Pholidosauridae

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Pholidosauridae
Temporal range:
Ma
Possible Middle Jurassic (Bathonian) and Paleocene (Danian) records.
Oceanosuchus
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauria
Clade: Pseudosuchia
Clade: Crocodylomorpha
Clade: Crocodyliformes
Suborder: Tethysuchia
Family: Pholidosauridae
Zittel and Eastman, 1902
Subgroups
Pholidosaurus purbeckensis fossil.

Pholidosauridae is an extinct

crocodylomorphs. Fossils have been found in Europe (Denmark, England, France, Germany, Spain and Sweden), Africa (Algeria, Niger, Mali, Morocco and Tunisia), North America (Canada and the United States) and South America (Brazil and Uruguay). The pholidosaurids first appeared in the fossil record during the Bathonian stage of the Middle Jurassic. Jouve & Jalil (2020) described postcranial material of a pholidosaurid from the Paleocene (Danian) of Ouled Abdoun Basin (Morocco), representing the most recent record of the family. The authors also reinterpreted putative Maastrichtian dyrosaurid Sabinosuchus as a pholidosaurid, and argued that at least two independent pholidosaurid lineages reached the Maastrichtian, among which one survived the Cretaceous–Paleogene extinction event.[1] Before the publication of this study it was thought that the family became extinct during the Late Turonian stage of the Late Cretaceous.[2]

). The Cenomanian Terminonaris was the Pholidosaurid species that appeared to be the most common during the Late Cretaceous.[8]

Phylogeny

Pholidosauridae is usually considered to be most closely related to the

polyphyletic[10] in relation to Dyrosauridae. For example, Fortier, Perea & Schultz (2011) found the family to be monophyletic, and include to main lineages: the ElosuchusMeridiosaurus lineage and the Pholidosaurus lineage. The cladogram below shows their phylogenetic analysis, which is based on an expanded version of Pol and Gasparini (2009) analysis.[2]

Neosuchia
Atoposauridae

Alligatorium

Theriosuchus pusillus

Goniopholididae

Bernissartia fagesii

Eusuchia

Thalattosuchia

Pholidosauridae

Pholidosaurus purbeckensis

Sarcosuchus imperator

Terminonaris robusta

Oceanosuchus boecensis

Elosuchus cherifiensis

Meridiosaurus vallisparadisi

Dyrosauridae

Sokotosuchus ianwilsoni

Dyrosaurus phosphaticus

Hyposaurus rogersii

Rhabdognathus

de Andrade et al. (2011) recovered a

paraphyletic traditional Pholidosauridae. In their analysis the "Elosuchus lineage" was found to be basal to the "Pholidosaurus lineage"+Dyrosauridae. They used the name Elosuchidae for the Elosuchus lineage and restricted Pholidosauridae to its type genus. The following cladogram simplified after their analysis.[9]

Neosuchia
Atoposauridae

Theriosuchus pusillus

Theriosuchus guimarotae

Stolokrosuchus

Goniopholididae

Thalattosuchia

Tethysuchia
Elosuchidae

Sarcosuchus hartti

Sarcosuchus imperator

Pholidosauridae

Pholidosaurus schaumburgensis

Dyrosauridae

Congosaurus

Guarinisuchus

Dyrosaurus maghribensis

Dyrosaurus phosphaticus

References

  1. ^ .
  2. ^ .
  3. .
  4. .
  5. ^ Buffetaut, E. (2007). "The spinosaurid dinosaur Baryonyx (Saurischia, Theropoda) in the Early Cretaceous of Portugal." Geological Magazine, 144(6): 1021-1025.
  6. ISSN 1175-5334
    .
  7. .
  8. ^ .
  9. .

External links