Tianzhenosaurus

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Tianzhenosaurus
Temporal range:
Ma
Reconstructed skeleton
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Thyreophora
Clade: Ankylosauria
Family: Ankylosauridae
Subfamily: Ankylosaurinae
Tribe:
Ankylosaurini
Genus: Tianzhenosaurus
Pang & Cheng, 1998
Species:
T. youngi
Binomial name
Tianzhenosaurus youngi
Pang & Cheng, 1998
Synonyms

Tianzhenosaurus (meaning “

junior synonym of Saichania, an ankylosaurine known from the Barun Goyot and Nemegt Formation.[2]

Discovery and naming

Holotype skull of Tianzhenosaurus

In 1983, Pang Qiqing and Cheng Zhengwu discovered articulated cervical vertebrae of an ankylosaurid from the

Hebei GEO University, Shijiazhuang.[1]

The generic name, Tianzhenosaurus, is derived from the

In 1999, Sullivan considered Tianzhenosaurus as a

Currie (2015) also reaffirmed Tianzhenosaurus as a junior synonym of Saichania based on similar reasons provided by Sullivan (1999), but noted that if the humerus of Tianzhenosaurus is later shown to differ from that of Saichania, then could be considered as a distinct taxon.[2]

Description

caudal vertebrae that ended with a tail club; rectangular, plate-like scapula; proximal and distal ends of humerus are moderately expanded and not twisted; thick femur lacking the fourth trochanter; tarsometatarsal and digits that are typical for ankylosaurs.[1]

Reconstructed skull

Pang & Cheng (1998) noted that the overall skull morphology of Tianzhenosaurus was similar to that of Saichania as both taxa have an isosceles triangle shaped skull, a similar location of the orbit at the mid-posterior part of the skull,

horizontal extension of the opisthotic and having no ventral curvature.[1]

Classification

Pang & Cheng (1998) originally placed Tianzhenosaurus into Ankylosauridae, but did not specify it's relationship with other ankylosaurids.[1] Both Sullivan (1999) and Arbour & Currie (2015) considered Tianzhenosaurus as a junior synonym of Saichania based on the overall similar morphology of the skulls.[3][2] Vickaryous et al. (2004) classified Tianzhenosaurus as an ankylosaurine, sister taxon to Pinacosaurus mephistocephalus, while Thompson et al. (2012) recovered Tianzhenosaurus as sister taxon to Talarurus.[4][5] However, Wiersma and Irmis (2018) recovered Tianzhenosaurus as a valid taxon and interpreted it as sister taxon to Pinacosaurus grangeri.[6]

A phylogenetic analysis conducted by Thompson et al., 2012 is reproduced below.[5]

Huayangosaurus taibaii

Stegosaurus armatus

Nodosauridae

Ankylosauridae

Minmi paravertebra

Liaoningosaurus paradoxus

Cedarpelta bilbeyhallorum

Gobisaurus domoculus

Shamosaurus scutatus

Zhongyuansaurus luoyangensis

Tsagantegia longicranialis

Shanxia tianzhensis

"Crichtonsaurus" benxiensis

Dyoplosaurus acutosquameus

Pinacosaurus mephistocephalus

Ankylosaurus magniventris

Euoplocephalus tutus

Minotaurasaurus ramachandrani

Pinacosaurus grangeri

Nodocephalosaurus kirtlandensis

Talarurus plicatospineus

Tianzhenosaurus youngi

Saichania chulsanensis

Tarchia gigantea

The results of an earlier analysis by Vickaryous et al., 2004 is reproduced below.[4]

Paleoenvironment

Stratigraphic map of the Huiquanpu Formation

The holotype specimen of Tianzhenosaurus was recovered from the Huiquanpu Formation, which possibly dates to the Cenomanian or Campanian stage of the Late Cretaceous. However, the precise dating of the formation is problematic due to the absence of index fossils and the Cretaceous age is partially based on the discovery of indeterminate pliosaur material.[7] The Huiquanpu Formation consists of grey siltstones interbedded with red medium to coarse grained sandstones that has extensive cross-bedding.[7] Based on the sedimentation, the formation represents a fluvial environment, with the upper part of the formation being extensively reworked by a number of early Tertiary volcanic intrusions.[7]

Tianzhenosaurus would have coexisted with the macronarian sauropod Huabeisaurus,[8] the hadrosauroid ornithopod Datonglong,[9] the pantyrannosaurian tyrannosauroid Jinbeisaurus[10] and the ankylosaurine ankylosaurid Shanxia, which may represent a junior synonym of Tianzhenosaurus.[7][3]

See also

References

  1. ^ a b c d e f g h i j k l Pang, Q.; Chang, Z. (1998). "A new ankylosaur of the late Cretaceous from Tianzhen, Shanxi". Progress in Natural Science.
  2. ^
    S2CID 214625754
    .
  3. ^ .
  4. ^ a b Vickaryous, M. K., Maryanska, T., and Weishampel, D. B. (2004). Chapter Seventeen: Ankylosauria. in The Dinosauria (2nd edition), Weishampel, D. B., Dodson, P., and Osmólska, H., editors. University of California Press.
  5. ^
    S2CID 86002282
    .
  6. .
  7. ^ .
  8. ^ Pang, Q.; Cheng, Z.; Yang, J.; Xie, M.; Zhu, C.; Luo, J. (1996). "The preliminary report on Late Cretaceous dinosaur fauna expeditions in Tianzhen, Shanxi". Journal of Hebei College of Geology. 19 (3–4): 227–235.
  9. .
  10. .