Australopithecus garhi

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Australopithecus garhi
Temporal range: Gelasian, 2.6–2.5 Ma
Reconstruction of the skull at the National Museum of Ethiopia
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Suborder: Haplorhini
Infraorder: Simiiformes
Family: Hominidae
Subfamily: Homininae
Tribe: Hominini
Genus: Australopithecus
Species:
A. garhi
Binomial name
Australopithecus garhi
Asfaw et al., 1997

Australopithecus garhi is a species of

arboreality); and it is possible that, though unclear if, males were larger than females (exhibited sexual dimorphism
). One individual, presumed female based on size, may have been 140 cm (4 ft 7 in) tall.

A. garhi is the first pre-Homo hominin postulated to have manufactured tools—using them in butchering—and may be counted among a growing body of evidence for pre-Homo stone tool industries (the ability to manufacture tools was previously believed to have separated Homo from predecessors.) A. garhi possibly produced the Oldowan industry which was previously considered to have been invented by the later H. habilis, though this may have instead been produced by contemporary Homo.

Anatomy

Like other australopithecines, A. garhi had a brain volume of about 450 cc (27 cu in), a

post-canine megadontia), similar to or greater than those of other australopithecines and of the large-toothed Paranthropus robustus.[1]

Like the earlier A. afarensis from the same region, A. garhi had a humanlike humerus to femur ratio, and an apelike brachial index (lower to upper arm ratio) as well as curved

The BOU-VP-35/1 humerus specimen is notably larger than the humerus of the BOU-VP-12/1 specimen, which could potentially indicate size-specific sexual dimorphism with males larger than females to a similar degree to what is postulated in A. afarensis, but it is unclear if this does not represent normal size variation of the same sex as this is based on only two specimens. Nonetheless, on the basis of size, BOU-VP-12/130 is considered male and BOU-VP-17/1 female. Contemporary hominins from Kenya are about the same size as A. garhi.[1] BOU-VP-17/1 may have been about 140 cm (4 ft 7 in) tall.[3]

Australopithecus are thought to have had fast, apelike growth rates, lacking an extended childhood typical of modern humans. However, the legs of A. garhi are elongated, unlike those of other Australopithecus, and, in humans, elongated limbs develop during the delayed adolescent growth spurt. This could mean that A. garhi, compared to other Australopithecus, either had a slower overall growth rate, or a more rapid leg growth rate.[2]

Taxonomy

The Ethiopian Australopithecus garhi was first described in 1999 by palaeoanthropologists

locality of the formation, and American palaeoanthropologist David DeGusta discovered a humerus (BOU-VP-35/1) 1 km (0.62 mi) north of BOU-VP-17/1. However, BOU-VP-11, -12, and -35 cannot conclusively be attributed to A. garhi.[1]

The remains are dated to about 2.5 million years ago (mya) based on

synapomorphies (traits unique to only them).[4][5] In 2015, Homo was recorded from 2.8 mya, much earlier than A. garhi.[6]

Palaeoecology

The large teeth of Australopithecus species have historically been interpreted as having been adaptations for a diet of hard foods, but the durable teeth may instead have only served an important function during leaner times for harder fallback foods. That is, dental anatomy may not accurately portray normal Australopithecus diet, rather abnormal diet during times of famine.[7]

Though it was not found with any tools, mammalian bones associated with the A. garhi remains exhibit cut and percussion marks made from stone tools: the left

bovid with three successive, unambiguous cut marks presumably made while removing the tongue; a bovid tibia with cut marks, chop marks, and impact scars from a hammerstone, possibly inflicted to harvest the bone marrow; and a Hipparion (a horse) femur with cut marks consistent with dismemberment and filleting. As to why stone tools were not present, because the Hatayae locality was likely a featureless, grassy lake margin with so few raw materials for making stone tools, it is possible these hominins were creating and carrying tools some ways with them to butchering sites, intending to use them many times before discarding. It was previously believed that only Homo could manufacture tools;[8][3] but it is also possible that the butcherers were not manufacturing tools and simply used naturally sharp rocks.[9]

At the nearby Gona site, where there is an abundance of raw materials, several Oldowan tools (an industry previously believed to have been invented by H. habilis) were recovered from 1992 to 1994. The tools date to around 2.6–2.5 mya, the oldest evidence of manufacturing at the time, and since A. garhi was the only species identified in the vicinity at the time, this species was the best candidate for authorship.[10][11] However, in 2015, the earliest remains of Homo, LD 350-1, were discovered in Ledi-Geraru, also in the Afar Region, dating to 2.8–2.75 mya.[6] More stone tools were found in 2019 dating to about 2.6 mya in Ledi-Geraru, predating the Gona artifacts, and these may be attributed to Homo; the invention of sharp-edged Oldowan tools could actually be due to specific adaptations characteristic of Homo.[12] Nonetheless, other australopithecines have been associated with stone tool manufacturing, such as the 2010 discovery of cut marks dating to 3.4 mya attributed to A. afarensis,[9] and the 2015 discovery of the Lomekwi culture from Lake Turkana dating to 3.3 mya possibly attributed to Kenyanthropus.[13]

African
hominin timeline (in mya)
View references
H. sapiensH. nalediH. rhodesiensisH. ergasterAu. sedibaP. robustusP. boiseiH. rudolfensisH. habilisAu. garhiP. aethiopicusLD 350-1K. platyopsAu. bahrelghazaliAu. deyiremedaAu. africanusAu. afarensisAu. anamensisAr. ramidusAr. kadabba


See also

References

  1. ^
    PMID 10213683
    .
  2. ^ .
  3. ^ .
  4. .
  5. ^ Strait, D. S. (2001). "The systematics of Australopithecus garhi". Ludus Vitalis. 9 (15): 109–135.
  6. ^
    PMID 25739410
    .
  7. .
  8. .
  9. ^ .
  10. .
  11. .
  12. .
  13. .

External links