Pentacentron

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Pentacentron
Temporal range: Ypresian
Holotype infructescence, SR 93-08-02
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Order: Trochodendrales
Family: Trochodendraceae
Genus: Pentacentron
Species:
P. sternhartae
Binomial name
Pentacentron sternhartae
Manchester et al, 2018

Pentacentron is an

Washington state, United States. P. sternhartae are possibly the fruits belonging to the extinct trochodendraceous leaves Tetracentron hopkinsii
.

Distribution and paleoenvironment

Pentacentron sternhartae is known from specimens which are recovered from outcrops of the early Eocene,

golden larch, but also having notable traces of fir, spruce, cypress, and palm.[1]

Taxonomy

The species was described from a

matronym honoring Janet L. Hartford of Republic, Washington. Sternberg and Hartford have helped to facilitate the collecting of fossils at Republic, and promoted the research and education of fossils done through the Stonerose Interpretive center.[2]

Pe. sternhartae is one of between three and four trochodendraceae species that have been described from the Klondike Mountain Formation. Broadly circumscribed, three other species have been identified at Republic, Paraconcavistylon wehrii, Tetracentron hopkinsii, and Trochodendron nastae. Additionally the species Trochodendron drachukii is known from related Kamloops group shales at the McAbee Fossil Beds near Cache Creek, British Columbia. Manchester et al 2018 noted that Tr. drachukii is likely the fruits of Tr. nastae, while Pe. sternhartae are likely the fruits of Te. hopkinsii.[2] If fossils of the fruits and foliage in attachment are found, that would bring the species count down to three whole plant taxa.[2]

Description

Pentacentron sternhartae fruiting spikes range between 5.8–9.2 cm (2.3–3.6 in) in length with the fruits arranged long the axis in a helical pattern. Each capsule is sessile on the thin raceme. The capsular heads consist of five fruiting chambers, arranged pentagonally around the midline of the 1.3–1.5 mm (0.051–0.059 in) wide head. Growing from the middle area of each chamber is an apically and inwardly curving persistent style, each with an elliptical nectary bulge at its base. Rimming the base of each fruit are Perianth scars creating a raised flange. The fruits opened apically, through separation lines running from just above the styles up to the fruit tip where they met to form a pentagonal star like opening.[2]

References

  1. ^ .
  2. ^ .
  3. ^ Wolfe, J.A.; Tanai, T. (1987). "Systematics, Phylogeny, and Distribution of Acer (maples) in the Cenozoic of Western North America". Journal of the Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy. 22 (1): 1–246.
  4. ^ Wolfe, J.A.; Wehr, W.C. (1987). "Middle Eocene dicotyledonous plants from Republic, northeastern Washington". United States Geological Survey Bulletin. 1597: 1–25.