Mixosaurus

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Mixosaurus
Temporal range: Triassic
(Anisian to Ladinian, about 247–242 Mya)[1]
Dorsal view of fossil showing top of skull
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: †Ichthyosauria
Family: †Mixosauridae
Subfamily: †
Mixosaurinae
Genus: †Mixosaurus
Baur, 1887
Species
  • M. cornalianus (Bassani, 1886)
  • M. kuhnschnyderi (Brinkmann, 1998)
  • M. xindianensis?[2] Chen & Cheng, 2010

Mixosaurus is an

ichthyosaur. Its fossils have been found near the Italy–Switzerland border and in South China.[1]

The genus was named in 1887 by

transitional form between the eel-shaped ichthyosaurs such as Cymbospondylus and the later dolphin-shaped ichthyosaurs, such as Ichthyosaurus. Baur named Mixosaurus as a new genus because its forefin was sufficiently different from that of Ichthyosaurus.[3]

Mixosaurus includes three species.[2][1] Previously this number was bigger, and Mixosaurus was considered as the most common genus of Triassic ichthyosaurs,[3] whose fossils have been found all over the world, including China, Timor, Indonesia, Italy, Spitsbergen, Svalbard, Canada, as well as Alaska and Nevada in the US.

Description

Mixosaurus with a human hand to scale

Mixosaurus was a small ichthyosaur, measuring 73–100 cm (2.40–3.28 ft) long and weighing 2.2–5.7 kg (4.9–12.6 lb).[4] The largest specimens would have measured over 2 m (6.6 ft) in length.[5] It possessed a long tail with a low fin, suggesting it could have been a slow swimmer, but also possessed a dorsal fin for stability in the water.[6] The paddle-like limbs were made up of five toes each, unlike the three toes found in later Ichthyosaurs. Noteworthy, however, is that each toe had more individual bones than is usual in reptiles, and the front limbs were longer than the back limbs, both adaptations typical of later ichthyosaurs. The jaws were narrow, with several sharp teeth, that would have been ideal for catching fish.[7] They had relatively large skulls compared to their bodies, unlike the basal ichthyosaurs, but resembled fish-shaped ichthyosaurs that appeared later. They had around 50 vertebrae in front of the pelvic girdle, around twice as many as terrestrial diapsids.[8] Recent studies suggest that genus Mixosaurus may have lived near shore or in a shelf-like habitat as it possesses more compact

spongy bone within its long bones than other Ichthyosaurs.[9]

Species

There are three species of Mixosaurus currently recognised, Mixosaurus cornalianus, Mixosaurus xindianensis and Mixosaurus kuhnschnyderi.[10][2][11] They share many similar characteristics throughout the cranial and post cranial with the main differences morphology occurring in the dental region. Examples of the dental variation are the extent of the dental groove in the upper jaw, the shape and size of the teeth and the number of rows of teeth [3]

Previous authors assigned other species to the genus, including M. atavus (Quenstedt, 1852), M. callawayi Schmitz et al., 2004, M. panxianensis Jiang et al., 2006 and M. yangjuanensis Liu & Yin, 2008. These are now included in Contectopalatus, Phalarodon, Barracudasauroides and Nothosaurus, respectively.[12]

Mixosaurus species declared as nomen dubium, meaning the description was insufficient to fully classify them as a species, are M. maotaiensis, M. helveticus, M. timorensis, M. major, and M. nordenskioeldii.[10][13][14]

Mixosaurus cornalianus

Skeletal reconstruction of M. cornalianus showing soft tissue outline
Life restoration of M. cornalianus

Many specimens of Mixosaurus cornalianus have been found from the Middle Triassic of Monte San Giorgio and the Tessin areas on the border of Italy and Switzerland.

Mixosaurus cornalianus is the only Triassic ichthyosaur for which completely articulated skeletons have been found. Many specimens have been collected but Mixosaurus cornalianus is not well studied, this is because all of the known specimens have been compressed during the preservation process.

Mixosaurus cornalianus has a

fenestra. This indicates that it had exceptionally strong jaw muscles.[3]

Mixosaurus panxianensis

Mixosaurus panxianensis was discovered in the Middle Triassic of the

marls
.

The specimens found have important Mixosaurid characteristics such as a long sagittal crest along the top of the

quadratojugal
.

Articulated skeletons have been found and the centra of the

vertebrae are higher than they are long. This is evidence for its transitional position between basal early Triassic ichthyosaurs and more derived Jurassic and Cretaceous species; who have disc shaped circular centra of the vertebrae.[10] This species has been moved to its own genus, Barracudasauroides.[12][11]

Classification

M. cornalianus in Milan
M. cornalianus

In recent years the

phylogeny of mixosaurid ichthyosaurs has been a controversial topic. Most recently, Mixosauridae has been separated into Mixosaurinae and the sister group Phalarondontinae. Mixosaurus contains M. cornalianus, M. kuhnschnyderi and M. xindianensis, Barracudasauroides contains B. panxianensis, Phalarodon contains P. fraasi and P. callawayi, and Contectopalatus contains C. atavus.[2][12][11]

Mixosaurids are characterised by a relatively short and wide humerus and Phalarodon are characterised by the lack of a dental groove in the upper jaw. Phalarodon fossils are found in every major Mixosaur locality.

It was suggested that Tholodus schmidi should be included in Mixosauridae but only dental material has been found so it is difficult to assign it to a genus.[10]

See also

References

  1. ^
    S2CID 85621052
    .
  2. ^
    doi:10.1111/pala.12021
    .
  3. ^
    S2CID 130661969
    .
  4. S2CID 245444783
    .
  5. doi:10.1080/02724634.1999.10011160
    .
  6. S2CID 222285117
    .
  7. ISBN 978-1-84028-152-1
    .
  8. S2CID 4323012
    .
  9. PMID 24752508
    .
  10. ^
    S2CID 55171917
    .
  11. ^
    S2CID 219078178
    .
  12. ^ a b c Michael W. Maisch (2010). "Phylogeny, systematics, and origin of the Ichthyosauria – the state of the art" (PDF). Palaeodiversity. 3: 151–214.
  13. S2CID 84214133
    .
  14. ^ Prof. Dr. H.D Sues, ed. (2003). Handbook of Paeoherpetology, Part 8, Ichthyopterygia. Munchen: Friedrich Pfeil. p. 175.
  15. S2CID 90912678
    .
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