Dipper

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For other uses, see Dipper (disambiguation).

Dippers
American dipper (Cinclus mexicanus)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Superfamily: Muscicapoidea
Family: Cinclidae
Sundevall, 1836
Genus: Cinclus
Borkhausen, 1797
Type species
Cinclus hydrophilus[1]
Borkhausen, 1797
Distribution map
  White-throated dipper
  Brown dipper
  American dipper
  White-capped dipper
  Rufous-throated dipper
Phylogeny of the dippers[2]

Dippers are members of the genus Cinclus in the bird family Cinclidae, so-called because of their bobbing or dipping movements. They are unique among passerines for their ability to dive and swim underwater.

Taxonomy

The genus Cinclus was introduced by the German naturalist Moritz Balthasar Borkhausen in 1797 with the white-throated dipper (Cinclus cinclus) as the type species.[3][4] The name cinclus is from the Ancient Greek word kinklos that was used to describe small tail-wagging birds that resided near water.[5]

Cinclus is the only genus in the

Old English osle). Ouzel also survives as the name of a relative of the blackbird, the ring ouzel.[6]

The genus contains five species:[7]

A 2002

sister species as are the South American white-capped dipper and rufous-throated dipper. The study also showed that the dipper family, Cinclidae, is most closely related to the thrush family, Turdidae.[2]

Description

White-throated dipper (C. cinclus)

Dippers are small, chunky, stout, short-tailed, short-winged, strong-legged birds. The different species are generally dark brown (sometimes nearly black), or brown and white in colour, apart from the rufous-throated dipper, which is brown with a reddish-brown throat patch. Sizes range from 14–22 cm (5.5–8.7 in) in length and 40–90 g (1.4–3.2 oz) in weight, with males larger than females. Their short wings give them a distinctive whirring flight.[8][9][10] They have a characteristic bobbing motion when perched beside the water, giving them their name. While under water, they are covered by a thin, silvery film of air, due to small bubbles being trapped on the surface of the plumage.[9]

Distribution and habitat

Dippers are found in suitable freshwater habitats in the highlands of the Americas, Europe and Asia. In Africa they are only found in the Atlas Mountains of Morocco. They inhabit the banks of fast-moving upland rivers with cold, clear waters, though, outside the breeding season, they may visit lake shores and sea coasts.[9]

Adaptations

Unlike many water birds, dippers are generally similar in form to many terrestrial birds (for example, they do not have

preen gland for waterproofing their feathers. Relatively long legs and sharp claws enable them to hold on to rocks in swift water. Their eyes have well-developed focus muscles that can change the curvature of the lens to enhance underwater vision.[12] They have nasal flaps to prevent water entering their nostrils.[13]

The high

haemoglobin concentration in their blood gives them a capacity to store oxygen greater than that of other birds, allowing them to remain underwater for 30 seconds or more,[9] whilst their basal metabolic rate is approximately one-third slower than typical terrestrial passerines of similar mass.[14] One small population wintering at a hot spring in Suntar-Khayata Mountains of Siberia feeds underwater when air temperatures drop below −55 °C (−67 °F).[15]

Behaviour

Food

Dippers

Molluscs and crustaceans are also consumed, especially in winter when insect larvae are less available.[9]

Breeding

Linear breeding territories are established by pairs of dippers along suitable rivers, and maintained against incursion by other dippers. Within their territory the pair must have a good nest site and roost sites, but the main factor affecting the length of the territory is the availability of sufficient food to feed themselves and their broods. Consequently, the length of a territory may vary from about 300 metres (1,000 feet) to over 2,500 metres (8,200 feet).[9]

Dipper nests are usually large, round, domed structures made of moss, with an internal cup of grass and rootlets, and a side entrance hole. They are often built in confined spaces over, or close to, running water. The site may be on a ledge or bank, in a crevice or drainpipe, or beneath a bridge. Tree sites are rare.[9]

The usual clutch-size of the three northern dipper species is four or five; those of the South American species is not well known, though some evidence suggests that of the rufous-throated dipper is two.

altricial young which are brooded by the female alone for the next 12 to 13 days. The nestlings are fed by both parents and the whole fledging period is about 20–24 days. Young dippers usually become independent of their parents within a couple of weeks of leaving the nest. Dippers may raise second broods if conditions allow.[9] The maximum recorded age from ring-recovery data of a white-throated dipper is 10 years and 7 months for a bird ringed in Finland.[18] The maximum age for an American dipper is 8 years and 1 month for a bird ringed and recovered in South Dakota.[19]

Communication

Dippers' calls are loud and high-pitched, being similar to calls made by other birds on fast rivers; the

kHz, well above the torrent noise frequency of maximum 2 kHz.[20] Dippers also communicate visually by their characteristic dipping or bobbing movements, as well as by blinking rapidly to expose the white feathers on their upper eyelids as a series of white flashes in courtship and threat displays.[12]

Conservation

The rufous-throated dipper is considered Vulnerable by the IUCN

Dippers are completely dependent on fast-flowing rivers with clear water, accessible food and secure nest-sites. They may be threatened by anything that affects these needs such as water pollution, acidification and turbidity caused by erosion. River regulation through the creation of dams and reservoirs, as well as channelization, can degrade and destroy dipper habitat.[9]

Dippers are also sometimes

salmonids.[9]

Despite threats to local populations, the conservation status of most dipper species is considered to be of

least concern. The one exception, the rufous-throated dipper, is classified as vulnerable because of its small, fragmented and declining population which is threatened, especially in Argentina, by changes in river management.[21]

References

  1. ^ "Cinclidae". aviansystematics.org. The Trust for Avian Systematics. Retrieved 2023-07-15.
  2. ^
    doi:10.1046/j.1474-919X.2002.00084.x
    .
  3. ^ Borkhausen (1797). Deutsche Fauna, oder, Kurzgefasste Naturgeschichte der Thiere Deutschlands. Erster Theil, Saugthiere und Vögel (in German). Frankfurt am Main: Varrentrapp und Wenner. p. 300.
  4. ^ Mayr, Ernst; Greenway, James C. Jr, eds. (1960). Check-list of Birds of the World. Vol. 9. Cambridge, Massachusetts: Museum of Comparative Zoology. p. 374.
  5. ISBN 978-1-4081-2501-4
    .
  6. ^ "ouzel". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
  7. ^ Gill, Frank; Donsker, David, eds. (2019). "Dippers, leafbirds, flowerpeckers, sunbirds". World Bird List Version 9.1. International Ornithologists' Union. Retrieved 9 February 2019.
  8. ISBN 978-1-4067-4576-4
    .
  9. ^
    ISBN 0-85661-093-3
    .
  10. ^ Robbins, C.S.; Bruun, B.; & Zim, H.S. (1966). Birds of North America. Western Publishing Company: New York.
  11. ^ "Country diary: it looks like a songbird, but the dipper is aquatic to its bones". www.theguardian.com. 7 April 2018. Retrieved 22 October 2023.
  12. ^
    S2CID 7227306
    .
  13. S2CID 242827109
    . Retrieved 11 February 2019. The text is identical to Volume 10 of the print edition published in 2005.
  14. doi:10.1016/0010-406X(70)91009-1
    .
  15. S2CID 91058122
    .
  16. ^ Tyler, S.J. (1994). "The Yungas of Argentina: in search of Rufous-throated Dippers Cinclus schulzi" (PDF). Cotinga. 2: 38–41.
  17. ^ Salvador, S.; Narosky, S.; Fraga, R. (1986). "First description of the nest and eggs of the red-throated dipper in northwestern Argentina". Gerfaut. 76: 63–66.
  18. ^ "European Longevity Records". Euring. Retrieved 13 February 2019.
  19. ^ "Longevity Records of North American Birds". United States Geological Survey. Retrieved 13 February 2019.
  20. ^ J., Martens; Geduldig, G. (1990). "Acoustic adaptations of birds living close to Himalayan torrents". Proc. Int. 100 DO-G Meeting. Bonn: Current Topics Avian Biol. pp. 123–131.
  21. IUCN Red List of Threatened Species. International Union for Conservation of Nature
    . Retrieved 13 February 2019.

Further reading

External links
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