Cod

Source: Wikipedia, the free encyclopedia.
This article is about the genus of fish. For other uses, see Cod (disambiguation).

Atlantic cod

Cod (pl.: cod) is the common name for the demersal fish genus Gadus, belonging to the family Gadidae.[1] Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus Gadus is commonly not called cod (Alaska pollock, Gadus chalcogrammus).

The two most common species of cod are the

North Atlantic, and the Pacific cod (Gadus macrocephalus), found in both eastern and western regions of the northern Pacific. Gadus morhua was named by Linnaeus in 1758. (However, G. morhua callarias, a low-salinity, nonmigratory race
restricted to parts of the Baltic, was originally described as Gadus callarias by Linnaeus.)

Cod as food is popular in several countries. It has a mild flavour and a dense, flaky, white flesh. Cod livers are processed to make cod liver oil, a common source of vitamin A, vitamin D, vitamin E, and omega-3 fatty acids (EPA and DHA). Young Atlantic cod or haddock prepared in strips for cooking is called scrod. In the United Kingdom, Atlantic cod is one of the most common ingredients in fish and chips, along with haddock and plaice.

Species

At various times in the past, taxonomists included many species in the genus Gadus. Most of these are now either classified in other genera, or have been recognized as forms of one of three species. All these species have a number of common names, most of them ending with the word "cod", whereas other species, as closely related, have other common names (such as pollock and haddock). However, many other, unrelated species also have common names ending with cod. The usage often changes with different localities and at different times.

Cod in the genus Gadus

Three species in the genus Gadus are currently called cod:

This article is part of a series on
Commercial fish
Large pelagic
Forage
Demersal
Mixed
Cod in the genus Gadus
Common name Scientific name Maximum
length 		Common
length 		Maximum
weight 		Maximum
age 		Trophic
level 		Fish
Base
FAO
ITIS
IUCN status
Atlantic cod Gadus morhua Linnaeus, 1758 200 cm 100 cm 96.0 kg 25 years 4.4 [2] [3] [4] VU IUCN 3 1.svg Vulnerable[5]
Pacific cod Gadus macrocephalus Tilesius, 1810 119 cm cm 22.7 kg 18 years 4.0 [6] [7] [8] Not assessed
Greenland cod Gadus ogac Richardson, 1836 77.0 cm cm kg 12 years 3.6 [9] [10] [11] Not assessed

The fourth species of genus Gadus,

Gadus chalcogrammus, is commonly called Alaska pollock or walleye pollock. But there are also less widespread alternative trade names highlighting the fish's belonging to the cod genus, like snow cod[12][13][14] or bigeye cod.[13]

Related species

Cod forms part of the

Ranicipitidae
) has now been placed in Gadidae.

"Cod" in the order Gadiformes, but not part of Gadus
Common name Scientific name Maximum
length 		Common
length 		Maximum
weight 		Maximum
age 		Trophic
level 		Fish
Base
FAO
ITIS
IUCN status
Arctic cod
 Arctogadus glacialis (Peters, 1872) 32.5 cm cm kg years 3.8 [15] [16] Not assessed
East Siberian cod Arctogadus borisovi Dryagin, 1932 55.6 cm cm 1.5 kg years 3.9 [17] [18] Not assessed
Eucla cod Euclichthys polynemus McCulloch, 1926 35.0 cm 22.5 cm kg years 3.6 [19] [20] Not assessed
Common ling Molva molva (Linnaeus, 1758) 200 cm 106 cm 45.0 kg 25 years 4.3 [21] [22] [23] Not assessed
Pelagic cod Melanonus gracilis Günther, 1878 18.7 cm cm kg years 3.5 [24] [25] Not assessed
Polar cod
 Boreogadus saida (Lepechin, 1774) 40.0 cm 25.0 cm kg 7 years 3.1 [26] [27] [28] Not assessed
Poor cod Trisopterus minutus (Linnaeus, 1758) 40.0 cm 20.0 cm kg 5 years 3.8 [29] [30] Not assessed
Rock cod Lotella rhacina (Forster, 1801) 50.0 cm cm kg years 3.5 [31] [32] Not assessed
Saffron cod Eleginus gracilis (Tilesius, 1810) 55.0 cm cm 1.3 kg 15 years 4.1 [33] [34] [35] Not assessed
Small-headed cod Lepidion microcephalus Cowper, 1956 48.0 cm cm kg years 3.5 [36] [37] Not assessed
Tadpole cod
 Guttigadus globosus (Paulin, 1986) 18.1 cm cm kg 3.5 years [38] [39] Not assessed

Some fish have common names derived from "cod", such as

tomcod
. ("Codling" is also used as a name for a young cod.)

Other species

Some fish commonly known as cod are unrelated to Gadus. Part of this name confusion is market-driven. Severely shrunken Atlantic cod stocks have led to the marketing of cod replacements using culinary names of the form "x cod", according to culinary rather than phyletic similarity. The common names for the following species have become well established; note that all inhabit the Southern Hemisphere.

Perciformes

Fish of the order Perciformes that are commonly called "cod" include:

Rock cod, reef cod, and coral cod

Almost all

Lotella rhacina
, which as noted above actually is related to the true cod (it is a morid cod).

Scorpaeniformes

From the order Scorpaeniformes:

  • Ling cod
    Ophiodon elongatus
  • Red rock cod
    Scorpaena papillosa
  • Rock cod Sebastes

Ophidiiformes

The tadpole cod family,

Euclichthyidae, were formerly classified in the order Ophidiiformes, but are now grouped with the Gadiformes
.

Marketed as cod

Some fish that do not have "cod" in their names are sometimes sold as cod. Haddock and whiting belong to the same family, the Gadidae, as cod.

Characteristics

The Atlantic cod, Gadus morhua

Cods of the genus Gadus have three rounded

caudal
or tail fin. The back tends to be a greenish to sandy brown, and shows extensive mottling, especially towards the lighter sides and white belly. Dark brown colouration of the back and sides is not uncommon, especially for individuals that have resided in rocky inshore regions.

The Atlantic cod can change colour at certain water depths. It has two distinct colour phases: gray-green and reddish brown. Its average weight is 5–12 kilograms (11–26 pounds), but specimens weighing up to 100 kg (220 lb) have been recorded. Pacific cod are smaller than Atlantic cod[2][6] and are darker in colour.

Distribution

Atlantic cod (Gadus morhua) live in the colder waters and deeper sea regions throughout the North Atlantic. Pacific cod (Gadus macrocephalus) is found in both eastern and western regions of the Pacific.[41]

Atlantic cod could be further divided into several

West Greenland, Newfoundland, and Labrador stocks. There seems to be little interchange between the stocks, although migrations to their individual breeding grounds may involve distances of 300 kilometres (190 statute miles; 160 nautical miles) or more.[42] For instance, eastern Baltic cod shows specific reproductive adaptations to low salinity compared to Western Baltic and Atlantic cod.[43]

Atlantic cod occupy varied habitats, favouring rough ground, especially inshore, and are

demersal in depths between 6 and 60 metres (20 and 200 feet; 3 and 30 fathoms), 80 m (260 ft; 44 fathoms) on average, although not uncommonly to depths of 600 m (2,000 ft; 330 fathoms). Off the Norwegian and New England coasts and on the Grand Banks of Newfoundland, cod congregate at certain seasons in water of 30–70 m (100–200 ft; 20–40 fathoms) depth. Cod are gregarious and form schools, although shoaling
tends to be a feature of the spawning season.

Life cycle

Spawning of northeastern Atlantic cod occurs between January and April (March and April are the peak months), at a depth of 200 metres (660 ft) in specific spawning grounds at water temperatures between 4 and 6 °C (39 and 43 °F). Around the UK, the major spawning grounds are in the middle to southern North Sea, the start of the Bristol Channel (north of Newquay), the Irish Channel (both east and west of the Isle of Man), around Stornoway, and east of Helmsdale.

Prespawning courtship involves fin displays and male grunting, which leads to pairing.

isopods and small crabs. They increase in size to 8 cm (3 in) in the first six months, 14–18 cm (5+1⁄2–7 in) by the end of their first year, and to 25–35 cm (10–14 in) by the end of the second. Growth tends to be less at higher latitudes. Cod reach maturity at about 50 cm (20 in) at about 3 to 4 years of age. Changes in growth rate over decades of particular stocks have been reported, current eastern Baltic cod shows the lowest growth observed since 1955.[45]

Ecology

See also:
Diseases and parasites in cod
cod worms

Adult cod are active hunters, feeding on

molluscs
.

In the Baltic Sea the most important prey species are

sprat.[46] Many studies that analyze the stomach contents of these fish indicate that cod is the top predator, preying on the herring and sprat.[46] Sprat form particularly high concentrations in the Bornholm Basin in the southern Baltic Sea.[47] Although cod feed primarily on adult sprat, sprat tend to prey on the cod eggs and larvae.[48]

Cod and related species are plagued by parasites. For example, the

lumpsucker, which it captures with grasping hooks at the front of its body. It penetrates the fish with a thin filament, which it uses to suck the fish's blood. The nourished larvae then mate on the fish.[49][50] The female larva, with her now fertilized eggs, then finds a cod, or a cod-like fish such as a haddock or whiting. There the larva clings to the gills while it metamorphoses into a plump sinusoidal wormlike body with a coiled mass of egg strings at the rear. The front part of the worm's body penetrates the body of the cod until it enters the rear bulb of the host's heart. There, firmly rooted in the cod's circulatory system, the front part of the parasite develops like the branches of a tree, reaching into the main artery. In this way, the worm extracts nutrients from the cod's blood, remaining safely tucked beneath the cod's gill cover until it releases a new generation of offspring into the water.[49][50]

Fisheries

FAO 1950–2010[51]
The same chart as above, but showing embedded in light green, the collapse of the Atlantic northwest cod fishery[52]
Main article: Cod fisheries

The 2006 northwest Atlantic cod

total allowable catch (TAC) for the Gulf of Alaska and Aleutian Islands was 260,000 tons.[53]

Aquaculture

Farming of Atlantic cod has received a significant amount of interest due to the overall trend of increasing cod prices alongside reduced wild catches.[54] However, progress in creating large scale farming of cod has been slow, mainly due to bottlenecks in the larval production stage, where survival and growth are often unpredictable.[55] It has been suggested that this bottleneck may be overcome by ensuring cod larvae are fed diets with similar nutritional content as the copepods they feed on in the wild [56][57] Recent examples have shown that increasing dietary levels of minerals such as selenium, iodine and zinc may improve survival and/or biomarkers for health in aquaculture reared cod larvae.[58][59][60][61]

As food

Preserved codfish
Main article: Cod as food

Cod is popular as a food with a mild flavour and a dense, flaky white flesh. Cod livers are processed to make cod liver oil, an important source of vitamin A, vitamin D, vitamin E and omega-3 fatty acids (EPA and DHA).

Young Atlantic cod or haddock prepared in strips for cooking is called scrod. In the United Kingdom, Atlantic cod is one of the most common ingredients in fish and chips, along with haddock and plaice. Cod's soft liver can be tinned (canned) and eaten.

History

Sixteenth-century Flemish fishmonger displaying cod, by Joachim Beuckelaer

Cod has been an important economic commodity in

Sacred Cod of Massachusetts
, in its chambers.

Apart from the long history, cod differ from most fish because the fishing grounds are far from population centres. The large cod fisheries along the coast of

Bergen as the most important port.[65]

William Pitt the Elder, criticizing the Treaty of Paris in Parliament, claimed cod was "British gold"; and that it was folly to restore Newfoundland fishing rights to the French
.

In the 17th and 18th centuries in the New World, especially in Massachusetts and Newfoundland, cod became a major commodity, creating trade networks and cross-cultural exchanges. In 1733, Britain tried to gain control over trade between New England and the British Caribbean by imposing the Molasses Act, which they believed would eliminate the trade by making it unprofitable. The cod trade grew instead, because the "French were eager to work with the New Englanders in a lucrative contraband arrangement".[63] In addition to increasing trade, the New England settlers organized into a "codfish aristocracy". The colonists rose up against Britain's "tariff on an import".

In the 20th century, Iceland re-emerged as a fishing power and entered the Cod Wars. In the late 20th and early 21st centuries, fishing off the European and American coasts severely depleted stocks and become a major political issue. The necessity of restricting catches to allow stocks to recover upset the fishing industry and politicians who are reluctant to hurt employment.

Collapse of the Atlantic northwest cod fishery

Main article: Collapse of the Atlantic northwest cod fishery
The Atlantic fishery abruptly collapsed in 1992, following overfishing since the late 1950s, and an earlier partial collapse in the 1970s.[66][67]

On July 2, 1992, the Honourable

Federal Minister of Fisheries and Oceans, declared a two-year moratorium on the Northern Cod fishery,[68] a designated fishing region off the coast of Newfoundland, after data showed that the total cod biomass had suffered a collapse to less than 1% of its normal value.[69] The minister championed the measure as a temporary solution, allowing the cod population time to recover.[70] The fisheries had long shaped the lives and communities on Canada's Atlantic eastern coast for the preceding five centuries. Societies which are dependent on fishing have a strong mutual relationship with them: the act of fishing changes the ecosystems' balance, which forces the fishery and, in turn, the fishing societies to adapt to new ecological conditions.[69]

The near-complete destruction of the Atlantic northwest cod biomass off the shores devastated coastal communities, which had been overexploiting the same cod population for decades.[70] The fishermen along the Atlantic northwest had employed modern fishing technologies, including the ecologically-devastating practice of trawling, especially in the years leading up to the 1990s, in the misguided belief that fishing stocks are perpetually plentiful and unable to be depleted.[70][71][68] After this assumption was empirically and abruptly shown to be incorrect, to the dismay of government officials and rural workers, some 19,000 fishermen and cod processing plant workers in Newfoundland lost their employment.[70] The powerful economic engine of rural Newfoundland coughed, wheezed, and died. Nearly 40,000 workers and harvesters in the provinces of Newfoundland and Labrador applied for the federal relief program TAGS (the Atlantic Groundfish Strategy). Abandoned and rusting fishing boats still litter the coasts of Newfoundland and the Canadian northwest to this day.[68]

The fishery minister, John Crosbie, after delivering a speech on the day before the declaration of the moratorium, or July 1, 1992, was publicly heckled and verbally harassed by disgruntled locals at a fishing village.[72] The moratorium, initially lasting for only two years,[70] was indefinitely extended after it became evident that cod populations had not recovered at all but, instead, had continued to spiral downward in both size and numbers, due to the damage caused by decades of horrible fishing practices, and the fact that the moratorium had permitted exceptions for food fisheries for "personal consumption" purposes to this very day.[70] Some 12,000 tons of Northwest cod are still being caught every year along the Newfoundland coast by local fishermen.[68]

The collapse of the four-million ton biomass, which had persevered through several previous marine extinctions over tens of millions of years, in a timespan of no more than 20 years, is oft-cited by researchers as one of the most visible examples of the phenomenon of the "Tragedy of the Commons."[70] Factors which had been implicated as contributing to the collapse include: overfishing; government mismanagement; the disregard of scientific uncertainty;[70] warming habitat waters; declining reproduction; and plain human ignorance.[68] The Northern Cod biomass has been recovering slowly since the imposition of the moratorium. However, as of 2021, the growth of the cod population has been stagnant since 2017, and some scientists argue that the population will not rebound unless the Fisheries Department of Canada lower its yearly quota to 5,000 tons.[73]

See also

References

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Further reading

External videos
video icon Presentation by Mark Kurlansky on Cod: A Biography of the Fish That Changed the World, August 15, 1998, C-SPAN
  • Bavington, Dean L.Y. Managed Annihilation: An Unnatural History of the Newfoundland Cod Collapse (University of British Columbia Press; 2010) 224 pages. Links the collapse of Newfoundland and Labrador cod fishing to state management of the resource.
  • Cobb, John N (1916). Pacific Cod Fisheries. Bureau of Fisheries Document. Vol. 830. Washington, DC: Government Printing Office.
    OCLC 14263968
    .
  • Greenberg, Paul (2010). Four Fish: The Future of the Last Wild Food. New York: Penguin Press.
    OCLC 813929026
    .
  • Mark Kurlansky (1997). Cod: A Biography of the Fish That Changed the World.
  • Shields, Edward (2001). Salt of the sea: the Pacific Coast cod fishery and the last days of sail. Lopez Island, Wash.: Heritage House.
    ISBN 978-1-894384-35-3
    .

External links

Look up cod in Wiktionary, the free dictionary.
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Wikispecies has information related to Gadus.
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