Ancient North Eurasian

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Ancient North Eurasian
Mal'ta–Buret' culture ivory figurines (c. 24,000 BP-c. 15,000 BP). Some of the figurines wear hooded overalls with decorative stripes.[1] [2]
Approximate location of the Ancient North Eurasians c. 24,000~16,000 BP.[3][4][5]

In

Early East Eurasian' population (represented by the Tianyuan man, c. 40,000 BP).[8] The Ancient North Eurasians derive around 2/3 from an Early West Eurasian lineage and around 1/3 of their ancestry from an Early East Eurasian lineage.[a][b][9][10][11][12][13][14][15]

Around 20,000 to 25,000 years ago, a branch of Ancient North Eurasian people mixed with

Ancient East Asians, which led to the emergence of Ancestral Native American, Ancient Beringian and Ancient Paleo-Siberian populations. It is unknown exactly where this population admixture took place, and two opposing theories have put forth different migratory scenarios that united the Ancient North Eurasians with ancient East Asian populations.[16]

ANE ancestry has spread throughout Eurasia and the Americas in various migrations since the Upper Paleolithic, and more than half of the world's population today derives between 5 and 42% of their genomes from the Ancient North Eurasians.[17] Significant ANE ancestry can be found in Native Americans, as well as in regions of northern Europe, South Asia, Central Asia, and Siberia. It has been suggested that their mythology may have featured narratives shared by both Indo-European and some Native American cultures, such as the existence of a metaphysical world tree and a fable in which a dog guards the path to the afterlife.[18]

Genetic studies

Definition

The ANE lineage is defined by association with the "

Mal'ta boy" (MA-1), the remains of an individual who lived during the Last Glacial Maximum, 24,000 years ago in central Siberia, discovered in the 1920s. Together with the Yana Rhinoceros Horn Site samples, and Afontova Gora individuals, they are collectively referred to as 'Ancient North Siberians', although 'Ancient North Eurasian' is also used as collective name for both MA-1 and Yana remains.[19][20][21]

Mal'ta boy (MA-1), dated 24,000 BP, with tomb artifacts, Hermitage Museum (Hall 11),[22] Saint-Petersburg.[23]

The Ancient North Eurasians represent a distinct cluster of genetic diversity within the larger Eurasian gene pool, forming an early Siberian variation of modern humans, taking up an 'intermediate position' between West and East Eurasians.[24][25] It is suggested that the ANE ancestry found among modern human populations was largely contributed from a population linked to Afontova Gora (AG-3), rather than Malta (MA-1) or Yana.[26]

Formation

A qpGraph by Allentoft et al. 2024, showing the formation of Ancient North Siberians/Eurasians (ANS/ANE) and their subsequent contribution to Western Eurasian lineages.[14]
Formation of Ancient North Eurasians in the context of West and East Eurasian Core populations[21]

The formation of the Ancient North Eurasian/Siberian gene pool likely occurred very early by the combination of an '

Sunghir, or the Peștera Muierii woman,[28] while their East Eurasian-related component can be associated with ancestry found among a population related to the Paleolithic Tianyuan man, who is basal to contemporary East/Southeast Asians.[29][30][31][32][33][34][c][d][35][13][14]

A different but geographically close specimen, known as the Salkhit individual (c. 34,000 BP) from Northern Mongolia, was found to have approximately 83% Tianyuan-like ancestry and 17% Kostenki14-like ancestry, being the to date oldest found representative of contact between West and East Eurasians. The relationship between the Salkhit individual and Ancient North Eurasians is described as complex, with evidence for bidirectional geneflow between them.[36][25]

Lipson and Reich (2017) modeled the Malta sample to derive ancestry from a West Eurasian source, with additional admixture from a lineage related to East Asians (represented by

Kostenki-14, contributing around 60-68% ancestry, and from a lineage contemporary to the 'Basal-East Asian' Tianyuan man, contributing between 32-40%, while finding no evidence for a reversed geneflow from ANE into Tianyuan or modern East Asians.[37][38] Other studies could reproduce significant geneflow between these two sources and increased affinity for the ANE to the Tianyuan man (or other Upper Paleolithic East/Southeast Asian specimens), and also confirmed higher affinity for Eastern European Hunter-Gatherers (EHG) to the Tianyuan man, explained by them having received significant amounts of ANE ancestry.[29] The 'Basal East Asian' (Tianyuan-like) ancestry among EHGs (Sidelkino) has been estimated to be around 12,9–31%.[39][40]

Grebenyuk et al. summarized that the Ancient North Eurasians descended from the 'Ancient North Siberian' Yana population, which were "Early Upper Paleolithic tribes of hunters" and linked to similar groups associated with contemporaneous Southern Siberian sites. These communities of Southern Siberian and Central Asian hunters belonged to one of the earliest migration waves of the anatomically modern humans into Siberia. The authors summarized that "the initial peopling of Northeastern Asia by the anatomically modern humans could have happened both from West to East and from South to North".[41] Sikora et al. notes that the Ancient North Eurasians (Malta and Afontova Gora individuals) are unlikely to be direct descendants of the 'Ancient North Siberian' Yana population; rather, the study argues, both are sister lineages sharing a common ancestor. According to Sikora et al., the Malta sample may additionally also have received some 'early Caucasus hunter-gatherer' geneflow (c. 11%).[20] This scenario is questioned by Maier et al. 2023, who state that this conclusion is contradicted by other published articles, and that the direction of gene flow as well as observed affinity between ANE and CHG populations cannot be demonstrated by analysis of admixture graphs, but need further investigation.[42]

Vallini et al. 2024 notes that the Ancient North Eurasian lineage (represented by the Mal'ta and Yana specimens) is taking up an intermediate position between 'Ancient West Eurasians' (represented by a

Kostenki-14-like lineage) and 'Ancient East Eurasians' (represented by a Tianyuan-like lineage), and is the result of a Paleolithic admixture event between West and East Eurasians deriving around 50% ancestry from each lineage respectively.[21]

Distribution

Mal'ta figurines" with facial features, and 3D rendering.[43]

By c. 32kya, populations carrying ANE-related ancestry were probably widely distributed across northeast Eurasia. They may have expanded as far as Alaska and the Yukon, but were forced to abandon high latitude regions following the onset of harsher climatic conditions that came with the Last Glacial Maximum.[44]

Populations genetically similar to MA-1 and Afontova Gora were an important genetic contributor to

South Asians, and some East Asian groups, in order of significance.[45]
Lazaridis et al. (2016:10) note "a cline of ANE ancestry across the east-west extent of Eurasia". A 2016 study found that the global maximum of ANE ancestry occurs in modern-day Kets, Mansi, Native Americans, and Selkups.[6][45]

Deer tooth pendant of an ANE woman, from Denisova Cave, dated circa 24,700 years BP.[46]

The ancient Bronze-age-steppe

Afanasevo cultures were found to have a significant ANE-like component at c. 25–50% via their EHG ancestry.[47][48] According to Moreno-Mayar et al. 2018 between 14% and 38% of Native American ancestry may originate from gene flow from the Mal'ta–Buret' (ANE) population. This difference is caused by the penetration of posterior "Neo-Siberian" migrations into the Americas, with the lowest percentages of ANE ancestry found in Inuit and Alaskan Natives, as these groups are the result of migrations into the Americas roughly 5,000 years ago.[49] Estimates for ANE ancestry among first wave Native Americans show higher percentages,[50] such as 42% for those belonging to the Andean region in South America.[50] The other gene flow in Native Americans (the remainder of their ancestry) was of an East Asian-related origin, specifically diverged from other East Asians c. 30,000 years ago.[31] Gene sequencing of another south-central Siberian people (Afontova Gora-2) dating to approximately 17,000 years ago, revealed similar autosomal genetic signatures to that of Mal'ta boy-1, suggesting that the region was continuously occupied by humans throughout the Last Glacial Maximum.[31]

Genomic studies also indicate that the ANE component was brought to Western Europe by people related to the

Eastern European Hunter-Gatherers, which resulted in populations such as Scandinavian Hunter-Gatherers. [51] Western Hunter-Gatherers of the Villabruna cluster also carried the Y-haplogroup R1b, derived from the Ancient North Eurasian haplogroup R*, indicating "an early link between Europe and the western edge of the Steppe Belt of Eurasia."[52]

A deer tooth pendant impregnated with the genetic material of an ANE woman was found in the Denisova Cave, and dated to circa 24,700 years before present. She is closely related to Mal'ta and Afontova Gora specimens, found further east.[46]

An early Neolithic Central Asian specimen (Tutkaul1) from Tajikistan was found to be primarily derived from Ancient North Eurasians with some additional Neolithic Iranian-related inputs. The sample is closely related to Afontova Gora 3 (AG3) and Mal’ta 1, as well as to the West Siberian hunter-gatherers (Tyumen and Sosnoviy). While the sample also displays affinity for Eastern hunter-gatherers (EHGs), AG3 was found to be closer to EHGs than Tutkaul1, who instead may be a good proxy for ANE-related ancestry among ancient populations from the Iran and the Turan region.[53]

The Ancient Tianyuan Man and modern East/Southeast Asian populations were found to lack Upper Paleolithic Western Eurasian or ANE-related admixture, suggesting "resistance of those groups to the incoming UP population movements", or alternatively a subsequent reexpansion from a genetically East Asian-like population reservoir.[25]

Groups partially derived from the Ancient North Eurasians

Native American contribution

Main article: Genetic history of the Indigenous peoples of the Americas
The "Ancient North Eurasian" (ANE) network, consisted of several Paleolithic Siberian samples and contributed ancestry towards a wide variety of populations across Eurasia.
A qpGraph model by Maier et al. 2023, showing the possible formation of Ancient North Siberians/Eurasians (ANS/ANE) and their subsequent contribution to the Ancient Paleo-Siberians and Native Americans.[54]

According to Jennifer Raff, the Ancient North Eurasian population mixed with a daughter population of ancient East Asians, who they encountered around 25,000 years ago, which lead to the emergence of Native American ancestral populations. However, the exact location where the admixture took place is unknown, and the migratory movements that united the two populations are a matter of debate.[16]

One theory supposes that Ancient North Eurasians migrated south to

Southern Siberia, where they would have encountered and mixed with ancient East Asians. Genetic evidence from Lake Baikal in Mongolia supports this area as the location where the admixture took place.[55]

However, a third theory, the "Beringian standstill hypothesis", suggests that East Asians instead migrated north to Northeastern Siberia, where they mixed with ANE, and later diverged in Beringia, where distinct Native American lineages formed. This theory is supported by

paternal DNA evidence, which may reflect different population histories for paternal and maternal lineages in Native Americans, which is not uncommon and has been observed in other populations.[58]

The descendants of admixture between ANE and ancient East Asians include

Ancient Paleo-Siberians, populations represented by the Late Upper Paeolithic Lake Baikal Ust'Kyakhta-3 (UKY) 14,050-13,770 BP. They carried 30% ANE ancestry and 70% East Asian ancestry.[61]

Jōmon people, the pre-Neolithic population of Japan, mainly derived their ancestry from East Asian lineages, but also received geneflow from the ANE-related "Ancient North Siberians" (represented by samples from the Yana Rhinoceros Horn Site) prior to the migration from the Asian mainland to the Japanese archipelago. Jōmon ancestry is still found among the inhabitants of present-day Japan: most markedly among the Ainu people, who are considered the direct descendants of the Jōmon people, and to a small, but significant degree among the majority of the Japanese population.[62][63]

Siberian and Asian Holocene populations

Altai hunter-gatherer is the name given to Middle Holocene Siberian hunter-gatherers within the

Malta-Buret people.[65]

West Siberian Hunter-Gatherer (WSHG) is a specific archaeogenetic lineage that was first reported by Narasimhan et al. (2019). It can be modeled as 20% EHG, 73% ANE and 6%

Urals dated ca. 5,000 BCE, high-levels of WSHG-like ancestry can be detected in various populations of Central Asia until the Bronze Age. The population of the Botai culture, while probably not directly descended from WSHG, displays a high affinity with the WSHG lineage.[66] The European-Siberian cline defined by Eastern hunter-gatherer-like ancestry streched from Central Europe to Siberia and was already established 10,000 years ago, including the West Siberian hunter-gatherers, all deriving their ancestry primarily from Paleolithic Siberians (ANE).[67]

"Princess of Xiaohe", one of the Tarim mummies, the "best representatives" of Ancient North Eurasians.[68]

Lake Baikal Holocene - Among the Ancient Northeast Asians (ANA) of the Neolithic to Early Bronze Age period, Baikal Eneolithic (Baikal_EN) and Baikal Early Bronze Age (Baikal_EBA) derived 6.4% to 20.1% ancestry from ANE, while the rest of their ancestry was derived from ANA. Fofonovo_EN near by Lake Baikal were mixture of 12-17% ANE ancestry and 83-87% ANA ancestry.[69]

Tarim mummies

A 2021 genetic study on the

Baikal populations).[68]

The Tarim mummies are thus one of the rare

Mal'ta and Afontova Gora populations), despite their distance in time (around 14,000 years).[68] Having survived in a type of "genetic bottleneck" in the Tarim basin where they preserved and perpetuated their ANE ancestry, the Tarim mummies, more than any other ancient populations, can be considered as "the best representatives" of the Ancient North Eurasians.[68]

West Asian populations

Mesolithic Iranian hunter-gatherers and Neolithic Iranian farmers as well as

Kostenki-14; WEC), with the WEC2 component staying in the region of the Iranian Plateau, while the proper WEC component expanded into Europe.[71]

European populations

Further information: Eastern Hunter-Gatherer, Scandinavian Hunter-Gatherer, and Western Steppe Herders
Eastern Hunter Gatherers (EHGs) are the population with the highest genetic affinity in Europe and the main source for ANE-like ancestry among modern Europeans.
Modeling of modern European populatios, plotting them according to their proportions of ancestry from each of three inferred ancestral populations; Early European Farmers, Ancient Northern Euroasians and West European Hunter Gathers, as inferred from the model-based analysis.[72]

Lazaridis et al. (2014) detected ANE ancestry among modern European populations in proportions up to 20%.

Yamnaya people[47] but not of Western or Central Europeans predating the Corded Ware culture:[74] ANE ancestry was introduced in the European gene pool with the Eastern Hunter-Gatherer (EHG) lineage which derived significant ancestry from the ANE, c. 70%, with the remaining ancestry from a group more closely related to, but distinct from, Western Hunter-Gatherers (WHGs).[45][75][53] It is represented by multiple individuals, such as from Yuzhny Oleny in Karelia, one of Y-haplogroup R1a-M417, dated c. 8.4 kya, the other of Y-haplogroup J, dated c. 7.2 kya; and one individual from Samara, of Y-haplogroup R1b-P297, dated c. 7.6 kya, as well as individuals from Sidelkino and Popovo. After the end of the Last Glacial Maximum, the Western Hunter-Gatherers (WHG) and EHG lineages merged in Eastern Europe, accounting for early presence of ANE-derived ancestry in Mesolithic Europe. Evidence suggests that as Ancient North Eurasians migrated westward from Eastern Siberia, they absorbed Western Hunter-Gatherers and other West Eurasian populations as well.[76][53]

Scandinavian Hunter-Gatherer (SHG) is represented by several individuals buried at Motala, Sweden ca. 6000 BC. They were descended from Western Hunter-Gatherers who initially settled Scandinavia from the south, and received later admixture from EHG who entered Scandinavia from the north through the coast of Norway.[72][47][77][51][78]

Western Steppe Herders (WSH) is the name given to a distinct ancestral component that represents descent closely related to the Yamnaya culture of the Pontic–Caspian steppe.[e] This ancestry is often referred to as Yamnaya ancestry or Steppe ancestry, and was formed from EHG and CHG (Caucasus hunter-gatherer) in about equal proportions.[30]

Phenotype prediction

Genomic studies by Raghavan et al. (2014) and Fu et al. (2016) suggested that Mal'ta boy may have had brown eyes, and relatively dark hair and dark skin,[80][81] while cautioning that this analysis was based on an extremely low coverage of DNA that might not give an accurate prediction of pigmentation.[82] Mathieson, et al. (2018) could not determine if Mal'ta 1 boy had the derived allele associated with blond hair in ANE descendants, as they could obtain no coverage for this SNP.[83]

Anthropologic research

One of the Tarim mummies, a "Beauty of Loulan" dated c. 2000 BCE

Kozintsev (2020) argues that the historical Southern Siberian

Okunevo population, which derives most of their ancestry from Ancient North Eurasians and their closest relatives, as possessing a distinct craniometric phenotype, which he dubbed "Americanoid", which represents the variation of the first humans in Siberia. He further argues that "As the geography and chronology of the ANE component show, it is misleading to describe it as Western Eurasian and associate it solely with ancient Caucasoids. To all appearances, it emerged before the Caucasoid-Mongoloid split."[48]

Zhang et al. (2021) proposed that the 'Western' like features of the earlier Tarim mummies could be attributed to their Ancient North Eurasian ancestry.[84] Previous craniometric analyses on the early Tarim mummies found that they formed their own cluster, and clustered with neither European-related Steppe pastoralists of the Andronovo and Afanasievo cultures, nor with inhabitants of the Western Asian BMAC culture, nor with East Asian populations further east.[85]

Evolution of blond hair

Blond hair is associated with a

Mal'ta boy lacks the sequence coverage to make this determination).[91] The allele then appears later in ANE-derived Eastern Hunter-Gatherer (EHG) populations at Samara, Motala and Ukraine, circa 10,000 BP, and then in populations with Steppe ancestry.[83] Mathieson, et al. (2018) thus argued that this allele originated in the Ancient North Eurasian population, before spreading to western Eurasia.[83]

Geneticist

Yamnayas, were responsible for transmitting this gene to Europeans.[86] The gene was also found among the Tarim mummies.[93]

The mutation for blond hair is thought to have originated among the Afontova Gora population of the Ancient North Eurasian (ANE) cline of south-central Siberia

Comparative mythology

Mal'ta–Buret' culture centrally perforated ivory plaques with abstract circles, and three snakes[1] According to archeologist Don Hitchcock "the snake is rare in northern hemisphere Paleolithic art, presumably because the cold conditions precluded a wide distribution of snakes. In addition, it can be seen that the snakes have very broad heads, as though they belong to the Cobra group - yet Cobras are now known only in southern asian localities." It has yet to be clarrified how the creators of these ivory plaques know snakes, or if there are other possible interpretations for these.[94]

Since the term 'Ancient North Eurasian' refers to a genetic bridge of connected mating networks, scholars of comparative mythology have argued that they probably shared myths and beliefs that could be reconstructed via the comparison of stories attested within cultures that were not in contact for millennia and stretched from the Pontic–Caspian steppe to the American continent.[18]

The

Sarvarā, and Garmr. In Zoroastrianism, two four-eyed dogs guard the bridge to the afterlife called Chinvat Bridge. Anthony and Brown note that it might be one of the oldest mythemes recoverable through comparative mythology.[18]

A second canid-related series of beliefs, myths and rituals connected dogs with healing rather than death. For instance, Ancient Near Eastern and Turkic-Kipchaq myths are prone to associate dogs with healing and generally categorised dogs as impure. A similar myth-pattern is assumed for the Eneolithic site of Botai in Kazakhstan, dated to 3500 BC, which might represent the dog as absorber of illness and guardian of the household against disease and evil. In Mesopotamia, the goddess Nintinugga, associated with healing, was accompanied or symbolized by dogs. Similar absorbent-puppy healing and sacrifice rituals were practiced in Greece and Italy, among the Hittites, again possibly influenced by Near Eastern traditions.[18]

See also

References

  1. ^
    doi:10.3390/arts2020046
    .
  2. S2CID 231980510
    .
  3. ^ "Paleolithic to Bronze Age Siberians Reveal Connections with First Americans and across Eurasia" (PDF). CELL. 2020.
  4. ISSN 2770-5005
    .
  5. S2CID 235460793
    .
  6. ^ a b c d Flegontov et al. 2016.
  7. ^ Jeong et al. 2019
  8. ISBN 978-3-030-90060-1
    .
  9. PMID 36859553
    .
  10. S2CID 174809069
    .
  11. ^ Vallini et al. 2022, Supplementary Information, p. 17.
  12. PMID 28074030
    .
  13. ^
    PMID 30415837
    .
  14. ^
    PMID 38200295
    .
  15. ^
    ISSN 2500-2295
    .
  16. ^
    ISBN 978-1-5387-4970-8
    .
  17. ^ Reich 2018, p. 81
  18. ^ a b c d Anthony & Brown 2019, pp. 104–106.
  19. S2CID 235460793
    .
  20. ^
    S2CID 174809069
    .
  21. ^
    PMID 38528002
    .
  22. ^ "Hall 11 panorama". State Hermitage Museum.
  23. S2CID 231980510
    .
  24. ISSN 1563-0110
    .
  25. ^ a b c d Vallini et al. 2022
  26. PMID 27135931
    .
  27. S2CID 250502011
    .
  28. S2CID 234793812
    .
  29. ^
    PMID 36859553
    .
  30. ^ a b Jeong et al. 2019.
  31. ^ a b c Raghavan et al. 2013.
  32. ^ Balter 2013.
  33. ISSN 2770-5005
    .
  34. S2CID 174809069
    .
  35. ^
    PMID 28074030
    .
  36. PMID 33122380
    .
  37. S2CID 218649510
    .
  38. PMID 29033327
    .
  39. S2CID 263672903
    .
  40. PMID 29545507
    .
  41. ^
    ISBN 978-3-030-90061-8
  42. ^ Maier et al. 2023.
  43. ^ "Figurine of a woman with the "Spanish rose" hairdo". Art of Mal'ta. Novosibirsk State University.
  44. S2CID 249355636
    .
  45. ^ a b c Lazaridis et al. 2016.
  46. ^
    PMID 37138083
    .
  47. ^ a b c d e Haak et al. 2015.
  48. ^
    ISSN 1563-0110
    .
  49. ^ Moreno-Mayar et al. 2018.
  50. ^ a b Wong et al. 2017.
  51. ^ a b Günther et al. 2018.
  52. doi:10.24412/2500-2295-2023-3-95-117
    .
  53. ^
    PMID 36859578
    .
  54. PMID 37057893
    .
  55. ^ Raff 2022, p. 188
  56. ^ Raff 2022, pp. 188–189
  57. ISBN 978-3-030-90060-1
    .
  58. S2CID 3519553
    .
  59. ^ Moreno-Mayar et al. 2018.
  60. PMID 32843717
    .
  61. ^ Yu et al. 2020.
  62. ^ Osada & Kawai 2021.
  63. PMID 34533991
    .
  64. S2CID 255750546
    .
  65. doi:10.17746/1563-0110.2021.49.4.140-151
    .
  66. ^ Narasimhan 2019.
  67. PMID 37558731
    .
  68. ^
    PMID 34707286
    .
  69. ^ Jeong et al. 2020.
  70. PMID 38200295
    .
  71. PMID 38528002
    .
  72. ^ a b Lazaridis et al. 2014.
  73. PMID 25230663
    .
  74. ^ Lazaridis, Patterson & Mittnik 2014.
  75. ^ Wang 2019.
  76. ^ Mathieson et al. 2018.
  77. ^ Mathieson 2015.
  78. ^ Mittnik 2018.
  79. ^ Jeong et al. 2018.
  80. ^ Raghavan & Skoglund et al. 2014.
  81. PMID 27135931
    .
  82. PMID 24256729
    .
  83. ^ a b c Mathieson et al. 2018 "Supplementary Information page 52
  84. PMID 34707286
    .
  85. ^ "A Craniometric Investigation of The Bronze Age Settlement of Xinjiang - Important | PDF". Scribd. Retrieved 4 April 2023.
  86. ^ a b Carlberg & Hanel 2020.
  87. ^ Hartz P. "KIT LIGAND; KITLG". Online Mendelian Inheritance in Man.
  88. ^ Khan R. "KITLG makes you white skinned?". Discover Magazine.
  89. ^ "P21583". Uniprot.
  90. ^ Guenther et al. 2014.
  91. ISBN 978-1-78607-623-6
    .
  92. ISBN 978-0-19-882125-0
    .
  93. PMID 34707286
    .
  94. ^ "The Mal'ta - Buret' venuses and culture in Siberia". donsmaps.com.

Notes

  1. ^ Vallini et al. (2022) created a TreeMix model in which "admixture events in varying proportions between sister groups of Kostenki14 and Tianyuan is supported, and can indeed explain this observation (fig. 1A, purple leaves; Supplementary fig. S5 and Section 3.6, Supplementary Material online)".
  2. ^ Sikora et al. (2019) model the Yana individuals as 22% East Eurasian and the remainder West Eurasian ("Using admixture graphs and outgroup-based estimation of mixture proportions, we find that ANS can be modelled as early West Eurasian with an approximately 22% contribution from early East Asians"). Massilani et al. (2020) model the Yana individuals as around one-third East Eurasian and two-thirds West Eurasian.Vallini et al. (2022) model Yana as 50% West Eurasian and 50% East Eurasian.
  3. ^ Vallini et al. (2022) created a TreeMix model in which "admixture events in varying proportions between sister groups of Kostenki14 and Tianyuan is supported, and can indeed explain this observation (fig. 1A, purple leaves; Supplementary fig. S5 and Section 3.6, Supplementary Material online)".
  4. ^ Sikora et al. (2019) model the Yana individuals as 22% East Eurasian and the remainder West Eurasian. Massilani et al. (2020) model the Yana individuals as around one-third East Eurasian and two-thirds West Eurasian.Vallini et al. (2022) model Yana as 50% West Eurasian and 50% East Eurasian.
  5. ^ "Recent paleogenomic studies have shown that migrations of Western steppe herders (WSH) beginning in the Eneolithic (ca. 3300–2700 BCE) profoundly transformed the genes and cultures of Europe and central Asia... The migration of these Western steppe herders (WSH), with the Yamnaya horizon (ca. 3300–2700 BCE) as their earliest representative, contributed not only to the European Corded Ware culture (ca. 2500–2200 BCE) but also to steppe cultures located between the Caspian Sea and the Altai-Sayan mountain region, such as the Afanasievo (ca. 3300–2500 BCE) and later Sintashta (2100–1800 BCE) and Andronovo (1800–1300 BCE) cultures."[79]

Bibliography

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