Source: Wikipedia, the free encyclopedia.

Temporal range: Campanian - recent[1]
Least mallow, Malva parviflora
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Malvales
Family: Malvaceae

See List of Malvaceae genera

  • Bombacaceae Kunth
  • Brownlowiaceae Cheek
  • Byttneriaceae R.Br.
  • Dombeyaceae Kunth
  • Durionaceae Cheek
  • Helicteraceae J.Agardh
  • Hermanniaceae Marquis
  • Hibiscaceae J.Agardh
  • Lasiopetalaceae Rchb.
  • Melochiaceae J.Agardh
  • Pentapetaceae Bercht. & J.Presl
  • Philippodendraceae A.Juss.
  • Plagianthaceae J.Agardh
  • Sparmanniaceae J.Agardh
  • Sterculiaceae Vent.
  • Theobromataceae J.Agardh
  • Tiliaceae Juss.

Malvaceae (

roselle and durian. There are also some genera containing familiar ornamentals, such as Alcea (hollyhock), Malva (mallow), and Tilia (lime or linden tree). The genera with the largest numbers of species include Hibiscus (434 species),[6] Pavonia (291 species),[7] Sida (275 species),[8] Ayenia (216 species),[9] Dombeya (197 species),[10] and Sterculia (181 species).[11][12]

Taxonomy and nomenclature

Swamp rose mallow
Swamp rose mallow
Common mallow (Malva sylvestris)
Common mallow (Malva sylvestris)

The circumscription of the Malvaceae is controversial. The traditional Malvaceae

sensu lato, has been more recently defined on the basis that genetics studies have shown the commonly recognised families Bombacaceae, Tiliaceae, and Sterculiaceae, which have always been considered closely allied to Malvaceae s.s., are not monophyletic groups. Thus, the Malvaceae can be expanded to include all of these families so as to compose a monophyletic group. Adopting this circumscription, the Malvaceae incorporate a much larger number of genera.[citation needed


This article is based on the second circumscription, as presented by the Angiosperm Phylogeny Website.[5] The Malvaceae s.l. (hereafter simply "Malvaceae") comprise nine subfamilies. A tentative cladogram of the family is shown below. The diamond denotes a poorly supported branching (<80%).

Byttnerioideae: 26 genera, 650 species, pan-tropical, especially South America

Grewioideae: 25 genera, 770 species, "pantropical"

Sterculioideae: 12 genera, 430 species, pan-tropical

temperate regions and Central America


Brownlowioideae: eight genera, about 70 species, especially palaeo-tropical

Helicteroideae: eight to 12 genera, 10 to 90 species, tropical, especially Southeast Asia

Malvoideae: 78 genera, 1,670 species, temperate to tropical

Bombacoideae: 12 genera, 120 species, tropical, especially Africa and America

Until recently, relationships between these subfamilies were either poorly supported or almost completely obscure. Continuing disagreements focused primarily on the correct circumscription of these subfamilies, including the preservation of the family Bombacaceae.[13] A study published in 2021 presented a fully resolved phylogenetic framework for Malvaceae s.l. using genomic data for all nine subfamilies.[14]

Regarding the traditional Malvaceae s.s., the subfamily Malvoideae approximately corresponds to that group.


245 genera are currently accepted.[15]


The relationships between the "core Malvales" families used to be defined on the basis of shared "malvean affinities". These included the presence of malvoid teeth, stems with mucilage canals, and stratified wedge-shaped phloem.[16] These affinities were problematic because they were not always shared within the core families.[17] Later studies revealed more unambiguous synapomorphies within Malvaceae s.l.. Synapomorphies identified within Malvaceae s.l. include the presence of tile cells, trichomatous nectaries, and an inflorescence structure called a bicolor unit.[18] Tile cells consist of vertically positioned cells interspersed between and dimensionally similar to procumbent ray cells. Evidence of Malvean wood fossils has confirmed their evolutionary link in Malvaceae s.l., as well as explained their diverse structures.[19] Flowers of Malvaceae s.l. exhibit nectaries consisting of densely arranged multicellular hairs resembling trichomes. In most of Malvaceae s.l., these trichomatous nectaries are located on the inner surface of the sepals, but flowers of the subfamily Tiliodeae also have present nectaries on the petals.[20] Malvean flowers also share a unifying structure known as a bicolor unit, named for its initial discovery in the flowers of Theobroma bicolor. The bicolor unit consists of an ordered inflorescence with determinate cymose structures. The inflorescence can branch off the main axis, creating separate orders of the flowers, with the main axis developing first. Bracts on the peduncle subtend axillary buds that become these lateral stalks. One bract within this whorl is a sterile bract. The bicolor unit is a variable structure in complexity, but the presence of fertile and sterile bracts is a salient characteristic.[21]


The English common name 'mallow' (also applied to other members of Malvaceae) comes from Latin

Arabic: ملوخية (mulukhiyah) and modern Hebrew: מלוחיה (molokhia).[22][23]


Alcea rosea, the hollyhock, is a common garden flower in Malvaceae
Pterospermum acerifolium

Most species are herbaceous plants or shrubs, but some are trees or lianas.

Leaves and stems

Stellate hairs on the underside of a dried leaf of Malva alcea

Leaves are generally alternate, often palmately lobed or compound and palmately veined. The margin may be entire, but when dentate, a vein ends at the tip of each tooth (malvoid teeth). Stipules are present. The stems contain mucous canals and often also mucous cavities. Hairs are common, and are most typically stellate.[citation needed] Stems of Bombacoideae are often covered in thick prickles.[24]


Close-up photograph of Theobroma cacao flowers.

The flowers are commonly borne in definite or indefinite axillary

carpels. The ovary
is superior, with axial placentation, with capitate or lobed stigma. The flowers have
nectaries made of many tightly packed glandular hairs, usually positioned on the sepals.[20]


Durian fruits

The fruits are most often

loculicidal capsules, schizocarps or nuts


Self-pollination is often avoided by means of

entomophilous (pollinated by insects). Bees from the tribe Emphorini of the Apidae (including Ptilothrix, Diadasia, and Melitoma
) are known to specialize on the plants.


A number of species are pests in

baobabs are edible, as is the fruit of the durian. A number of species, including Hibiscus syriacus, Hibiscus rosa-sinensis and Alcea rosea
are garden plants.

See also


  1. ^ "Malvales". Retrieved 2023-07-20.
  2. .
  3. ^ "Family: Malvaceae". Germplasm Resources Information Network (GRIN) [Online Database]. United States Department of Agriculture Agricultural Research Service, Beltsville, Maryland. 17 January 2017. Retrieved 7 June 2017.
  4. .
  5. ^ a b "Angiosperm Phylogeny Website". Retrieved 15 July 2014.
  6. ^ Hibiscus L. Plants of the World Online. Retrieved 26 June 2023.
  7. ^ Pavonia Cav. Plants of the World Online. Retrieved 26 June 2023.
  8. ^ Sida L. Plants of the World Online. Retrieved 26 June 2023.
  9. ^ Ayenia L. Plants of the World Online. Retrieved 26 June 2023.
  10. ^ Dombeya Cav. Plants of the World Online. Retrieved 26 June 2023.
  11. ^ Sterculia L. Plants of the World Online. Retrieved 26 June 2023.
  12. .
  13. .
  14. .
  15. ^ Malvaceae Juss. Plants of the World Online. Retrieved 20 June 2023.
  16. ^ Kubitzki, K. (2003). The Families and Genera of Vascular Plants. Berlin: Springer-Verlag.
  17. S2CID 40348158
  18. .
  19. IAWA Bulletin
    : 23–28 – via ResearchGate.
  20. ^ .
  21. ^ .
  22. ^ a b Douglas Harper. "mallow". Online Etymology Dictionary. Retrieved February 3, 2012.
  23. ^ Khalid. "Molokheya: an Egyptian National Dish". The Baheyeldin Dynasty. Retrieved September 10, 2011.
  24. .

External links