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Land slug Laevicaulis alte (Veronicelloidea)
Sea slug
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Superorder: Eupulmonata
Order: Systellommatophora

See text



The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005).[1]

They are

Onchidioidea are primarily marine (except for five land-dwelling or freshwater species),[2] whereas slugs in the superfamily Veronicelloidea
are primarily terrestrial.


No species in this clade have shells as adults.[2][3] These slugs are distinguished by the location of the anus at the rear of the body.[4]

Although neither superfamily bears shells as adults,[3] the Onchidioidea do possess a vestigial, non-mineralized shell sac[5] and possess a larval shell.[6] It is not known whether or not the veronicellids bear a larval shell.[7]


According to the previous

suborder or an order.[8]

The following two superfamilies and families have been recognized in the taxonomy of Bouchet & Rocroi (2005):


  1. ^ MolluscaBase eds. (2020). MolluscaBase. Stylommatophora. Accessed through: World Register of Marine Species at: on 2020-05-02
  2. ^ .
  3. ^ a b Schilthuizen M.; Thome J. W. (2008). "Valiguna flava (Heynemann, 1885) from Indonesia and Malaysia: Redescription and Comparison with Valiguna siamensis (Martens, 1867) (Gastropoda: Soleolifera: Veronicellidae)". Veliger. 50 (3): 163–170.
  4. .
  5. ^ TILLIER, S (1983). "A new mountain Platevindex from Philippine Islands (Pulmonata: Onchidiidae)". Journal of Molluscan Studies. 49 (supp12A). Malacological Soc London: 198.
  6. .
  7. .
  8. ^ "Systellommatophora". Integrated Taxonomic Information System.