Esociformes Species List

Umbra longidorsalis †

Proumbra irthyshensis †

Umbra perpusilla † (syn. Palaeoesox perpusillus)

Novumbra oregonensis †

Dallia admirabilis

Dallia delicatissima

Palaeoesox fritzschei † (type species and therefore senior to P. weileri)

Umbra prochazkai †

Umbra oderiensis †

Boltyshia brevicauda †

Boltyshia truncata †

Palaeoesox weileri † (probable syn. Palaeoesox fritzschei) (previously named Umbra weileri)

Estesesox foxi † (some fossil material misdescribed as Platacodon nanus in the family Sciaenidae; the family is not within Esociformes)

Oldmanesox canadensis †

Esox tiemani †

Esox longkouensis †

Esox columbianus †

Esox kronneri †

Esox cisalpinus

Esox nogaicus †

Esox sibiricus †

Esox moldavicus †

Palaeoesox praekrameri † (previously named in the genus Umbra)

Palaeoesox acutirostris † (previously named Mikroumbra acutirostris, then later Palaeumbra acutirostris (? May be valid name, just with the same species name as Palaeoesox acutirostris))

Palaeumbra acutirostris † (may be synonym of Palaeoesox acutirostris?)

Palaeoesox perpusillus † (described as Lebias perpusillus) (correct name for syn. Umbra perpusilla (?))

Palaeoesox densus †

Umbra valida † (possibly in the genus Palaeoesox)

Palaeoesox oligocenicus †

Palaeoesox longirostratus †

Esteesox sp. † (species name not mentioned in genetic study source) (probable typo of Estesesox)

Esox primaevus †

Esox papyraceus †

Esox waltschanus †

Esox desiructus †

Esox lepidotus †

Palaeoesox goellheimensis †

Palaeumbra moguntina †

"genus Umbridarum" crassus †

Umbra n. sp. † (unnamed)

Genera

Esox

Umbra (the oldest known representative of the genus Umbra (U. prochazkai), is dated from the Late Oligocene)

Dallia

Novumbra

Oldmanesox † (more closely related to Esox than Estesesox

Proumbra † (intermediate between Umbra and Palaeoesox)

Palaeoesox †

Boltyshia †

Estesesox †

Palaeumbra † (genus may be invalid if Palaeumbra acutirostris is a synonym of Palaeoesox acutirostris)

"Umbridarum" † (invalid genus?)

Esteesox † (probable typo of Estesesox)

Families

Umbridae (paraphyletic; Umbra, Novumbra, Dallia) (recently proposed to only include Umbra) (Boltyshia is a basal member of the Umbridae lineage(?)) (Palaeoesox “ belongs clearly to the lineage leading to the genus Umbra”, though I’m unsure if it is in its own family (Palaeoesocidae) or is in Umbridae) (genetic study source recommends including Boltyshia, Palaeoesox, Proumbra, and Umbra in Umbridae)

Esocidae (Esox) (genetic study source asserts this family should include Novumbra and Dallia) (Mesozoic fossil study places Estesesox and Oldmanesox within Esocidae)

Paleoesocidae † (Palaeoesox(?), Boltyshia) (incartae sedis within Umbridae)

Sources

Translate to English

Kovalchuk, O., & Rekovets, L. EWOLUCJA I PALEOEKOLOGIA FAUNY RYB SŁODKOWODNYCH W GÓRNYM PALEOCENIE–DOLNYM EOCENIE EUROPY WSCHODNIEJ 1 EVOLUTION AND PALEOECOLOGY OF FRESHWATER FISH FAUNA IN THE UPPER PALEOCENE–LOWER EOCENE. ZESZYTY NAUKOWE UNiWErSYTETU prZYrOdNicZEgO, 27. [Mentions Boltyshia brevicauda]

https://www.persee.fr/doc/geolm_0397-2844_1978_num_5_2_1047 [Mentions Esox primaevus, Esox papyraceus, Esox waltschanus, Esox desiructus, and Esox lepidotus]

https://www.researchgate.net/profile/Bettina-Reichenbacher/publication/287699893_Biostratigraphy_and_palaeoecology_of_fish_otoliths_in_the_Upper_Cerithium_Beds_Lower_Miocene_near_Gollheim_Mainz_Basin/links/56839f0908ae051f9af02bd6/Biostratigraphy-and-palaeoecology-of-fish-otoliths-in-the-Upper-Cerithium-Beds-Lower-Miocene-near-Gollheim-Mainz-Basin.pdf [Mentions "genus Umbridarum" crassus, Palaeoesox fritzschei, Palaeumbra moguntina, Palaeoesox goelheimensis, Palaeoesox acutirostris]

English

Gaudant, J. (2012). An attempt at the palaeontological history of the European mudminnows (Pisces, Teleostei, Umbridae). Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen, 93-109. [Mentions Boltyshia brevicauda, Palaeoesox fritzschei, Umbra prochazkai, Palaeumbra acutirostris, Palaeoesox perpusillus, Palaeoesox praekrameri, Umbra prochazkai, Proumbra irtyshensis, Umbra n. sp., Palaeoesox densus, Palaeoesox acutirostris]

Matschiner, M., Musilová, Z., Barth, J. M., Starostová, Z., Salzburger, W., Steel, M., & Bouckaert, R. (2016). Bayesian node dating based on probabilities of fossil sampling supports trans-Atlantic dispersal of cichlid fishes. BioRxiv, 038455. [Supplementary text mentions age of fossil Esociformes]

Marić, S., Stanković, D., Wanzenböck, J. et al. Phylogeography and population genetics of the European mudminnow (Umbra krameri) with a time-calibrated phylogeny for the family Umbridae. Hydrobiologia 792, 151–168 (2017). https://doi.org/10.1007/s10750-016-3051-9 [Extant Umbra spp. split based on genetic research; mentions Esteesox, taxonomy of other extinct species.]

Wilson, Mark & Brinkman, Donald & Neuman, Andrew. (1992). Cretaceous Esocoidei (Teleostei): Early radiation of the pikes in North American fresh waters. Journal of Paleontology. 66. 839-846. 10.1017/S0022336000020849. [Metions Estesesox, Oldmanesox. Mudminnows (which the study classifies in the paraphyletic sense of Umbridae) may be descended from ancestors that had depressible teeth like those of esocids. This is still likely given the reported presence of depressible teeth in Palaeoesox in this study.]

Reichenbacher, B. (1999). Preliminary otolith-zonation in continental Tertiary deposits of the Paratethys and adjacent areas. Neues Jahrbuch Geologie Paläontologie Abhandlungen, 214(3), 375-390. [Mentions Palaeoesox oligocenicus, Palaeoesox longirostratus]

Cladograms

Extant Esociformes Species

Esociformes Genera

^§Estesesox and Oldmanesox are assigned to Esocoidea, however taxonomic placement of Esocoidea is messy at best.