Wheeler Shale
Wheeler Shale | |
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Ma | |
Elrathia kingii, famed trilobite of the Wheeler Shale. | |
Type | Geological formation |
Thickness | 100–200 m (330–660 ft) |
Lithology | |
Primary | Calcareous shale |
Other | Mudstone, shaley limestone and limestone |
Location | |
Coordinates | 39°15′N 113°20′W / 39.25°N 113.33°W |
Region | House Range and Drum Mountains, Millard Co., west Utah |
Country | United States |
Type section | |
Named for | House Amphitheater (Geographic feature and type locality) |
Named by | Charles Doolittle Walcott |
Part of a series on |
The Cambrian explosion |
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The Wheeler Shale (named by
Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms 490 to 610 m (1,610 to 2,000 ft) of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America.[5]
At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thin, flaggy limestone.[6] The Wheeler Formation (although the Marjum & Weeks Formations are missing) extends into the Drum Mountains, northwest of the House Range where similar fossils and preservation are found.[6]
Taphonomy and sedimentology
Detailed work recognises a number of ~10 m thick lagerstätten sequences in the formation, each of which formed at a sea-level high stand[7] in deep water.[8] The lagerstätte were deposited by turbidities and mudslides onto an oxygenated sea floor.[7] The productive layers comprise mud and clay particles, with a tiny fraction of wind-blown quartz.[9]
Stratigraphy
The Wheeler Shale spans the Ptychagnostus atavus[10] and uppermost-Middle Cambrian Bolaspidella trilobite zones (See House Range for full stratigraphy).
Fauna
Incomplete list of the fauna of the Wheeler Shale:[6][11][12][13][14][15][16] (Note: the preservation of hard bodied trilobite remains and soft bodied animals seems to be mutually exclusive within particular horizons.)[4][17]
Protista
- Marpolia spissa - cyanobacteria or green algae
- Morania fragmenta - cyanobacteria
Arthropoda
- Branchiocaris pretiosa - hymenocarine
- Branchiocaris sp.
- myriapod
- Canadaspis perfecta - hymenocarine
- Dicerocaris opisthoeces
- vicissicaudatan
- Isoxys
- Perspicaris dilatus hymenocarine
- Pseudoarctolepis sharpi - possible hymenocarine
- Tuzoia? peterseni hymenocarine
- Waptia fieldensis hymenocarine
- Alalcomenaeus cambricus - megacheiran; or alalcomenaeid
- Dicranocaris guntherorum - megacheiran; or alalcomenaeid
- unnamed 'Molli Sonia symmetrica'
- Leanchoilia superlata - megacheiran
- Sidneyia inexpectans - vicissicaudatan
Dinocaridida
- Amplectobelua cf. A. stephenensis – radiodont[18]
- Anomalocarididae gen. et sp. nov. - radiodont[18]
- Buccaspinea cooperi? - radiodont[18]
- Caryosyntrips durus - radiodont[18]
- Caryosyntrips serratus - radiodont[18]
- Peytoia nathorsti - radiodont[18]
- Pahvantia hastata - radiodont[18]
- Stanleycaris sp. - radiodont[18]
- Utahnax vannieri - kerygmachelid lobopodian, also possible that is from Marjum Formation[19]
- Utaurora comosa - opabiniid[20]
Trilobita
- nectaspid
- agnostid
- Peronopsis amplaxis - peronopsid agnostid
- Peronopsis bidens
- Peronopsis fallax
- Peronopsis gaspensis
- Peronopsis intermedius
- Peronopsis interstrictus
- Peronopsis montis
- Peronopsis segmentis
- Ptychagnostus atavus (= Acidusus atavus) - ptychagnostid agnostid
- Ptychagnostus germanus
- Ptychagnostus gibbus
- Ptychagnostus intermedius
- Ptychagnostus michaeli
- Ptychagnostus occultatus
- Ptychagnostus seminula
- Glyphaspis concavus - asaphid
- Bathyuriscus fimbriatus - dolichometopid corynexochid
- Bathyuriscus sp.
- Kootenia sp. - dorypygid corynexochid, perhaps a synonym of Olenoides
- Olenoides expansus - dorypygid corynexochid
- Olenoides nevadensis
- Olenoides serratus
- Tonkinella breviceps
- Zacanthoides divergens - zacanthoidid corynexochid
- Zacanthoides sp.
- Altiocculus harrisi - ptychopariid (specific name may be confused with Alokistocare)
- Alokistocare harrisi - alokistocarid ptychopariid
- Asaphiscus wheeleri - ptychopariid; second-most common species in the formation
- Bathyocos housensis - ptychopariid
- Bolaspidella drumensis
- Bolaspidella housensis
- Bolaspidella sp.
- Bolaspidella wellsvillensis
- Brachyaspidion microps
- Brachyaspidion sulcatum
- Cedaria minor - known from the Warrior Formation
- Elrathia kingii - alokistocarid ptychopariid
- Elrathia sp.
- Elrathinawheeleri = Ptychoparella wheeleri? - ptychopariid
- Jenkinsonia varga
- Modocia brevispina
- Modocia laevinucha
- Modocia typicalis
- Ptychoparella sp. - ptychopariid
- Ptychoparella wheeleri
- Spencella sp. - ptychopariid
Brachiopoda
- Acrothele subsidua
Chordata
Cnidaria
- Cambromedusa sp. - jellyfish
Mollusca
- Pelagiella sp. - pelagiellid helcionelloid
Echinodermata
- carpoid
- Castericystis sp.
- Cothurnocystis sp. - stylophoran
- Ctenocystis sp. - ctenocystoid
- eocrinoid
- Eocrinoid holdfasts believed to belong to Gogia spiralis; may belong to other species
Porifera
- Choia carteri - choiid monaxonid demosponge
- Choia utahensis
- Crumillospongia sp. - hazeliid monaxonid demosponge
- Diagonellasp.
Priapulida
- Ottoia prolifica - a stem group and it was an archaeopriapulid
- Selkirkia sp. - archaeopriapulid
- "Selkirkia willoughbyi" (Note: S. columbia is the only recognized species)
Unclassified
- Hallucigenia sparsa - ?xenusiid lobopod
- Allonnia cf. tintinopsis - a chancelloriid[21]
- Chancelloria pentacta - chancelloriid coeloscleritophoran, perhaps a sponge?
- paropsonemid cambroernid
- Skeemella clavula - Possible vetulicolian
- Hylolithellus sp. - annelid?
- Wiwaxia corrugata - halwaxiid? lophotrochozoan
- Yuknessia simplex - pterobranch
- Margaretia dorus - possibly organic tube associated with hemichordate[22]
References
- ISBN 978-1-55591-451-6
- doi:10.1130/G19926.1
- ^ Robert R. Gaines; Derek E.G. Briggs; Zhao Yuanlong (2008), "Cambrian Burgess Shale–type deposits share a common mode of fossilization", Geology, 36 (10): 755–758,
- ^
- ^ Robison, R.A. (1964), "Late Middle Cambrian faunas from western Utah", Journal of Paleontology, 38 (3): 510–566
- ^ a b c Gunther, L.F.; Gunther, V.G. (1981), "Some Middle Cambrian Fossils of Utah", Brigham Young University Geology Studies, 28: 1–81
- ^ .
- .
- .
- S2CID 130145927.
- ^ Briggs, D.E.G.; Robison, R.A. (1984), Exceptionally preserved nontrilobite arthropods and Anomalocaris from the Middle Cambrian of Utah, The Paleontological Institute, The University of Kansas
- ^ Photos of Wheeler Shale fossils from UC Berkeley
- JSTOR 1303938
- ^ Utah's Cambrian Life from University of Kansas Natural History Museum
- ^ Cambrian fossils from Utah by the University of Utah
- ^ Comprehensive treatment from The Virtual Fossil Museum
- ^ Gaines, Robert R.; Droser, Mary L.; Kennedy, Martin J. (2001), "Taphonomy of soft-bodied preservation and ptychopariid Lagerstätte in the Wheeler Shale (Middle Cambrian), House Range, USA; controls and implications", PaleoBios, 21 (Suppl.2): 1–55
- ^ PMID 33552709.
- S2CID 250076505.
- PMID 35135344.
- S2CID 251544727.
- PMID 27383414.