List of crocidurines

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Gray shrew
Greater white-toothed shrew (Crocidura russula)

Soricinae. They are found in Africa, Europe, and Asia, primarily in forests, savannas, shrublands, and grasslands, though some species can also be found in wetlands, rocky areas, or coastal areas. They range in size from the Etruscan shrew, one of the smallest mammal species at 3 cm (1 in) plus a 2 cm (1 in) tail,[1] to the hero shrew and Thor's hero shrew, at 15 cm (6 in) plus a 10 cm (4 in) tail. Crocidurines primarily eat invertebrates, small mammals, frogs, toads, and lizards. Almost no crocidurines have population estimates, but 22 species are categorized as endangered species, while the Andaman shrew, Christmas Island shrew, Harenna shrew, Jenkins's shrew, Nicobar shrew, and Wimmer's shrew are categorized as critically endangered
.

The 235 extant species of Crocidurinae are divided into nine genera; 192 of them are in Crocidura, the most species of any mammal genus.[2] A few extinct prehistoric Crocidurinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[3]

Conventions

 CR 
Critically Endangered (6 species)
 EN Endangered (22 species)
 VU Vulnerable (12 species)
 NT Near threatened (8 species)
 LC Least concern (126 species)
Other categories
 DD Data deficient (60 species)
 NE Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the crocidurine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

The

Solisorex, with one species, Suncus, containing nineteen species; and Sylvisorex
, containing fifteen species.

Subfamily

Crocidurinae

  • Genus Crocidura (white-toothed shrews): 191 species
  • Genus Diplomesodon (piebald shrew): one species
  • Genus
    Feroculus
     (Kelaart's long-clawed shrew): one species
  • Genus Paracrocidura (large-headed shrews): three species
  • Genus
    Ruwenzorisorex
     (Ruwenzori shrew): one species
  • Genus Scutisorex (hero shrews): two species
  • Genus
    Solisorex
     (Pearson's long-clawed shrew): one species
  • Genus Suncus (musk shrews and pygmy shrews): nineteen species
  • Genus Sylvisorex (forest shrews): fifteen species
Crocidurinae[4]
  

Feroculus

Solisorex

Crocidurines

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[5]

Genus Crocidura – Wagler, 1832 – 192 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
African black shrew


C. nigrofusca
Matschie, 1895 		Central and eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[7]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[7]

African dusky shrew


C. caliginea
Hollister, 1916 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[8]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[8]

African giant shrew

C. olivieri
(Lesson, 1827)

20 subspecies
  • C. o. anchietae
  • C. o. bueae
  • C. o. cara
  • C. o. cinereoaenea
  • C. o. darfurea
  • C. o. giffardi
  • C. o. guineensis
  • C. o. hansruppi
  • C. o. hedenborgiana
  • C. o. kivu
  • C. o. manni
  • C. o. martiensseni
  • C. o. nyansae
  • C. o. occidentalis
  • C. o. odorata
  • C. o. olivieri
  • C. o. spurelli
  • C. o. sururae
  • C. o. toritensis
  • C. o. zuleika
Sub-Saharan Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, savanna, and shrubland[9]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[9]

Andaman shrew


C. andamanensis
Miller, 1902 		South Andaman Island in India
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[10]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[10]

Andaman spiny shrew


C. hispida
Thomas, 1913 		South Andaman Island in India
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[11]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Unknown[11]

Annamite shrew


C. annamitensis
Jenkins, Lunde, & Moncrieff, 2009 		Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[12]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[12]

Ansell's shrew


C. ansellorum
Hutterer & Dippenaar, 1987 		South-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[13]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[13]

Arabian shrew


C. arabica
Hutterer, 1988 		Oman and Yemen
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[14]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[14]

Armenian shrew


C. armenica
Gureev, 1963 		Armenia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas[15]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[15]

Asian gray shrew


C. attenuata
H. Milne-Edwards, 1872 		Eastern and southeastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[16]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[16]

Asian lesser white-toothed shrew

Brown shrew

C. shantungensis
Miller, 1901

Two subspecies
  • C. s. quelpartis
  • C. s. shantungensis
Eastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[17]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[17]

Bailey's shrew


C. baileyi
Osgood, 1936 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[18]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[18]

Bale shrew


C. bottegoides
Hutterer & Yalden, 1990 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and forest[19]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[19]

Banka shrew


C. vosmaeri
Jentink, 1888 		Bangka Island in Indonesia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[20]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[20]

Batak shrew


C. batakorum
Hutterer, 2007 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[21]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[21]

Bates's shrew


 C. batesi
(Dollman, 1915) 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[22]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[22]

Beccari's shrew


C. beccarii
Dobson, 1886 		Sumatra
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[23]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[23]

Beletta shrew


C. yaldeni
Lavrenchenko, Voyta, & Hutterer, 2016 		Ethiopia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[24]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[24]

Bicolored musk shrew


C. fuscomurina
(Heuglin, 1865) 		Sub-Saharan Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna, grassland, and desert[25]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[25]

Bicolored shrew

Brown shrew

C. leucodon
(Hermann, 1780) 		Europe and western Asia
Map of range 		Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[26]

Habitat: Shrubland and grassland[27]

Diet: Small mammals, frogs, toads, lizards, and invertebrates[26] 		 LC 


Unknown Unknown[27]

Black-footed shrew

Gray shrew

C. nigripes
Miller & Hollister, 1921

Two subspecies
  • C. n. lipara
  • C. n. nigripes
Sulawesi island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[28]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[28]

Blackish white-toothed shrew


C. nigricans
Bocage, 1889 		Angola
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[29]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[29]

Bornean shrew


C. foetida
Peters, 1870

Three subspecies
  • C. f. doriae
  • C. f. foetida
  • C. f. kelabit
Borneo
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[30]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[30]

Bottego's shrew


C. bottegi
Thomas, 1898 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[31]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[31]

Buettikofer's shrew


C. buettikoferi
Jentink, 1888 		West-central Afria
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[32]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[32]

Butiaba naked-tailed shrew


C. littoralis
Heller, 1910 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[33]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[33]

Cameroonian shrew


C. picea
Sanderson, 1940 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[34]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[34]

Canarian shrew

Gray shrew

C. canariensis
Hutterer, López-Jurado, & Vogel, 1987 		Canary Islands
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Coastal marine, desert, and shrubland[35]

Diet: Snails and insects[35] 		 EN 


Unknown Population declining[35]

Caspian shrew


C. caspica
Thomas, 1907 		Iran and Azerbaijan
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[36]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[36]

Chinese white-toothed shrew


C. rapax
Allen, 1923

Four subspecies
  • C. r. kurodai
  • C. r. lutaoensis
  • C. r. rapax
  • C. r. tadae
Eastern Asia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[37]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[37]

Christmas Island shrew


C. trichura
Dobson, 1889 		Christmas Island
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[38]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


0–200 Population declining[38]

Cinderella shrew


C. cinderella
Thomas, 1911 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[39]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[39]

Congo white-toothed shrew


C. congobelgica
Hollister, 1916 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[40]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[40]

Cranbrook's white-toothed shrew


C. cranbrooki
Jenkins, Lunde, & Moncrieff, 2009 		Northern Myanmar Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[41]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[41]

Cretan shrew


C. zimmermanni
Wettstein, 1953 		Island of Crete in Greece
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown and rocky areas[42]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[42]

Crosse's shrew


C. crossei
Thomas, 1895 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[43]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[43]

Cyrenaica shrew


C. aleksandrisi
Vesmanis, 1977 		Northern Libya
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[44]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[44]

Dent's shrew

Black and white picture of a schrew

C. denti
Dollman, 1915 		Western and central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[45]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[45]

Desert musk shrew


C. smithii
Thomas, 1895

Two subspecies
  • C. s. debalsaci
  • C. s. smithii
Western and Eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[46]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[46]

Desperate shrew


C. desperata
Hutterer, Jenkins, & Verheyen, 1991 		Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[47]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Population declining[47]

Dhofar shrew


C. dhofarensis
(Hutterer, 1988) 		Yemen and Oman Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[48]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[48]

Doucet's musk shrew


C. douceti
Heim de Balsac, 1958 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[49]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[49]

Dsinezumi shrew

Drawing of gray shrew

C. dsinezumi
(Temminck, 1842) 		Japan
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[50]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[50]

East African highland shrew


C. allex
Osgood, 1910 		Kenya and Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and inland wetlands[51]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[51]

Egyptian pygmy shrew


C. religiosa
(Geoffroy, 1827) 		Northern Egypt
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: [52]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[52]

Eisentraut's shrew


C. eisentrauti
Heim de Balsac, 1957 		Cameroon
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and forest[53]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Population steady[53]

Elgon shrew


C. elgonius
Osgood, 1910 		Kenya and Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[54]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[54]

Elongated shrew


C. elongata
Miller & Hollister, 1921 		Sulawesi island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[55]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[55]

Fingui white-toothed shrew


C. fingui
Ceríaco et al., 2015 		Príncipe in São Tomé and Príncipe Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[56]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[56]

Fischer's shrew


C. fischeri
Pagenstecher, 1885 		Kenya and Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[57]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[57]

Flat-headed shrew


C. planiceps
Heller, 1910 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[58]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[58]

Flower's shrew


C. floweri
Dollman, 1915 		Northern Egypt
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: [59]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[59]

Fox's shrew


C. foxi
Dollman, 1915 		Western and central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna and forest[60]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[60]

Fraser's musk shrew


C. poensis
(Fraser, 1843) 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and forest[61]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[61]

Gathorne's shrew


C. gathornei
Jenkins, 2013 		Himalayas in India Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[62]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[62]

Glass's shrew


C. glassi
Heim de Balsac, 1966 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[63]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Population declining[63]

Gmelin's white-toothed shrew


C. gmelini
Pallas, 1811 		Central Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[64]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[64]

Goliath shrew


C. goliath
Thomas, 1906 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[65]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[65]

Gracile naked-tailed shrew


C. maurisca
Thomas, 1904 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands and forest[66]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[66]

Grasse's shrew


C. grassei
Brosset, Dubost, & Heim de Balsac, 1965 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[67]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[67]

Greater Mindanao shrew


C. grandis
Miller, 1911 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[68]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[68]

Greater red musk shrew

Drawing of brown shrew

C. flavescens
(Geoffroy, 1827) 		Southern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna and grassland[69]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[69]

Greater white-toothed shrew

Brown shrews

C. russula
(Hermann, 1780) 		Europe and northwestern Africa
Map of range 		Size: 6–9 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[70]

Habitat: Shrubland[71]

Diet: Invertebrates, as well as small mammals and small lizards[70] 		 LC 


Unknown Population steady[71]

Greenwood's shrew


C. greenwoodi
Heim de Balsac, 1966 		Somalia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, savanna, and shrubland[72]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[72]

Guramba shrew


C. phaeura
Osgood, 1936 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[73]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Unknown[73]

Guy's shrew


C. guy
Jenkins, Lunde, & Moncrieff, 2009 		Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[74]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[74]

Güldenstädt's shrew


C. gueldenstaedtii
(Pallas, 1811) 		Europe and western Asia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail
amphipods, annelids, and small vertebrates[75]
 		 LC 


Unknown Unknown[75]

Hainan Island shrew


C. wuchihensis
Sung, 1966 		Southern China and northern Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[76]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[76]

Harenna shrew


C. harenna
Hutterer & Yalden, 1990 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[77]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[77]

Heather shrew


C. erica
Dollman, 1915 		Angola
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[78]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[78]

Hildegarde's shrew


C. hildegardeae
Thomas, 1904 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[79]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[79]

Hill's shrew


C. hilliana
Smith
, 1995 		Thailand and Laos
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[80]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[80]

Horsfield's shrew


C. horsfieldii
(Tomes, 1856) 		Scattered southern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[81]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[81]

Hun shrew


C. attila
Dollman, 1915 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[82]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[82]

Hutan shrew


C. hutanis
Ruedi & Vogel, 1995 		Sumatra island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[83]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[83]

Indochinese shrew


C. indochinensis
Kloss, 1922 		Southeastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[84]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[84]

Iranian shrew


C. susiana
Redding & Lay, 1978 		Iran
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[85]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[85]

Jackass shrew


C. arispa
Spitzenberger, 1971 		Turkey
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas[86]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[86]

Jackson's shrew


C. jacksoni
Thomas, 1904 		East-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[87]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[87]

Javan ghost shrew


C. umbra
Demos, Achmadi, Handika, Maharadatunkamsi, Rowe, & Esselstyn, 2016 		Java island in Indonesia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[88]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Unknown[88]

Javanese shrew


C. maxi
Sody, 1936 		Indonesia and East Timor
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[89]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[89]

Jenkins's shrew


C. jenkinsi
Chakraborty, 1978 		South Andaman Island in India
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[90]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[90]

Jouvenet's shrew


C. jouvenetae
Heim de Balsac, 1958 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[91]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[91]

Kashmir white-toothed shrew


C. pullata
Miller, 1911 		Northern India and Pakistan
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[92]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[92]

Katinka's shrew


C. katinka
Bate, 1937 		Western Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[93]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[93]

Kilimanjaro shrew


C. monax
Thomas, 1910
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[94]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[94]

Kinabalu shrew


C. baluensis
Thomas, 1898 		Northern Borneo
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[95]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[95]

Kivu long-haired shrew


C. lanosa
Heim de Balsac, 1968 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and inland wetlands[96]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Population declining[96]

Kivu shrew


C. kivuana
Heim de Balsac, 1968 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and inland wetlands[97]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Unknown[97]

Lamotte's shrew


C. lamottei
Heim de Balsac, 1968 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[98]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[98]

Large-headed forest shrew


C. grandiceps
Hutterer, 1983 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[99]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Unknown[99]

Latona's shrew


C. latona
Hollister, 1916 		Democratic Republic of the Congo
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[100]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[100]

Lesser Ryukyu shrew


C. watasei
Kuroda, 1924 		Southern Japan
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[101]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[101]

Lesser gray-brown musk shrew


C. silacea
Thomas, 1895 		Southern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas, grassland, shrubland, and savanna[102]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[102]

Lesser red musk shrew

Brown shrew

C. hirta
Peters, 1852 		Southern and southeastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna, grassland, and desert[103]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[103]

Lesser rock shrew


C. serezkyensis
Laptev, 1929 		Western Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[104]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[104]

Lesser white-toothed shrew

Gray shrew

C. suaveolens
(Pallas, 1811) 		Europe and Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, grassland, rocky areas, and coastal/supratidal marine[105]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[105]

Long-footed shrew


C. crenata
Brosset, Dubost, & Heim de Balsac, 1965 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[106]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[106]

Long-tailed musk shrew


C. dolichura
Peters, 1876 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[107]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[107]

Lucina's shrew


C. lucina
Dippenaar, 1980 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and inland wetlands[108]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Population steady[108]

Ludia's shrew


C. ludia
Hollister, 1916 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[109]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[109]

Luzon shrew


C. grayi
Dobson, 1890 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[110]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[110]

Lwiro shrew


C. lwiroensis
Peterhans & Hutterer, 2013 		Democratic Republic of the Congo Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[111]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[111]

MacArthur's shrew


C. macarthuri
St. Leger, 1934 		Eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[112]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[112]

MacMillan's shrew


C. macmillani
Dollman, 1915 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[113]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Unknown[113]

Makwassie musk shrew


C. maquassiensis
Roberts, 1946 		Southern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and rocky areas[114]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[114]

Malayan shrew


C. malayana
Robinson & Kloss, 1911 		Malaysia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[115]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[115]

Mamfe shrew


C. virgata
Sanderson, 1940 		West-central Africa Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[116]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[116]

Manenguba shrew


C. manengubae
Hutterer, 1982 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[117]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Population declining[117]

Mauritanian shrew


C. lusitania
Dollman, 1915 		Western and eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna and shrubland[118]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[118]

Mduma's shrew


C. mdumai
Stanley, Hutterer, Giarla, & Esselstyn, 2015 		Tanzania Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[119]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Unknown[119]

Mindanao shrew

Gray shrew

C. beatus
Miller, 1910 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[120]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[120]

Mindoro shrew


C. mindorus
Miller, 1910 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[121]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[121]

Montane white-toothed shrew


C. montis
Thomas, 1906 		Eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[122]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[122]

Moonshine shrew


C. luna
Dollman, 1910 		Southeastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[123]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[123]

Mossy forest shrew


C. musseri
Ruedi & Vogel, 1995 		Sulawesi island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[124]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[124]

Mount Nimba giant forest shrew


C. nimbasilvanus
Hutterer, 2003 		Western Africa Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[125]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[125]

Munissi's shrew


C. munissii
Stanley, Hutterer, Giarla, & Esselstyn, 2015 		Tanzania Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[126]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[126]

Narrow-headed shrew


C. stenocephala
Heim de Balsac & Dieterlen, 1979 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[127]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[127]

Negev shrew


C. ramona
Ivanitskaya, Shenbrot, & Nevo, 1996 		Israel
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Desert[128]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[128]

Negros shrew


C. negrina
Rabor, 1952 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[129]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[129]

Nicobar shrew


C. nicobarica
Miller, 1902 		Great Nicobar Island in India
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[130]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[130]

Nigerian shrew


C. nigeriae
Dollman, 1915 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[131]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[131]

Nimba shrew


C. nimbae
Heim de Balsac, 1956 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[132]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Population declining[132]

Niobe's shrew


C. niobe
Thomas, 1906 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[133]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[133]

North African white-toothed shrew

Brown shrew

C. pachyura
Küster, 1835 		Northern Africa and southern Europe
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[134]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[134]

Nyiro shrew


C. macowi
Dollman, 1915 		Kenya
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[135]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[135]

Oriental shrew

Brown shrew

C. orientalis
Jentink, 1890

Two subspecies
  • C. o. lawuana
  • C. o. orientalis
Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[136]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[136]

Palawan shrew


C. palawanensis
Taylor, 1934 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[137]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[137]

Pale gray shrew


C. pergrisea
Miller, 1913 		Pakistan
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[138]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[138]

Panay shrew


C. panayensis
Hutterer, 2007 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[139]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[139]

Peninsular shrew


C. negligens
Kloss, 1914 		Southeastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[140]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[140]

Peters's musk shrew


C. gracilipes
Peters, 1870 		Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[141]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[141]

Phan Luong's shrew


C. phanluongi
Jenkins, Abramov, Rozhnov, & Olsson, 2010 		Vietnam and Cambodia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[142]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[142]

Phu Hoc shrew


C. phuquocensis
Abramov, Jenkins, Rozhnov, & Kalinin, 2008 		Phú Quốc island in Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[143]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[143]

Pitman's shrew


C. pitmani
Barclay, 1932 		Zambia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[144]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[144]

Polia's shrew


C. polia
Hollister, 1916 		Democratic Republic of the Congo
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[145]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[145]

Rainey's shrew


C. raineyi
Heller, 1912 		Kenya
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[146]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[146]

Reddish-gray musk shrew


C. cyanea
(Duvernoy, 1838) 		Southern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, grassland, and caves[147]

Diet: Invertebrates[147] 		 LC 


Unknown Population steady[147]

Roosevelt's shrew


C. roosevelti
(Heller, 1910) 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[148]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[148]

Ryukyu shrew


C. orii
Kuroda, 1924 		Southern Japan
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[149]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[149]

Sa Pa shrew

C. sapaensis
Jenkins, Abramov, Bannikova, & Rozhnov, 2013 		Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[150]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[150]

Saharan shrew


C. tarfayensis
Vesmanis & Vesmanis, 1980 		Mauritania and Morocco
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas and desert[151]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Population declining[151]

Sahelian tiny shrew


C. pasha
Dollman, 1915 		Scattered northern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[152]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[152]

Savanna dwarf shrew


C. nanilla
Thomas, 1909 		Western and eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[153]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[153]

Savanna path shrew


C. viaria
(Geoffroy, 1834) 		Western, central, and eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[154]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[154]

Savanna shrew


C. fulvastra
(Sundevall, 1843) 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[155]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[155]

Savanna swamp shrew


C. longipes
Hutterer & Happold, 1983 		West-central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[156]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[156]

Siberian shrew


C. sibirica
Dukelsky, 1930 		Central Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, grassland, inland wetlands, and desert[157]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[157]

Sibuyan shrew


C. ninoyi
Esselstyn & Goodman, 2010 		Philippines
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[158]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[158]

Sicilian shrew

Brown shrew

C. sicula
Miller, 1900

Four subspecies
  • C. s. aegatensis
  • C. s. calypso
  • C. s. esuae
  • C. s. sicula
Italy and Malta
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[159]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[159]

Sinharaja white-toothed shrew


C. hikmiya
Meegaskumbura, Meegaskumbura, Pethiyagoda, Manamendra-Arachchi, & Schneider, 2007 		Sri Lanka
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[160]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[160]

Small-footed shrew


C. parvipes
Osgood, 1910 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, savanna, and shrubland[161]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[161]

Smoky white-toothed shrew


C. fumosa
Thomas, 1904 		Kenya
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[162]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[162]

Sokolov's shrew


C. sokolovi
Jenkins, Abramov, Rozhnov, & Makarova, 2007 		Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[163]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[163]

Somali dwarf shrew


C. nana
Dobson, 1890 		Ethiopia and Somalia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[164]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[164]

Somali shrew


C. somalica
Thomas, 1895 		Eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[165]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[165]

Southeast Asian shrew

Gray shrew

C. fuliginosa
(Blyth, 1856) 		Southeastern Asian
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, grassland, and rocky areas[166]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[166]

Sri Lankan long-tailed shrew


C. miya
Phillips, 1929 		Sri Lanka
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[167]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[167]

Sulawesi shrew


C. lea
Miller & Hollister, 1921 		Sulawesi island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[168]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[168]

Sulawesi tiny shrew


C. levicula
Miller & Hollister, 1921 		Sulawesi island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[169]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[169]

Sulawesi white-handed shrew


C. rhoditis
Miller & Hollister, 1921 		Sulawesi island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[170]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[170]

Sumatran giant shrew


C. lepidura
Lyon
, 1908 		Sumatra island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[171]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[171]

Sumatran long-tailed shrew


C. paradoxura
Dobson, 1886 		Sumatra island in Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[172]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population declining[172]

Sunda shrew

C. monticola
Peters, 1870 		Southeastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[173]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[173]

Swamp musk shrew


C. mariquensis
(Smith, 1844)

Three subspecies
  • C. m. mariquensis
  • C. m. neavei
  • C. m. shortridgei
Southern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[174]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[174]

São Tomé shrew

Brown shrew

C. thomensis
(Bocage, 1887) 		São Tomé Island in São Tomé and Príncipe
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[175]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[175]

Taiwanese gray shrew


C. tanakae
Kuroda, 1938 		Eastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[176]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[176]

Tanzanian shrew


C. tansaniana
Hutterer, 1986 		Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[177]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 NT 


Unknown Unknown[177]

Tarella shrew


C. tarella
Dollman, 1915 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[178]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 EN 


Unknown Population declining[178]

Telford's shrew


C. telfordi
Hutterer, 1986 		Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[179]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Population declining[179]

Thalia's shrew


C. thalia
Dippenaar, 1980 		Ethiopia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[180]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[180]

Therese's shrew


C. theresae
Heim de Balsac, 1968 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[181]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[181]

Thick-tailed shrew


C. brunnea
Jentink, 1888

Two subspecies
  • C. b. brunnea
  • C. b. pudjonica
Indonesia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[182]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[182]

Timor shrew


C. tenuis
(Müller, 1840) 		Timor
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[183]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[183]

Turbo shrew


C. turba
Dollman, 1910 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[184]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[184]

Ugandan lowland shrew


C. selina
(Dollman, 1915) 		Kenya and Uganda
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and inland wetlands[185]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[185]

Ugandan musk shrew


C. mutesae
Heller, 1910 		Central Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[186]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[186]

Ultimate shrew


C. ultima
Dollman, 1915 		Kenya
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[187]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[187]

Upemba shrew


C. zimmeri
Osgood, 1936 		Democratic Republic of the Congo
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[188]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[188]

Usambara shrew


C. usambarae
Dippenaar, 1980 		Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[189]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 VU 


Unknown Unknown[189]

Ussuri white-toothed shrew

Brown shrew

C. lasiura
(Dobson, 1890) 		Eastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[190]

Diet: Insects and other invertebrates, as well as small vertebrates[190] 		 LC 


Unknown Unknown[190]

Voi shrew


C. voi
Osgood, 1910 		Central and eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[191]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[191]

Voracious shrew

Brown shrew

C. vorax
Allen, 1923 		Southeastern Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[192]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[192]

West African long-tailed shrew


C. muricauda
(Miller, 1900) 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[193]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[193]

West African pygmy shrew


C. obscurior
Heim de Balsac, 1958 		Western Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[194]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[194]

Whitaker's shrew


C. whitakeri
De Winton, 1898 		Northwestern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Coastal marine, intertidal marine, and shrubland[195]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[195]

Wimmer's shrew


C. wimmeri
Heim de Balsac & Aellen, 1958 		Ivory Coast
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[196]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Unknown[196]

Xanthippe's shrew


C. xantippe
Osgood, 1910 		Kenya and Tanzania
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna and shrubland[197]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[197]

Yankari shrew


C. yankariensis
Hutterer & Jenkins, 1980 		Central and eastern Africa
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[198]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Population steady[198]

Zaphir's shrew


C. zaphiri
Dollman, 1915 		Ethiopia and Kenya
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[199]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 DD 


Unknown Unknown[199]

Zarudny's rock shrew

Gray shrew

C. zarudnyi
Ognew, 1928 		Central Asia
Map of range 		Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Desert[200]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6] 		 LC 


Unknown Unknown[200]

Genus Diplomesodon – Brandt, 1852 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Piebald shrew

Gray and white shrew

D. pulchellum
(Lichtenstein, 1823) 		Central Asia
Map of range 		Size: 5–8 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[201]

Habitat: Desert[202]

Diet: Insects, as well as small lizards[202] 		 LC 


Unknown Population steady[202]

Genus
Feroculus – Kelaart
, 1852 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Kelaart's long-clawed shrew


F. feroculus
(Kelaart, 1850) 		Southern India and Sri Lanka
Map of range 		Size: 10–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[203]

Habitat: Forest, grassland, and inland wetlands[204]

Diet: Earthworms, small vertebrates, and plants[203] 		 EN 


Unknown Population declining[204]

Genus Paracrocidura – Heim de Balsac, 1956 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Grauer's large-headed shrew


P. graueri
Hutterer, 1986 		Democratic Republic of the Congo
Map of range 		Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[205]

Habitat: Forest[206]

Diet: Insects[205] 		 DD 


Unknown Unknown[206]

Greater large-headed shrew


P. maxima
Heim de Balsac, 1959 		Central Africa
Map of range 		Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[205]

Habitat: Forest and inland wetlands[207]

Diet: Insects[205] 		 LC 


Unknown Population declining[207]

Lesser large-headed shrew


P. schoutedeni
Heim de Balsac, 1956 		West-central Africa
Map of range 		Size: 6–10 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[205]

Habitat: Forest[208]

Diet: Insects[205] 		 LC 


Unknown Unknown[208]

Genus
Ruwenzorisorex – Hutterer
, 1986 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Ruwenzori shrew


R. suncoides
(Osgood, 1936) 		Central Africa
Map of range 		Size: 9–10 cm (4–4 in) long, plus 5–7 cm (2–3 in) tail[209]

Habitat: Inland wetlands and forest[210]

Diet: Mollusks and small vertebrates[203] 		 VU 


Unknown Unknown[210]

Genus Scutisorex – Thomas, 1910 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Hero shrew

Black-and-white picture of a shrew

S. somereni
(Thomas, 1910) 		Central Africa
Map of range 		Size: 12–15 cm (5–6 in) long, plus 6–10 cm (2–4 in) tail[211]

Habitat: Forest[212]

Diet: Insects, earthworms, and small vertebrates[211] 		 LC 


Unknown Unknown[212]

Thor's hero shrew


S. thori
Stanley, Malekani, &Gambalemoke
, 2013 		Democratic Republic of the Congo Size: 12–15 cm (5–6 in) long, plus 6–10 cm (2–4 in) tail[213]

Habitat: Forest[214]

Diet: Small animals and plants[213] 		 DD 


Unknown Unknown[214]

Genus
Solisorex – Thomas
, 1924 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Pearson's long-clawed shrew


S. pearsoni
Thomas, 1924 		Sri Lanka
Map of range 		Size: 12–15 cm (5–6 in) long, plus 5–7 cm (2–3 in) tail[201]

Habitat: Forest and grassland[215]

Diet: Insects, earthworms, and small vertebrates[201] 		 EN 


Unknown Population declining[215]

Genus Suncus – Ehrenberg, 1832 – eighteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Anderson's shrew


S. stoliczkanus
(Anderson, 1877) 		Southern Asia
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest and grassland[217]

Diet: Insects[216] 		 LC 


Unknown Unknown[217]

Asian highland shrew


S. montanus
(Kelaart, 1850) 		Southern India and Sri Lanka
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest and grassland[218]

Diet: Insects[216] 		 VU 


Unknown Population declining[218]

Asian house shrew

Gray shrew

S. murinus
(Linnaeus, 1766) 		Asia and Africa (introduced in red)
Map of range 		Size: 10–5 cm (4–2 in) long, plus 5–9 cm (2–4 in) tail[219]

Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[220]

Diet: Insects and small mammals, as well as plants[221] 		 LC 


Unknown Population steady[220]

Black shrew


S. ater
(Medway, 1965) 		Northern Borneo
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest[222]

Diet: Insects[216] 		 DD 


Unknown Unknown[222]

Bornean pygmy shrew


S. hosei
(Thomas, 1893) 		Borneo
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest[223]

Diet: Insects[216] 		 DD 


Unknown Unknown[223]

Climbing shrew


S. megalura
(Jentink, 1888) 		Sub-Saharan Africa
Map of range 		Size: 5–6 cm (2 in) long, plus 8–10 cm (3–4 in) tail[219]

Habitat: Forest and savanna[224]

Diet: Insects[219] 		 LC 


Unknown Unknown[224]

Day's shrew


S. dayi
(Dodson, 1888) 		Southern India
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest and grassland[225]

Diet: Insects[216] 		 EN 


Unknown Population declining[225]

Etruscan shrew

Gray shrew

S. etruscus
(Savi, 1822) 		Europe, northern Africa, and Asia
Map of range 		Size: 3–5 cm (1–2 in) long, plus 2–3 cm (1 in) tail[1]

Habitat: Forest, shrubland, grassland, and desert[226]

Diet: Insects[227] 		 LC 


Unknown Unknown[226]

Flores shrew


S. mertensi
Kock, 1974 		Flores island in Indonesia
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest[228]

Diet: Insects[216] 		 EN 


Unknown Population declining[228]

Greater dwarf shrew

Gray shrew

S. lixus
(Thomas, 1898) 		Southern Africa
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[216]

Habitat: Forest, savanna, shrubland, and grassland[229]

Diet: Insects[216] 		 LC 


Unknown Unknown[229]

Hutu-Tutsi dwarf shrew


S. hututsi
Peterhans & Hutterer, 2009 		Central Africa Size: 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[216]

Habitat: Forest and shrubland[230]

Diet: Insects[216] 		 DD 


Unknown Unknown[230]

Jungle shrew


S. zeylanicus
Phillips, 1928 		Sri Lanka
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest[231]

Diet: Insects 		 EN 


Unknown Population declining[231]

Least dwarf shrew


S. infinitesimus
(Heller, 1912) 		South Africa and Kenya
Map of range 		Size: 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[216]

Habitat: Forest and grassland[232]

Diet: Insects[216] 		 LC 


Unknown Unknown[232]

Lesser dwarf shrew


S. varilla
(Thomas, 1895) 		Southern Africa
Map of range 		Size: 5–7 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[216]

Habitat: Forest, savanna, shrubland, and grassland[233]

Diet: Insects[216] 		 LC 


Unknown Unknown[233]

Malayan pygmy shrew


S. malayanus
(Kloss, 1917) 		Malaysia and Thailand
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest[234]

Diet: Insects[216] 		 LC 


Unknown Unknown[234]

Remy's pygmy shrew


S. remyi
Brosset, Dubost, & Heim de Balsac, 1965 		West-central Africa
Map of range 		Size: 4–5 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail[216]

Habitat: Forest[235]

Diet: Insects[216] 		 LC 


Unknown Unknown[235]

Sri Lankan shrew


S. fellowesgordoni
Phillips, 1932 		Sri Lanka
Map of range 		Size: 5–9 cm (2–4 in) long, plus 3–10 cm (1–4 in) tail[216]

Habitat: Forest and grassland[236]

Diet: Insects[216] 		 EN 


Unknown Population declining[236]

Taita shrew


S. aequatorius
(Heller, 1912) 		Kenya
Map of range 		Size: 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[237]

Habitat: Forest[238]

Diet: Insects[239] 		 EN 


Unknown Unknown[238]

Genus Sylvisorex – Thomas, 1904 – fifteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bamenda pygmy shrew


S. silvanorum
(Hutterer, Riegert, & Sedláček, 2009) 		Cameroon Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[209]

Habitat: Forest and grassland[240]

Diet: Insects[241] 		 DD 


Unknown Unknown[240]

Bioko forest shrew


S. isabellae
Heim de Balsac, 1968 		West-central Africa
Map of range 		Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[209]

Habitat: Forest and grassland[242]

Diet: Insects[241] 		 VU 


Unknown Population declining[242]

Cameroonian forest shrew


S. camerunensis
Heim de Balsac, 1968 		West-central Africa
Map of range 		Size: 5–6 cm (2 in) long, plus 5–7 cm (2–3 in) tail[243]

Habitat: Forest and grassland[244]

Diet: Insects[241] 		 VU 


Unknown Population declining[244]

Corbet's forest shrew


S. corbeti
Hutterer, 2009 		West-central Africa Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[209]

Habitat: Forest[245]

Diet: Insects[241] 		 DD 


Unknown Unknown[245]

Dudu Akaibe's pygmy shrew


S. akaibei
Mukinzi, Hutterer, & Barriere, 2009 		Democratic Republic of the Congo Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[209]

Habitat: Forest[246]

Diet: Insects[241] 		 DD 


Unknown Unknown[246]

Grant's forest shrew


S. granti
Thomas, 1907

Two subspecies
  • S. g. granti
  • S. g. mundus
Central Africa
Map of range 		Size: 4–7 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[247]

Habitat: Forest[248]

Diet: Insects[241] 		 LC 


Unknown Unknown[248]

Greater forest shrew


S. ollula
Thomas, 1913 		Central Africa
Map of range 		Size: 5–8 cm (2–3 in) long, plus 1–2 cm (0–1 in) tail[249]

Habitat: Forest[250]

Diet: Insects[241] 		 LC 


Unknown Unknown[250]

Howell's forest shrew


S. howelli
Jenkins, 1984 		Tanzania
Map of range 		Size: 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[251]

Habitat: Forest[252]

Diet: Insects[241] 		 LC 


Unknown Population declining[252]

Johnston's forest shrew


S. johnstoni
(Dobson, 1888) 		West-central and central Africa
Map of range 		Size: 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[243]

Habitat: Forest[253]

Diet: Insects[241] 		 LC 


Unknown Unknown[253]

Kongana shrew


S. konganensis
Ray & Hutterer, 1996 		Central Africa
Map of range 		Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[209]

Habitat: Forest[254]

Diet: Insects[241] 		 DD 


Unknown Unknown[254]

Lesser forest shrew

S. oriundus
Hollister, 1916 		Central Africa
Map of range 		Size: 4–10 cm (2–4 in) long, plus 4–9 cm (2–4 in) tail[209]

Habitat: Forest[255]

Diet: Insects[241] 		 DD 


Unknown Unknown[255]

Moon forest shrew


S. lunaris
Thomas, 1906 		Central Africa
Map of range 		Size: 7–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[256]

Habitat: Forest[257]

Diet: Insects[241] 		 NT 


Unknown Population declining[257]

Mount Cameroon forest shrew


S. morio
(Gray, 1862) 		West-central Africa
Map of range 		Size: 6–8 cm (2–3 in) long, plus 5–6 cm (2 in) tail[258]

Habitat: Forest[259]

Diet: Insects[241] 		 EN 


Unknown Population declining[259]

Rain forest shrew


S. pluvialis
Hutterer & Schlitter, 1996 		West-central Africa
Map of range 		Size: 6–8 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[256]

Habitat: Forest[260]

Diet: Insects[241] 		 DD 


Unknown Unknown[260]

Volcano shrew


S. vulcanorum
Hutterer & Verheyan, 1985 		Central Africa
Map of range 		Size: 4–6 cm (2 in) long, plus 4–7 cm (2–3 in) tail[243]

Habitat: Forest and inland wetlands[261]

Diet: Insects[241] 		 LC 


Unknown Population declining[261]

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Retrieved from "https://en.wikipedia.org/w/index.php?title=List_of_crocidurines&oldid=1220991482"