Camarillasaurus

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Camarillasaurus
Temporal range:
Ma
Holotype
fossils
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: Spinosauridae (?)
Genus: Camarillasaurus
Sánchez-Hernández & Benton,
2014
Type species
Camarillasaurus cirugedae
Sánchez-Hernández & Benton, 2014

Camarillasaurus (meaning "

Teruel Province, in what is now northeastern Spain. Described in 2014, it was originally identified as a ceratosaurian theropod, but later studies suggested affinities to the Spinosauridae. If it does represent a spinosaur, Camarillasaurus would be one of five spinosaurid taxa known from the Iberian peninsula, the others being Iberospinus, Protathlitis, Riojavenatrix, and Vallibonavenatrix.[1]

Discovery and naming

Holotype sacrum and centrum

Fossils of Camarillasaurus were discovered in the

Michael J. Benton. The generic name, "'Camarillasaurus", combines a reference to the geologic formation in which the holotype was found with the Greek "sauros", meaning "lizard". The specific name, "cirugedae", honors Pedro Cirugeda Buj, the discoverer of the holotype specimen.[2]

Classification

Life restoration
as a spinosaur

Camarillasaurus has a complicated taxonomic history. Having originally been described as a ceratosaur,[2] most later studies have recovered it in various positions within the Spinosauridae.[3][1]

In their 2014

phylogenetic position using a ceratosaurian dataset, and recovered the following results:[2]

Ceratosauria

However, in an

sister taxon to Camarillasaurus, or synonymous with it.[3]

In contrast, in a 2024 review of theropod fossils from India focusing on potential

noasaurid bones, Mohabey et al. included Camarillasaurus in a phylogenetic analysis, recovering it within the ceratosaurian clade Noasauridae, as the sister taxon to a noasaurid from the Tiourarén Formation of Niger. However, their analysis did not include any spinosaurids.[7]

In the 2024 description of the Iberian spinosaurid Riojavenatrix by Isasmendi et al., Camarillasaurus was once again recovered in the Spinosauridae, either as the basalmost spinosaurine or a basal spinosaurid as the sister to Baryonychinae and Spinosaurinae. The results of their phylogenetic analyses are shown in the cladogram below:[1]

References