Dihoplus
| Dihoplus Temporal range:
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|---|---|
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| Dihoplus schleiermacheri holotype skull | |
Scientific classification 
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| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Perissodactyla |
| Family: | Rhinocerotidae |
| Tribe: | Dicerorhinini
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| Genus: | †Dihoplus Brandt, 1878 |
| Type species | |
| †Rhinoceros schleiermacheri Kaup, 1832
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| Species | |
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Dihoplus is an extinct genus of rhinoceros that lived in Eurasia from the Late Miocene to Pliocene.[1]
Description
Species of Dihoplus were large rhinoceroses with two horns. The nasal septum was not ossified, with a nasal notch above the frontmost premolars. The toothrow is placed posteriorly within the skull. The first upper premolar is absent, though the lower second incisor is present.[1]
Taxonomy
Members of Dihoplus were long placed in Dicerorhinus (which contains the living Sumatran rhinoceros). Sometimes these species are placed in the related Stephanorhinus. The genus is now generally considered distinct,[2] though there is still debate as to which species should be included; for example, Deng (2011) listed Merck's rhinoceros (Stephanorhinus kirchbergensis) under Dihoplus.[3] Species recently placed in the genus include:[4]
- D. schleiermacheri (Kaup, 1832) the type species of the genus, known from Late Miocene of Europe, with the type specimen being known from Eppelsheim, Germany.[4]
- D. pikermiensis (Toula, 1906), known from the Late Miocene of Europe, originally placed in the genus Stephanorhinus.[4]
- ‘D.’ bethlehemsis Pandolfi, Rivals and Rabinovich, 2023, known from Pliocene aged deposits from Bethlehem in the Levant.[4]
"D." megarhinus (de Christol, 1834) from the Late Miocene-Pliocene of Europe, Anatolia, China and Transbaikalia,[1][4] since 2021 has been placed instead in the genus Pliorhinus along with "Dicerorhinus" miguelcrusafonti from the Pliocene of Europe.[5][6] "D." ringstoemi is either considered a synonym of "D." megarhinus[1] or placed as a separate species within Pliorhinus.[7]
The monophyly of the genus has been questioned, with some studies suggesting that D. pikermiensis is more closely related to Stephanorhinus and Coelodonta (which contains the woolly rhinoceros) than to the type species D. schleiermacheri.[6]
Morphological phylogeny after Pandolfi (2023), excluding living African rhinoceros species.[6]
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Ecology
D. pikermiensis and D. bethlehemsis are suggested to have been browsers.[8][4]
References