Coelophysidae

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Coelophysids
Temporal range:
Ma
Skull comparison of Coelophysidae (
"Syntarsus" kayentakatae
at the bottom right)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Superfamily: Coelophysoidea
Family: Coelophysidae
Nopcsa, 1923
Type species
Coelurus bauri
, 1887
Genera
Synonyms
  • Procompsognathidae Nopcsa, 1923
  • Segisauridae Camp, 1936

Coelophysidae is a family of primitive carnivorous

theropod dinosaurs. Most species were relatively small in size. The family flourished in the Late Triassic and Early Jurassic periods, and has been found on numerous continents. Many members of Coelophysidae are characterized by long, slender skulls and light skeletons built for speed.[2] One member genus, Coelophysis, displays the earliest known furcula in a dinosaur.[3]

Under

type specimen was destroyed in a fire and can no longer be compared to new finds.[5]

Classification

Characteristics

Coelophysids are characterized by slender, skinny builds and long, narrow skulls with large fenestrae to allow for a lighter skull.[6] They are fairly primitive theropods, and so have fairly basal characteristics, such as hollow air sacs in the cervical vertebrae and obligate bipedalism.[6] Their slender builds allowed them to be fast and agile runners. All known members of Coelophysidae are carnivores. One species, Coelophysis bauri has the oldest known furcula (wishbone) of any dinosaur.[3]

It has also been speculated that some species within Coelophysidae, namely Coelophysis bauri, displayed cannibalism, although the fossil evidence behind these claims has been heavily debated (Rinehart et al., 2009; Gay, 2002; Gay, 2010).[7][8][9]

Phylogenetics

Coelophysidae is part of the larger superfamily of Coelophysoidea, which contains Dilophosauridae, Liliensternus, and Zupaysaurus in addition to Coelophysidae.[2][10][11] Coelophysoidea, in turn, is part of the larger clade of Neotheropoda.[2]

The cladogram below was recovered in a study by Matthew T. Carrano, John R. Hutchinson and Scott D. Sampson, 2005.[12]

The cladogram below follows the topology from a 2011 analysis by paleontologists Martin D. Ezcurra and Stephen L. Brusatte, modified with additional data by You Hai-Lu and colleagues in 2014.[1][13]

"Syntarsus" kayentakatae

Panguraptor lufengensis

Coelophysis bauri

Coelophysis rhodesiensis

Camposaurus arizonensis

This cladogram follows the 2017 analysis by Ezcurra,[14] with added results of the Martinez & Apaldetti analysis from their description of Lucianovenator.[15]

"Syntarsus" kayentakatae

Panguraptor lufengensis

Powellvenator podocitus

Procompsognathus triassicus

Coelophysis bauri

Lepidus praecisio

Segisaurus halli

Megapnosaurus rhodesiensis

Lucianovenator bonoi

Camposaurus arizonensis

Biogeography

Fossils of members of Coelophysidae have been found across many continents, including North America, South America, Europe, Asia, and Africa. Powellvenator podocitus was discovered in Northwestern Argentina.[16] Procompsognathus triassicus was discovered in Germany, and Camposaurus arizonensis is from Arizona in North America.[17][13] No coelophysid fossils were known from Asia until the discovery of Panguraptor lufengensis in 2014 in the Yunnan Province of China.[1] The genus Coelophysis has been found in North America, South Africa, and Zimbabwe.[18]

See also

References

  1. ^
    PMID 25544219
    .
  2. ^ .
  3. ^ .
  4. ^ Tykoski, Ronald S. (2005). Anatomy, Ontogeny, and Phylogeny of Coelophysoid Theropods (PhD). University of Texas at Austin.
  5. ^ Sereno, P. (1999). "Taxon Search: Coelophysidae Archived 2007-10-07 at the Wayback Machine". Accessed 2009-09-02.
  6. ^
    PMID 20007898
    .
  7. ^ Rinehart, L.F.; Lucas, S.G.; Heckert, A.B.; Spielmann, J.A.; Celesky, M.D. (2009). "The paleobiology of Coelophysis bauri (Cope) from the Upper Triassic (Apachean) Whitaker quarry, New Mexico, with detailed analysis of a single quarry block". New Mexico Museum of Natural History & Science, a division of the Department of Cultural Affairs Bulletin. 45: 260.
  8. ^ Gay, R.J. (2002). "The myth of cannibalism in Coelophysis bauri". Journal of Vertebrate Paleontology. 22 (3): 57A.
  9. .
  10. .
  11. .
  12. ^ .
  13. ISSN
    0002-7014.
  14. doi
    :10.5710/AMGH.09.04.2017.3065.
  15. .
  16. .
  17. ^ Bristowe, A.; Raath, M.A. (2004). "A juvenile coelophysoid skull from the Early Jurassic of Zimbabwe, and the synonymy of Coelophysis and Syntarsus". Palaeontologica Africana. 40: 31–41.