Borophagus
Appearance
Borophagus Ma
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Borophagus secundus skull | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Canidae |
Subfamily: | †Borophaginae |
Tribe: | †Borophagini |
Subtribe: | †Borophagina |
Genus: | †Borophagus Cope, 1892 |
Type species | |
†Borophagus diversidens | |
Other species | |
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Synonyms | |
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Borophagus ("gluttonous eater") is an extinct
Mya.[1]
Evolution
Borophagus, like other borophagines, are loosely known as "bone-crushing" or "
Aenocyon dirus. Early species of Borophagus were placed in the genus Osteoborus until recently, but the genera are now considered synonyms.[1]
Description


Typical features of this genus are a bulging forehead and powerful jaws; Borophagus has been considered to be probably a
hyena of the Old World. However, Borophagus fossils are so abundant and geographically widespread that some paleontologists now argue that Borophagus must have been both the dominant carnivore of its time, and thus an active predator because carrion feeding alone could not have sustained such a large population.[3] They note that not all carnivores with bone-cracking ability are scavengers, such as the modern spotted hyena; instead, they interpret the bone-cracking ability as an adaptation to social hunting where complete utilization of a carcass was favored.[3] Coprolites from Borophagus further vindicate its bone-crushing abilities, while simultaneously indicating it occupied a niche no longer seen in the present-day ecosystems of North America. The discovery of these coprolites also indicates that Borophagus may have been a social pack-hunter.[4]
The adult animal is estimated to have been about 80 cm in length, similar to a coyote, although it was much more powerfully built.[5]
Species

- Borophagus diversidens existed for 2.5 million years (synonymous with Felis hillianus, Hyaenognathus matthewi, Hyaenognathus pachyodon, Hyaenognathus solus, Porthocyon dubius)
- Borophagus dudleyi existed for 2 million years
- Borophagus hilli existed for 0.5 million years (synonymous with Osteoborus crassapineatus, Osteoborus progressus)
- Borophagus littoralis existed for 3 million years (syn. Osteoborus diabloensis)
- Borophagus orc existed for 2 million years
- Borophagus parvus existed for 2 million years
- Borophagus pugnator existed for 4 million years (synonymous with Osteoborus galushai)
- Borophagus secundus existed for 4 million years (synonymous with Hyaenognathus cyonoides, Hyaenognathus direptor, Osteoborus secundus)
Existence based on Figure 141 of Wang et al. (1999).[1]
Paleoecology
In North America, in places such as
machairodont cat Amphimachairodus coloradensis and fellow canid Epicyon. All of these animals were potential competitors that would have occasionally conflicted with Borophagus for food and territory, though it may also have readily scavenged their kills. Prey for Borophagus included herbivores like the camel Aepycamelus, the pronghorn antelope Cosoryx, horses like Neohipparion and Nannippus, the ancient peccary Prosthennops and even rhinoceroses like the hippo-like Teleoceras, all of which could provide a suitable meal through hunting or scavenging.[6][7]
References
- ^ a b c d Wang, Xiaoming; Richard Tedford; Beryl Taylor (1999-11-17). "Phylogenetic systematics of the Borophaginae" (PDF). Bulletin of the American Museum of Natural History. 243. Archived from the original (PDF) on September 30, 2007. Retrieved 2007-07-08.
- ISBN 0-8160-1125-7.
- ^ a b Wang, Xiaoming; and Tedford, Richard H. Dogs: Their Fossil Relatives and Evolutionary History. New York: Columbia University Press, 2008. pp112-3
- PMID 29785931.
- ISBN 1-84028-152-9.
- ISBN 9780253010421.
- ISBN 0-231-10228-3.
- Alan Turner, "National Geographic: Prehistoric Mammals" (Washington, D.C.: Firecrest Books Ltd., 2004), pp. 112–114. ISBN 0-7922-7134-3
- Xiaoming Wang, "The Origin and Evolution of the Dog Family" Accessed 1/30/06.
Further reading
- Picture of an Osteoborus skull in a museum, from "World of the Wolf." (Accessed 6/19/06)
- Russell Hunt, "Ecological Polarities Of the North American Family Canidae: A New Approach to Understanding Forty Million Years of Canid Evolution" (Accessed 1/30/06).
- Wang et al., "Phylogenetic Systematics of the Borophaginae (Carnivora:Canidae)." Bulletin of the American Museum of Natural History, No. 243, Nov. 17 1999. (PDF) (Accessed 4/11/06) Archived 2007-03-20 at the Wayback Machine
- Xiaoming Wang, Richard H. Tedford, Mauricio Antón, Dogs: Their Fossil Relatives and Evolutionary History, New York : Columbia University Press, 2008; ISBN 978-0-231-13528-3