Mekosuchinae

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Mekosuchinae
Temporal range:
Ma
Restoration of the skull of Quinkana timara at the Central Australian Museum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauromorpha
Clade: Archosauriformes
Order: Crocodilia
Clade: Mekosuchinae
Willis, Molnar & Scanlon, 1993
Type species
Balouet
& Buffetaut, 1987
Genera

Mekosuchinae is an extinct

fossil record in the Eocene in Australia, and survived until the arrival of humans: in the Late Pleistocene in Australia and within the Holocene in the Pacific islands of Fiji, New Caledonia and Vanuatu
.

Mekosuchine crocodiles are a diverse group. One of the last species, Mekosuchus inexpectatus from Holocene New Caledonia, may have been arboreal.[2] The early Miocene species Harpacochampsa camfieldensis may have resembled a false gharial. Another mekosuchine fossil, currently undescribed, has been found in Miocene deposits from New Zealand. One genus, Mekosuchus, managed to spread to the islands of the Pacific; it is believed to have island-hopped across the Coral Sea, moving first to a now submerged island known as Greater Chesterfield Island, then New Caledonia and onwards. In the Pleistocene, Quinkana was one of the top terrestrial predators of the Australian continent.

Mekosuchines underwent a drastic decline in post-Miocene Australia, with all genera, except for Quinkana and

Quaternary extinction event) becoming extinct in Australia by the end of the Pliocene
. After the demise of Quinkana and Pallimnarchus, the group survived on Vanuatu and New Caledonia until the arrival of humans, who are presumed to have driven them to extinction.

While historically considered to be true

Crocodylidae), modern research places them as an independent group within or closely related to Longirostres, which contains both crocodiles and gavialids.[1]

Phylogeny

Mekosuchus inexpectatus

Mekosuchinae is

Mekosuchus inexpectatus, and all of its descendants.[3]

Mekosuchinae is traditionally thought to be included as a

stratigraphic (fossil age) data established the inter-relationships within Crocodilia,[5] which was expanded upon in 2021 by Hekkala et al. using paleogenomics by extracting DNA from the extinct Voay.[6]

The below cladogram shows the results of the Lee and Yates study (2018), which placed Mekosuchinae outside of Crocodyloidea, as more basal than Longirostres (the combined group of crocodiles and gavialids).[5]

Crocodylia

Alligatoroidea

Prodiplocynodon

Asiatosuchus

"Crocodylus" affinis

"Crocodylus" depressifrons

"Crocodylus" acer

Brachyuranochampsa

Mekosuchinae

Australosuchus

Kambara taraina

Kambara implexidens

Kambara murgonensis

Kalthifrons

Pallimnarchus

Baru wickeni

Baru darrowi

Baru Alcoota

Bullock Creek taxon

Baru huberi

Longirostres

Other studies have recovered Mekosuchinae within Longirostres, belonging to Crocodyloidea, as shown in the alternative cladogram below.[1]

Anteophthalmosuchus epikrator

Theriosuchus pusillus

Susisuchidae

Bernissartia fagesii

Eusuchia

Hylaeochampsidae

Allodaposuchidae

Portugalosuchus azenhae

Crocodylia

Planocraniidae

Prodiplocynodon langi

'Crocodylus' affinis

'Crocodylus' depressifrons

Asiatosuchus germanicus

'Crocodylus' acer

Brachyuranochampsa eversolei

Longirostres
Gavialoidea

Kentisuchus spenceri

Megadontosuchus arduini

Gunggamarandu maunala

Dollosuchoides densmorei

Maomingosuchus petrolica

Harpacochampsa camfieldensis

Thecachampsa antiquus

'Tomistoma' cairense

Gavialidae

Crocodyloidea

'Crocodylus' megarhinus

Mekosuchinae

Kambara implexidens

Kambara taraina

Kambara murgonensis

Australosuchus clarkae

Kalthifrons aurivellensis

Mekosuchini

Trilophosuchus rackhami

Mekosuchus whitehunterensis

Mekosuchus sanderi

Baru wickeni

Baru darrowi

Paludirex gracilis

Paludirex vincenti

Quinkana meboldi

Quinkana fortirostrum

Quinkana timara

Crocodylidae

References