Bayan Mandahu Formation

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Bayan Mandahu Formation
Ma
TypeGeological formation
Unit ofGobi Desert
Lithology
PrimarySandstone
Location
Coordinates41.75N 106.75E
RegionInner Mongolia
Country China
Type section
Named forBayan Mandahu, Urad Rear Banner, Inner Mongolia

The Bayan Mandahu Formation (also known as Wulansuhai Formation or Wuliangsuhai Formation) is a

Period. Laid down in the Campanian, it is dated somewhat uncertainly to between 75 and 71 mya (million years ago).[1]

Description

The paleoenvironment it preserves was semi-

arid and characterized by alluvial (stream-deposited) and eolian (wind-deposited) sediments. The formation is known for its vertebrate fossils, most of which are preserved in unstructured sandstone, indicating burial by wind-blown sandstorms.[2]

Paleofauna of the Bayan Mandahu Formation

The

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon
Ootaxon
Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Crocodylomorphs

Crocodylomorphs
of the Bayan Mandahu Formation
Genus Species Location Material Notes Images
Shamosuchus[2] A mesoeucrocodylian.

Lizards

An

iguanian species is known from the formation.[2]

Lizards of the Bayan Mandahu Formation
Genus Species Location Material Notes Images
Adamisaurus[2] A. magnidentatus[2] An
agamid
.
Bainguis[2] A diploglossan.
Telmasaurus[2] A monitor lizard.

Turtles

Turtles of the Bayan Mandahu Formation
Genus Species Location Material Notes Images
Basilemys[2] A softshell turtle.
"Zangerlia" "Z." neimongolensis Bayan Mandahu "Partial to complete skeletons of multiple individuals."[3] A
nanhsiungchelyid
.

Mammals

A taeniolabidoidea multituberculate is known from the formation.[2]

Mammals of the Bayan Mandahu Formation
Genus Species Location Material Notes Images
Guibaatar[4] G. castellanus A djadochtatheriid multituberculate.
Kennalestes[2] K. gobiensis[2] A placental. Also present in the Djadochta Formation.
Kryptobaatar[1] K. mandahuensis A
multituberculate
.

Dinosaurs

Alvarezsaurs

Genus Species Location Material Notes Images
Alvarezsauridae indet.[5] Indeterminate[5] Bayan Mandahu[5] An articulated series of partial anterior cervical vertebrae, an anterior cervical vertebra, isolated fragmentary anterior caudal vertebrae, a partial scapula, and pedal phalanges.[5] An unnamed parvicursorine
alvarezsaurid.[5]
Linhenykus L. monodactylus[6] Wuliangsuhai [6] Cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, a scapulocoracoid, a sternum, much of the forelimbs, a partial pelvis, nearly complete hindlimbs, and some unidentified fragments.[6] A monodactyl parvicursorine alvarezsaurid.[6]

Ankylosaurs

Genus Species Location Material Notes Images
Pinacosaurus P. grangeri[7] Bayan Mandahu[7] [Four] juvenile specimens consisting of skulls, manidibles, cervical half-rings and almost complete skeletons, and an incomplete subadult skull.[7] A basal ankylosaurine ankylosaurid also known from the Djadochta Formation.
P. mephistocephalus[8] Bayan Mandahu[8] A nearly complete articulated skeleton, with in situ cervical dermal armour and tail-club, that lies in a natural with limb bones doubled under the body.[8] A basal ankylosaurine ankylosaurid.

Birds

Genus Species Location Material Notes Images
cf. Gobipteryx Indeterminate "Eggs similar to those attributed to Gobipteryx minuta from the Djadochta Formation."[2]

Ceratopsians

Genus Species Location Material Notes Images
Genus Species Location Material Notes Images
Bagaceratops B. rozhdestvenskyi Bayan Mandahu A partial skull.[9] A
protoceratopsid also known from the Barun Goyot Formation and possibly the Djadochta Formation
.
Magnirostris[1]
M. dodsoni[1] Bayan Mandahu[10] A partial skull. A
protoceratopsid now considered synonym of Bagaceratops.[9]
Protoceratops P. hellenikorhinus[11] Bayan Mandahu[11] Skulls and sparse body remains from multiple individuals.[12][11] A
protoceratopsid
.
Udanoceratops? Indeterminate Bayan Mandahu "Nearly 1 meter long skull."[2] A giant
leptoceratopsid. Reported remains probably represent Protoceratops hellenikorhinus.[13]

Dromaeosaurs

Genus Species Location Material Notes Images
Linheraptor[14] L. exquisitus[14] Wuliangsuhai[14] A skull, cervical vertebrae, caudal vertebrae, a scapula, a humerus, a radius, metacarpals, manual phalanxes, sternal plate, a pubis, a femur, a tibiotarsus, metatarsals, and pedal phalanxes.[14] A possible velociraptorine dromaeosaurid.[14]
Velociraptor V. osmolskae[15] Bayan Mandahu[1] A paired maxillae and left lacrimal.[1] A velociraptorine dromaeosaurid.[1]

Hadrosaurs

Genus Species Location Material Notes Images
Undescribed Hadrosauridae[10] Indeterminate[10] Bayan Mandahu[10] A hadrosaurid.

Oviraptorosaurs

Genus Species Location Material Notes Images
Machairasaurus M. leptonychus[10] Bayan Mandahu[10] A nearly complete articulated right forearm and manus, pedal phalanges, manual phalanges, caudal vertebrae, dorsal ribs, chevrons, and fragments of metatarsals III and IV.[10] A heyuanniine
oviraptorid.[10]
Oviraptoridae indet. Indeterminate Bayan Mandahu "Partial individual sitting atop an egg clutch."[16] An
oviraptorid that represent the second nesting oviraptorid. Sometimes referred to Machairasaurus.[10][17]
Wulatelong W. gobiensis[18] Wuliangsuhai[18] A partial skull, dorsal vertebrae, caudal vertebrae, partial scapulocoracoids, a partial sternum, a partial humerus, an incomplete manus, a nearly complete pelvic girdle, and the right hindlimb.[18] A basal
oviraptorosaur.[18]

Sauropods

Genus Species Location Material Notes Images
Undescribed Sauropoda[10] Indeterminate[10] Bayan Mandahu[10] A
sauropod
.

Troodontids

Genus Species Location Material Notes Images
Linhevenator[19] L. tani[19] Wuliangsuhai[19] A skull and mandible, anterior dorsal vertebrae, middle dorsal vertebrae, a scapula, a humerus, incomplete ischia, a femur, nearly complete pes, and other fragmentary bones.[19] A
troodontid with short forearms.[19]
Papiliovenator P. neimengguensis[20] Bayan Mandahu[20] An almost complete skull, cervical vertebrae, anterior dorsal vertebrae, partial scapulae, partial coracoid, fragmentary pelvic girdle, partial humeri, ulna, radius, partial manus, femur, tibia, fibula, pes, and a pedal phalanx II-3.[20] A troodontid that is part of a clad that includes all other Late Cretaceous troodontids except Almas.[20]
Philovenator P. curriei[21] Wuliangsuhai[21] A nearly complete femur, a partial tibia, a partial fibula, astragalus-calcaneum complex, distal tarsals, metatarsals, and pedal phalanges.[21] A
troodontid.[21]

Tyrannosaurs

Genus Species Location Material Notes Images
Undescribed Tyrannosauridae[10] Indeterminate[10] Bayan Mandahu[10] A
tyrannosaurid
.

See also

References

  1. ^ .
  2. ^ .
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  4. .
  5. ^ .
  6. ^ .
  7. ^ .
  8. ^ a b c Godefroit, P.; Pereda-Suberbiola, X.; Li, H.; Dong, Z. M. (1999). "A new species of the ankylosaurid dinosaur Pinacosaurus from the Late Cretaceous of Inner Mongolia (P.R. China)" (PDF). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 69 (supp. B): 17–36.
  9. ^
    S2CID 132780322. Archived from the original
    (PDF) on 2021-07-08. Retrieved 2021-03-03.
  10. ^ .
  11. ^ a b c Lambert, O.; Godefroit, P.; Li, H.; Shang, C.-Y.; Dong, Z. (2001). "A new Species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China)" (PDF). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 71: 5−28.
  12. .
  13. .
  14. ^ .
  15. .
  16. .
  17. .
  18. ^ a b c d Xu Xing; Tan Qing-Wei; Wang Shuo; Corwin Sullivan; David W. E. Hone; Han Feng-Lu; Ma Qing-Yu; Tan Lin & Xiao Dong (2013). "A new oviraptorid from the Upper Cretaceous of Nei Mongol, China, and its stratigraphic implications" (PDF). Vertebrata PalAsiatica. 51 (2): 85–101.
  19. ^
    PMID 21915256
    .
  20. ^ .
  21. ^ a b c d Xu Xing; Zhao Ji; Corwin Sullivan; Tan Qing-Wei; Martin Sander; Ma Qing-Yu (2012). "The taxonomy of the troodontid IVPP V 10597 reconsidered" (PDF). Vertebrata PalAsiatica. 50 (2): 140–150.

Bibliography