Apocrita
Apocrita Temporal range:
| |
---|---|
Seleucus cuneiformis (Ichneumonidae) | |
Vespula germanica (Vespidae) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
(unranked): | Unicalcarida
|
Suborder: | Apocrita Gerstaecker, 1867[1] |
Subgroups | |
|
Apocrita is a
petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma (or gaster) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor.[2] The ovipositor
either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell provisioned by their mothers.
Apocrita has historically been split into two groups,
paraphyletic
: the clade would contain the Aculeata. "Parasitica" is therefore a rankless grouping in many present classifications, if it appears at all.
Parasitica comprises the majority of hymenopteran insects, its members living as
biological control agents to control pests, such as caterpillars, true bugs and hoppers, flies, and weevils.[6]
Aculeata is a
social hymenopterans.[7]
Among the nonparasitic and nonsocial Aculeata, larvae are fed with captured prey (typically alive and paralyzed) or may be fed pollen and nectar. The fungi
, or nonviable eggs (ants).
Extant families and superfamilies
The Apocrita contains a large number of families. Some traditional taxa such as the
paraphyletic. Parasitoidism evolved once, and it is found today across most Apocritan families, though it has been secondarily lost several times. The phylogenetic tree gives a condensed overview of the phylogeny, illustrated with major groups. The sawflies are paraphyletic as the Apocrita evolved inside that group. The tree is not fully resolved.[7][8][9][3]
Cladogram of Apocrita after Peters et al.(2017)[3]
Apocrita |
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- Suborder Apocrita
- Infraorder Aculeata
- Superfamily Apoidea (bees and sphecoid wasps)
- Family cockroach wasps)
- Family Andrenidae (mining bees)
- Family honeybees)
- Family plasterer bees)
- Family Crabronidae (sand wasps, bee wolves, etc.)
- Family Halictidae ("sweat bees")
- Family Heterogynaidae
- Family leaf-cutting bees)
- Family Melittidae
- Family Stenotritidae
- Family Sphecidae (digger wasps)
- Family
- Superfamily Chrysidoidea
- Family Bethylidae
- Family Chrysididae (cuckoo wasps)
- Family Dryinidae
- Family Embolemidae
- Family Plumariidae
- Family Sclerogibbidae
- Family Scolebythidae
- Superfamily Formicoidea
- Family Formicidae (ants)
- Superfamily Pompiloidea
- Family velvet ants)
- Family Myrmosidae
- Family Pompilidae (spider wasps)
- Family Sapygidae
- Family
- Superfamily Scolioidea
- Family Scoliidae
- Superfamily Tiphioidea
- Family Bradynobaenidae
- Family Sierolomorphidae
- Family Tiphiidae
- Superfamily Thynoidea
- Family Chyphotidae
- Family Thynnidae
- Superfamily Vespoidea
- Family Rhopalosomatidae
- Family Vespidae (paper wasps, potter wasps, hornets, pollen wasps, yellowjackets)
- Superfamily Apoidea (bees and sphecoid wasps)
- Proctotrupomorpha
- Superfamily Chalcidoidea
- Family Agaonidae (fig wasps)
- Family Aphelinidae
- Family Chalcididae (chalcid wasps)
- Family Encyrtidae
- Family Eucharitidae
- Family Eulophidae
- Family Eupelmidae
- Family seed chalcids)
- Family Leucospidae
- Family Mymaridae (fairyflies) – the smallest of all insects
- Family Ormyridae
- Family Perilampidae
- Family Pteromalidae
- Family Rotoitidae
- Family Signiphoridae
- Family Tanaostigmatidae
- Family Tetracampidae
- Family Torymidae
- Family Trichogrammatidae
- Superfamily Cynipoidea
- Family Austrocynipidae
- Family Cynipidae (gall wasps)
- Family Figitidae
- Family Ibaliidae
- Family Liopteridae
- Superfamily Diaprioidea
- Family Austroniidae
- Family Diapriidae
- Family Maamingidae
- Family Monomachidae
- Family
- Superfamily Mymarommatoidea
- Family Mymarommatidae
- Superfamily Platygastroidea
- Family Geoscelionidae
- Family Janzenellidae
- Family Neuroscelionidae
- Family Nixoniidae
- Family Platygastridae
- Family †Proterosceliopsidae
- Family Scelionidae
- Family Sparasionidae
- Superfamily Proctotrupoidea
- Family Heloridae
- Family Pelecinidae
- Family Peradeniidae
- Family Proctorenyxidae
- Family Proctotrupidae
- Family Roproniidae
- Family Vanhorniidae
- Superfamily
- Superfamily Stephanoidea
- Family Stephanidae
- Superfamily Trigonaloidea
- Family †Maimetshidae
- Family Trigonalidae
- Superfamily Ceraphronoidea
- Family Ceraphronidae
- Family Megaspilidae
- Superfamily Evanioidea
- Family Aulacidae
- Family ensign wasps)
- Family Gasteruptiidae
- Superfamily Ichneumonoidea
- Family Braconidae
- Family ichneumon wasps)
- Superfamily Megalyroidea
- Family Megalyridae
- Infraorder Aculeata
References
- ^ Gerstaecker, C.E.A. (1867). "Ueber die Gattung Oxybelus Latr. und die bei Berlin vorkommenden Arten derselben". Zeitschrift für die Gesammten Naturwissenschaften (in German). 30 (7): 1–144.
- ISBN 978-0-521-82149-0.
- ^ PMID 28343967.
- PMID 21540117.
- ISBN 978-0-12-374144-8.
- ^ "Parasitoid Wasps (Hymenoptera)". University of Maryland. Archived from the original on 27 August 2016. Retrieved 6 June 2016.
- ^ PMID 28376325.
- .
- ^ Schulmeister, S. "Symphyta". Archived from the original on 21 June 2010. Retrieved 28 November 2016.
External links
Look up apocrita in Wiktionary, the free dictionary.
Wikimedia Commons has media related to Apocrita.
Wikispecies has information related to Apocrita.
- Suborder Apocrita – Ants, Bees and Wasps – BugGuide.Net — images and other information
- Tree of Life
- Balades Entomologiques — "entomological walks" with images (in French)