Mesotherium

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Mesotherium
Temporal range: Early Pleistocene–Middle Pleistocene
Skull of Mesotherium cristatum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Notoungulata
Family: Mesotheriidae
Subfamily:
Mesotheriinae
Genus: Mesotherium
Serres, 1867
Type species
Mesotherium cristatum
Serres, 1857
Synonyms
  • Mesotherium:
    • Typotherium
  • Mesotherium cristatum
    • M. hystatum Ameghino 1904
    • M. maendrum Ameghino 1887
    • M. pachygnathum Gervais & Ameghino 1880

Mesotherium ("Middle Beast")

Middle Pleistocene.[2]

Etymology

Serres named Mesotherium so due to his belief that it was an intermediate between rodents and pachyderms (or ungulates), due to its large upper

incisors, and its size and proportions.[1] "Serres—by a happy inspiration proposed calling it Mesotherium—as being a common centre towards which all mammalia got happily confounded," Hugh Falconer wrote Darwin in April 1863.[3]
"Bravard sent it home under the name of Typotherium as being the central type from which all mammals diverged." It was Serres' view that there was only one underlying animal type.

Despite Serres' having officially named the genus Mesotherium, it was known from the late 19th century to the early 20th century under the name "Typotherium", given it by the French palaeontologist living in Argentina, Auguste Bravard;[4] under this name, Bravard sent the skull he found to Paris, which led to the family being named "Typotheriidae", and served as the basis for the order Typotheria.[1] As the name Mesotherium had been published earlier in the same year as Typotherium, Mesotherium was declared the valid name of the genus, (Simpson, 1980[4]) and Mesotheriidae the valid name of the family.[1] Nevertheless, as the rules do not apply to anything above the family, the name of the order Typotheria is still in use, but refers to a wider range of rodent-like notoungulates.[1]

Description

Restoration of the skull and jaws in lateral view

Mesotherium was likely the size of a small sheep, and weighed around 55 kilograms (121 lb),[1] or possibly up to 100 kilograms (220 lb), making it the largest known member of Typotheria.[5] Like most rodents, it had superficially long upper incisors, which met at the tips,[1] however, it had enamel on both the labial and lingual surfaces of the incisors, while rodents only have enamel on the labial surface.[1] The lower incisors of Mesotherium were reminiscent of those of a rabbit's.[1]

The ankle joint of Mesotherium was made up of a "ball-and-socket" arrangement between the

great toe.[6]

Mesotherium was likely fossorial, in that it dug, possibly to find food,[1][6] or to construct dens.[5]

Ecology

Mesotherium is suggested to have been a mixed feeder that likely consumed a large quantity of grasses, with its wide snout allowing for bulk feeding. In comparison to earlier mesotheriids, it probably consumed less hard food items.[5]

Chronology

The oldest known specimens of the genus date to the Early Pleistocene. The youngest known specimen of the genus dates to around 220,000 years ago during the late Middle Pleistocene.[7]

References

  1. ^ a b c d e f g h i j "D. Croft SA Mammals: Mesotheriidae". D. Croft. 2007-01-02. Archived from D. Croft SA Mammals: Mesotheriidae the original on 2009-12-24. Retrieved 2008-06-05. {{cite web}}: Check |url= value (help)
  2. S2CID 249531139
    .
  3. ^ Darwin Correspondence Project: Letter dated 20 April [1863][permanent dead link],
  4. ^ Retrieved on 2008-05-02
  5. ^ .
  6. ^ a b c "Analysis of function in the absence of extant functional homologues: a case study using mesotheriid notoungulates (Mammalia)". Bruce Shockey, Darin Croft, and Frederico Anaya. Spring 2007. Retrieved 2008-07-10.[permanent dead link]
  7. S2CID 254913691
    .

Further reading

External links