lb), and females 35 to 40 kg (77 to 88 lb).[26] Occasionally, large males reaching 75 kg (165 lb) have been recorded, and small females under 25 kg (55 lb).[27]
Its
canine teeth are 28.6 mm (1.13 in) long and are more slender than those of the other Panthera species.[28]
In relation to the length of their
high-altitude hypoxia.[30]

The snow leopard shows several adaptations for living in a cold, mountainous environments. Its small rounded ears help to minimize heat loss. Their broad paws well distribute the body weight for walking on snow, and have fur on their undersides to enhance the grip on steep and unstable surfaces; they also help to minimize heat loss. Its long and flexible tail helps to balance the cat in the rocky terrain. The tail is very thick due to fat storage, and is covered in a thick layer of fur, which allows the cat to use it like a blanket to protect its face when asleep.[31][32]

The snow leopard differs from the other Panthera species by a shorter muzzle, an elevated

vocal folds provide little resistance to airflow.[34][35]

Distribution and habitat

The snow leopard is distributed from the west of

Himalayas in India, Nepal and Bhutan. In Mongolia, they inhabit the Mongolian and Gobi Altai Mountains and the Khangai Mountains. In Tibet, they occur up to the Altyn-Tagh in the north.[26][36]
They inhabit
subalpine zones at elevations of 3,000 to 4,500 m (9,800 to 14,800 ft), but also lives at lower elevations in the northern part of their range.[37]
Potential snow leopard habitat in the Indian Himalayas is estimated at less than 90,000 km2 (35,000 sq mi) in Jammu and Kashmir, Ladakh, Uttarakhand, Himachal Pradesh, Sikkim and Arunachal Pradesh, of which about 34,000 km2 (13,000 sq mi) is considered good habitat, and 14.4% is protected. In the beginning of the 1990s, the Indian snow leopard population was estimated at 200–600 individuals living across about 25 protected areas.[36]

In summer, the snow leopard usually lives above the

alpine meadows and in rocky regions at elevations of 2,700 to 6,000 m (8,900 to 19,700 ft). In winter, they descend to elevations around 1,200 to 2,000 m (3,900 to 6,600 ft). They prefer rocky, broken terrain, and can move in 85 cm (33 in) deep snow, but prefers to use existing trails made by other animals.[27]

Snow leopards were recorded by camera traps at 16 locations in northeastern Afghanistan's isolated Wakhan Corridor.[38]

Behavior and ecology

Scent rubbing
Walking on snow

The snow leopard's vocalizations include meowing, grunting, prusten and moaning. They can purr when exhaling.[25]

It is

radio-collared snow leopards largely overlapped, though they rarely met. Their individual home ranges ranged from 12 to 39 km2 (4.6 to 15.1 sq mi). Males moved between 0.5 and 5.45 km (0.31 and 3.39 mi) per day, and females between 0.2 and 2.25 km (0.12 and 1.40 mi), measured in straight lines between survey points. Since they often zigzagged in the precipitous terrain, they actually moved up to 7 km (4.3 mi) in a single night.[39]
Up to 10 individuals inhabit an area of 100 km2 (40 sq mi); in habitats with sparse prey, an area of 1,000 km2 (400 sq mi) usually supports only five individuals.[40]

A study in the Gobi Desert from 2008 to 2014 revealed that adult males used a mean home range of 144–270 km2 (56–104 sq mi), while adult females ranged in areas of 83–165 km2 (32–64 sq mi). Their home ranges overlapped less than 20%. These results indicate that about 40% of the 170 protected areas in their range countries are smaller than the home range of a single male snow leopard.[41]

Snow leopards leave

3-octanone, nonanal and indole, which possibly play a role in chemical communication.[42]

Hunting and diet

Snow leopard with a gray marmot
in Kyrgyzstan

The snow leopard is a

domestic sheep comprises less than 20% of its diet, although wild prey has been reduced and interactions with people are common.[45]

Snow leopards actively pursue prey down steep mountainsides, using the momentum of their initial leap to chase animals for up to 300 m (980 ft). They drag the prey to a safe location and consume all edible parts of the carcass. They can survive on a single Himalayan blue sheep for two weeks before hunting again, and one adult individual apparently needs 20–30 adult blue sheep per year.[1][27] Snow leopards have been recorded to hunt successfully in pairs, especially mating pairs.[48]

The snow leopard is capable of killing most animals in its range, with the probable exception of the adult male yak. They also eat a significant amount of vegetation, including grass and twigs. They have not been reported to attack humans, and is easily driven away from livestock and readily abandons kills, often without defending themselves.[27]

Reproduction and life cycle

Zurich Zoo

Snow leopards become

gestation period of 90–100 days, and the cubs are born between April and June.[27]
A
litter usually consists of two to three cubs, in exceptional cases there can be up to seven.[49]

The female gives birth in a rocky den or crevice lined with fur shed from her underside. The cubs are born blind and helpless, although already with a thick coat of fur, and weigh 320 to 567 g (11.3 to 20.0 oz). Their eyes open at around seven days, and the cubs can walk at five weeks and are fully weaned by 10 weeks. The cubs leave the den when they are around two to four months of age.[27] Three radio-collared snow leopards in Mongolia's Tost Mountains gave birth between late April and late June. Two female cubs started to part from their mothers at the age of 20 to 21 months, but reunited with them several times for a few days over a period of 4–7 months. One male cub separated from its mother at the age of about 22 months, but stayed in her vicinity for a month and moved out of his natal range at 23 months of age.[50]

The snow leopard has a generation length of eight years.[51]

Threats

Major threats to the population includes poaching and illegal trade of its skins and body parts.

Undercover operations in the country revealed an illegal trade network with links to Russia and China via Kazakhstan. The major skin trade center in the region is the city of Kashgar in Xinjiang.[52] In Tibet and Mongolia, skins are used for traditional dresses, and meat in traditional Tibetan medicine to cure kidney problems; bones are used in traditional Chinese and Mongolian medicine for treating rheumatism, injuries and pain of human bones and tendons. Between 1996 and 2002, 37 skins were found in wildlife markets and tourist shops in Mongolia.[53] Between 2003 and 2016, 710 skins were traded, of which 288 skins were confiscated. In China, an estimated 103 to 236 animals are poached every year, in Mongolia between 34 and 53, in Pakistan between 23 and 53, in India from 21 to 45, and in Tajikistan 20 to 25. In 2016, a survey of Chinese websites revealed 15 advertisements for 44 snow leopard products; the dealers offered skins, canine teeth, claws and a tongue.[54] In September 2014, nine snow leopard skins were found during a market survey in Afghanistan.[55]

alpine zone, which may reduce snow leopard habitat by an estimate 30%.[56]

Where snow leopards prey on domestic livestock, they are subject to human–wildlife conflict.[1] The loss of natural prey due to overgrazing by livestock, poaching, and defense of livestock are the major drivers for the ever decreasing snow leopard population.[27] Livestock also cause habitat degradation, which, alongside the increasing use of forests for fuel, reduces snow leopard habitat.[57]

Conservation

Global snow leopard population
Country Year Estimate
Afghanistan 2016 50–200[58]
Bhutan 2016 79–112[59]
China 2016 4,500[60]
India 2016 516–524[61]
Kazakhstan 2016 100–120[62]
Kyrgyzstan 2016 300–400[63]
Mongolia 2016 1,000[64]
Nepal 2016 301–400[65]
Pakistan 2016 250-420[66]
Russia 2016 70–90[67]
Tajikistan 2016 250–280[68]
Uzbekistan 2016 30–120[69]

The snow leopard is listed in

CITES Appendix I.[26] They have been listed as threatened with extinction in Schedule I of the Convention on the Conservation of Migratory Species of Wild Animals since 1985.[53]
Hunting snow leopards has been prohibited in Kyrgyzstan since the 1950s.
Wildlife Protection Act, 1972, and hunting is sentenced with imprisonment of 3–7 years.[61] In Nepal, they have been legally protected since 1973, with penalties of 5–15 years in prison and a fine for poaching and trading them.[70]
Since 1978, they have been listed in the Soviet Union’s Red Book and is still inscribed today in the Red Data Book of the Russian Federation as threatened with extinction. Hunting snow leopards is only permitted for the purposes of conservation and monitoring, and to eliminate a threat to the life of humans and livestock. Smuggling of snow leopard body parts is punished with imprisonment and a fine.[71] Hunting snow leopards has been prohibited in Afghanistan since 1986.[55] In China, they have been protected by law since 1989; hunting and trading snow leopards or their body parts constitute a criminal offence that is punishable by the confiscation of property, a fine and a sentence of at least 10 years in prison.[72] They have been protected in Bhutan since 1995.[59]

At the end of 2020, 35 cameras were installed on the outskirts of Almaty, Kazakhstan in hopes to catch footage of snow leopards. In November 2021, it was announced by the Russian World Wildlife Fund (WWF) that snow leopards were spotted 65 times on these cameras in the Trans-Ili Alatau mountains since the cameras were installed.[73][41][74][75][76]

Global Snow Leopard Forum

In 2013, government leaders and officials from all 12 countries encompassing the snow leopard's range (Afghanistan, Bhutan, China, India, Kazakhstan, Kyrgyzstan, Mongolia, Nepal, Pakistan, Russia, Tajikistan, and Uzbekistan) came together at the Global Snow Leopard Forum (GSLF) initiated by the then-President of Kyrgyzstan

World Wild Fund for Nature, the United States Agency for International Development, and Global Environment Facility.[77]

At the GSLF meeting, the 12 range countries signed the Bishkek Declaration, which stated: "[We] acknowledge that the snow leopard is an irreplaceable symbol of our nations' natural and cultural heritage and an indicator of the health and sustainability of mountain ecosystems; and we recognize that mountain ecosystems inhabited by snow leopards provide essential

ecosystem services
, including storing and releasing water from the origins of river systems benefitting one-third of the world’s human population; sustaining the pastoral and agricultural livelihoods of local communities which depend on biodiversity for food, fuel, fodder, and medicine; and offering inspiration, recreation, and economic opportunities."

In captivity

Snow leopard in the San Diego Zoo

The Moscow Zoo exhibited the first captive snow leopard in 1872 that had been caught in Turkestan. In Kyrgyzstan, 420 live snow leopards were caught between 1936 and 1988 and exported to zoos around the world. The Bronx Zoo housed a live snow leopard in 1903; this was the first ever specimen exhibited in a North American zoo.[78] The first captive bred snow leopard cubs were born in the 1990s in the Beijing Zoo.[52] The Snow Leopard Species Survival Plan was initiated in 1984; by 1986, American zoos held 234 individuals.[79][80]

Cultural significance

Snow leopard on the reverse of the old 10,000-Kazakhstani tenge banknote

The snow leopard is widely used in

Nur-Sultan. In Kyrgyzstan, they have been used in highly stylized form in the modern emblem of the capital Bishkek, and the same art has been integrated into the badge of the Kyrgyzstan Girl Scouts Association. They are also considered to be a sacred creature by the Kyrgyz people. A crowned snow leopard features in the arms of Shushensky District in Russia. It is the state animal of Ladakh and Himachal Pradesh in India.[81][82][83][84][85]

See also

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Further reading

External links

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