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Period (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. Early species had long, fully toothed jaws and long tails, while later forms had a highly reduced tail, and some lacked teeth. Many sported furry coats made up of hair-like filaments known as pycnofibres, which covered their bodies and parts of their wings. Pterosaurs spanned a wide range of adult sizes, from the very small Nemicolopterus to the largest known flying creatures of all time, including Quetzalcoatlus and Hatzegopteryx