Seashell

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Part of a series on
Seashells
Mollusc shells
Clam shells Chiton shells Snail shells Tusk shells
About mollusc shells
Conchology Nacre Valve
Other seashells
Brachiopod shells Crustacean shells Horseshoe crabs Sea urchin tests
v t e

A seashell or sea shell, also known simply as a shell, is a hard, protective outer layer usually created by an animal or organism that lives in the sea. The shell is part of the body of the animal. Empty seashells are often found washed up on beaches by beachcombers. The shells are empty because the animal has died and the soft parts have decomposed or been eaten by another animal.

A seashell is usually the

Apart from

Seashells have been used by humans for many different purposes throughout history and prehistory. However, seashells are not the only kind of shells; in various habitats, there are shells from freshwater animals such as

When the word "seashells" refers only to the shells of marine mollusks, then studying seashells is part of conchology. Conchologists or serious collectors who have a scientific bias are in general careful not to disturb living populations and habitats: even though they may collect a few live animals, most responsible collectors do not often over-collect or otherwise disturb ecosystems.

The study of the entire molluscan animal (as well as the shell) is known as

Occurrence

Seashells are commonly found in beach drift, which is natural detritus deposited along strandlines on beaches by the waves and the tides. Shells are very often washed up onto a beach empty and clean, the animal having already died.

Empty seashells are often picked up by beachcombers. However, the majority of seashells which are offered for sale commercially have been collected alive (often in bulk) and then killed and cleaned, specifically for the commercial trade.^[1] This type of large-scale exploitation can sometimes have a strong negative impact on local ecosystems, and sometimes can significantly reduce the distribution of rare species.

Shell synthesis

Seashells are created by the molluscs that use them for protection.^[2] Molluscs have an outside layer of tissues on their bodies – the mantle – which creates the shell material and which connects the shell to the mollusc. The specialized cells in the mantle form the shell using different minerals and proteins.^[2] The proteins are then used to create the framework that supports the growing shell. Calcium carbonate is the main compound of shell structure, aiding in adhesion.^[2]

Molluscan seashells

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The word seashell is often used to mean only the shell of a

Marine species of gastropods and bivalves are more numerous than land and freshwater species, and the shells are often larger and more robust. The shells of marine species also often have more sculpture and more color, although this is by no means always the case.

In the

Although there are a number of species of shelled mollusks that are quite large, there are vast numbers of extremely small species too, see micromollusks.

Not all mollusks are marine. There are numerous land and freshwater mollusks, see for example

Bivalves are often the most common seashells that wash up on large sandy beaches or in sheltered lagoons. They can sometimes be extremely numerous. Very often the two valves become separated.

There are more than 15,000 species of bivalves that live in both marine and freshwater. Examples of bivalves are clams, scallops, mussels, and oysters. The majority of bivalves consist of two identical shells that are held together by a flexible hinge. The animal's body is held protectively inside these two shells. Bivalves that do not have two shells either have one shell or they lack a shell altogether. The shells are made of calcium carbonate and are formed in layers by secretions from the mantle. Bivalves, also known as pelecypods, are mostly filter feeders; through their gills, they draw in water, in which is trapped tiny food particles. Some bivalves have eyes and an open circulatory system. Bivalves are used all over the world as food and as a source of pearls. The larvae of some freshwater mussels can be dangerous to fish and can bore through wood.

Certain species of gastropod seashells (the shells of sea snails) can sometimes be common, washed up on sandy beaches, and also on beaches that are surrounded by rocky marine habitat.

Polyplacophorans

Chiton plates or valves often wash up on beaches in rocky areas where chitons are common. Chiton shells, which are composed of eight separate plates and a girdle, usually come apart not long after death, so they are almost always found as disarticulated plates. Plates from larger species of chitons are sometimes known as "butterfly shells" because of their shape.

Cephalopods

Only a few species of cephalopods have shells (either internal or external) that are sometimes found washed up on beaches.

Some cephalopods such as

, and this often washes up on beaches in parts of the world where cuttlefish are common.

Spirula spirula

is a deep water squid-like cephalopod. It has an internal shell which is small (about 1 in or 24 mm) but very light and buoyant. This chambered shell floats very well and therefore washes up easily and is familiar to beachcombers in the tropics.

Nautilus is the only genus of cephalopod that has a well-developed external shell. Females of the cephalopod genus Argonauta create a papery egg case which sometimes washes up on tropical beaches and is referred to as a "paper nautilus".

The largest group of shelled cephalopods, the

ammonites, are extinct, but their shells are very common in certain areas as fossils

.

Molluscan seashells used by other animals

Empty molluscan seashells are a sturdy, and usually readily available, "free" resource which is often easily found on beaches, in the

second-hand by animals other than humans for various purposes, including for protection (as in hermit crabs

) and for construction.

Mollusks

• Carrier shells in the family Xenophoridae are marine shelled gastropods, fairly large sea snails. Most species of xenophorids cement a series of objects to the rim of their shells as they grow. These objects are sometimes small pebbles or other hard detritus. Very often shells of bivalves or smaller gastropods are used, depending on what is available on the particular substrate where the snail itself lives. It is not clear whether these shell attachments serve as camouflage, or whether they are intended to help prevent the shell sinking into a soft substrate.

• Small octopuses sometimes use an empty shell as a sort of cave to hide in, or hold seashells around themselves as a form of protection like a temporary fortress.

Invertebrates

Part of a series related to
Biomineralization
General Mineralized tissues Remineralisation Biocrystallization Biointerface Biofilm
Exoskeletons (shells) Arthropod exoskeleton cuticle Brachiopod shell Cephalopod shell cirrate shell cuttlebone gladius Lorica Choanoflagellate lorica Protist shell coccosphere coccolith diatom frustule foraminifera test testate amoebae Seashell echinoderm stereom mollusc shell nacre chiton shell gastropod shell small shelly fauna scaly-foot snail shell estuary shells Sponge spicule Test
Endoskeletons (bones) Vertebrate skeleton Bone mineral Ossification
Teeth, scales, tusks etc Limpet teeth Otolith otolithic membrane Scale microfossils Tusk
Calcification amorphous calcium carbonate marine biogenic calcification calcareous nannofossils Aragonite oolitic aragonite sand aragonite sea Calcite microbial calcite precipitation calcite sea Great Calcite Belt
Silicification biogenic silica siliceous ooze diatomaceous earth
Other forms Bone bed Kerogen alginite oil shale Phosphate phosphorite Pyrena
Related Mineral evolution In soil mineralization immobilization Ballast minerals Magnetofossil Magnetosome Magnetotactic bacteria Magnetoreception Microfossils engrailed gene Druse Cupriavidus metallidurans Biomineralising polychaetes Mineral nutrients Microbial mat Fossilization permineralization petrifaction Burgess Shale preservation
Category
v t e

• Almost all genera of hermit crabs use or "wear" empty marine gastropod shells throughout their lifespan, in order to protect their soft abdomens, and in order to have a strong shell to withdraw into if attacked by a predator. Each individual hermit crab is forced to find another gastropod shell on a regular basis, whenever it grows too large for the one it is currently using.

Some hermit crab species live on land and may be found quite some distance from the sea, including those in the tropical genus Coenobita.

Conchology

There are numerous popular books and

There are a number of clubs or societies which consist of people who are united by a shared interest in shells. In the US, these clubs are more common in southerly coastal areas, such as Florida and California, where the marine fauna is rich in species.

Identification

Seashells are usually identified by consulting general or regional shell-collecting field guides, and specific scientific books on different taxa of shell-bearing mollusks (monographs) or "iconographies" (limited text – mainly photographs or other illustrations). (For a few titles on this subject in the US, see the list of books at the foot of this article.) Identifications to the species level are generally achieved by examining illustrations and written descriptions, rather than by the use of Identification keys, as is often the case in identifying plants and other phyla of invertebrates. The construction of functional keys for the identification of the shells of marine mollusks to the species level can be very difficult, because of the great variability within many species and families. The identification of certain individual species is often very difficult, even for a specialist in that particular family. Some species cannot be differentiated on the basis of shell character alone.

Numerous smaller and more obscure mollusk species (see micromollusk) are yet to be discovered and named. In other words, they have not yet been differentiated from similar species and assigned scientific (binomial) names in articles in journals recognized by the International Commission on Zoological Nomenclature (ICZN). Large numbers of new species are published in the scientific literature each year. There are currently an estimated 100,000 species of mollusks worldwide.

Non-marine "seashells"

The term seashell is also applied loosely to mollusk shells that are not of marine origin, for example by people walking the shores of lakes and rivers using the term for the

Cultural significance

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Currency

Seashells have been used as a medium of exchange in various places, including many Indian Ocean and Pacific Ocean islands, also in North America, Africa and the Caribbean.

• The most common species of shells to be used as currency have been
  dentalium tusk shells
  , used in North Western North America for many centuries.
• Many of the tribes and nations all across the continent of Africa have historically used the cowry as their media of exchange. The cowry circulated, historically, alongside metal coins and goods, and foreign currencies. Being durable and easy to carry the cowry made a very favorable currency.
• Some tribes of the
  quahog
  clam.

Tools

Seashells have often been used as tools, because of their strength and the variety of their shapes.

• Giant clams (Family Tridacnidae) have been used as bowls, and when big enough, even as bathtubs and baptismal fonts.
• Melo melo, the "bailer volute", is so named because Native Australians used it to bail out their canoes.
• Many different species of bivalves have been used as scrapers, blades, clasps, and other such tools, due to their shape.
• Some marine gastropods have been used for oil lamps, the oil being poured in the aperture of the shell, and the siphonal canal serving as a holder for the wick.

Horticulture

Because seashells are in some areas a readily available bulk source of calcium carbonate, shells such as oyster shells are sometimes used as soil conditioners in horticulture. The shells are broken or ground into small pieces in order to have the desired effect of raising the pH and increasing the calcium content in the soil.

Religion and spirituality

Seashells have played a part in religion and spirituality, sometimes even as ritual objects.

• In Christianity, the
  Saint James the Great, see Pecten jacobaeus.^[6]
• In Hinduism, left-handed shells of Turbinella pyrum (the sacred shankha) are considered to be sacred to the god Vishnu. The person who finds a left-handed chank shell (one that coils to the left) is sacred to Vishnu, as well. The chank shell also plays an important role in Buddhism.
• Cowries have often been considered to be symbols of female fertility. They were often treated as actual fertility charms. The dorsum of the shell resembles a pregnant belly, and the underside of the shell resembles a vulva
  . In the South Indian state of Kerala, cowries are used for making astrological predictions.
• In the Santería religion, shells are used for divination.
• The
  Moche culture of ancient Peru worshipped animals and the sea, and often depicted shells in their art.^[7]
• In Christianity, the top of the sand dollar represents the Star of Bethlehem that led the Wise Men to the manger of Christ. Outside the "star" you will see the Easter Lily, a sign of Jesus' Resurrection. There are four holes that represent the holes in the Lord's hands and feet. The center hole is the Wound to His Sacred Heart by the spear of Longinus. On the other side of the sand dollar, you will see Poinsettia. Lastly, if you break open the sand dollar, five doves will come out, the doves of Peace and Joy.^[8]

Musical instruments

Main article:

Conch (musical instrument)

Seashells have been used as musical instruments,

wind instruments for many hundreds if not thousands of years. Most often the shells of large sea snails are used, as trumpets, by cutting a hole in the spire of the shell or cutting off the tip of the spire altogether. Various different kinds of large marine gastropod shells can be turned into "blowing shells"; however, the most commonly encountered species used as "conch

" trumpets are:

• The sacred chank, Turbinella pyrum, known in India as the shankha. In Tibet it is known as "dung-dkar".^[9]
• The
  Triton shell also known as "Triton's trumpet" Charonia tritonis which is used as a trumpet in Melanesian and Polynesian culture and also in Korea and Japan. In Japan this kind of trumpet is known as the horagai. In Korea it is known as the nagak. In some Polynesian islands it is known as "pu".^[9]
• The
  Lobatus gigas
  , has been used as a trumpet in the Caribbean.

Children in some cultures are often told the myth that you can hear the sound of the ocean by holding a seashell to ones ear. This is due to the effect of seashell resonance.

Personal adornment

Whole seashells or parts of sea shells have been used as jewelry or in other forms of adornment since prehistoric times.

• Shell necklaces have been found in Stone Age graves as far inland as the Dordogne Valley in France.
• Seashells are often used whole and drilled, so that they can be threaded like
  Palawa women elders. The shells used include pearly green and blue-green maireener (rainbow kelp) shells, brown and white rice shells, black cats' teeth shells and pink button shells.^[10]
• Naturally-occurring, beachworn,
  cone shell "tops" (the broken-off spire of the shell, which often has a hole worn at the tip) can function as beads without any further modification. In Hawaii these natural beads were traditionally collected from the beach drift in order to make puka shell jewelry. Since it is hard to obtain large quantities of naturally-occurring beachworn cone tops, almost all modern puka shell jewelry
  uses cheaper imitations, cut from thin shells of other species of mollusk, or even made of plastic.
• Shells historically have been and still are made into, or incorporated into, necklaces, pendants, beads, earrings,
  rings, hair combs
  , belt buckles and other uses.
• The shell of the large "bullmouth helmet" sea snail, scientific name Cypraecassis rufa, was historically, and still is, used to make valuable cameos.
• Haliotidae
  , and various pearly bivalves, has often been used in jewelry, buttons, etc.
• In London,
  mother-of-pearl or nacre
  . In recent years however, the majority of "pearl buttons" are imitations that are made of pearlescent plastic.

The making of shell work artifacts is a practice of Aboriginal women from La Perouse in Sydney, dating back to the 19th century. Shell work objects include baby shoes, jewelry boxes and replicas of famous landmarks, including the Sydney Harbour Bridge and the Sydney Opera House. The shellwork tradition began as an Aboriginal women's craft which was adapted and tailored to suit the tourist souvenir market, and which is now considered high art.^[11]

Architectural decoration

Small pieces of colored and iridescent shell have been used to create mosaics and inlays, which have been used to decorate walls, furniture and boxes. Large numbers of whole seashells, arranged to form patterns, have been used to decorate mirror frames, furniture and human-made shell grottos.

Art

A very large outdoor sculpture at

• 
  Enormous seashell sculpture at

  Akkulam, Thiruvananthapuram

  , India
• 
  Large sculpture of a scallop on the beach at Aldeburgh, by Maggi Hambling, 2003
• 
  Illustration from an 18th-century book, edited by Albertus Seba. These decorative arrangements were a popular way to display seashells at the time
• 
  Portrait of the Shell Collector Jan Govertsen van der Aer, by Hendrick Goltzius (1603)

Poultry feeds

Sea shells found in the creek and backwater of the coast of west India are used as an additive to poultry feed. They are crushed and mixed with jowar maize and dry fish.^{[citation needed]}

Use

Seashells, namely from bivalves^[12] and gastropods, are fundamentally composed of calcium carbonate. In this sense, they have potential to be used as raw material in the production of lime.

Along the Gulf Coast of the United States, oyster shells were mixed into cement to make "shellcrete" which could form bricks, blocks and platforms. It could also be applied over logs.^[13] A notable example is the 19th-century Sabine Pass Lighthouse in Louisiana, near Texas.^[14]

Shells of other marine invertebrates

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Arthropods

Many

Some

The

Pieces of the hard skeleton of corals commonly wash up on beaches in areas where corals grow.

The construction of the shell-like structures of corals are aided by a

Plant-like

• Bailey-Matthews Shell Museum
• Marine biogenic calcification
• Mollusk shell
• Ocean acidification
• Seashell resonance
• Seashell surface, a mathematical construct
• Shell growth in estuaries
• Shell purse
• Small shelly fauna

References

Citations

Sources

Books

• Abbott R. Tucker & S. Peter Dance, 1982, Compendium of Seashells, A full color guide to more than 4,200 of the World's Marine shells, E.P. Dutton, Inc, New York,
  ISBN 0-525-93269-0
  .
• Abbott R. Tucker, 1985, Seashells of the World: a guide to the better-known species, 1985, Golden Press, New York,
  ISBN 0-307-24410-5
  .
• Abbott, R. Tucker, 1986, Seashells of North America, St. Martin's Press, New York,
  ISBN 1-58238-125-9
  .
• Abbott, R. Tucker, 1974, American Seashells, Second edition, Van Nostrand Rheinhold, New York,
  ISBN 0-442-20228-8
  .

External links

Look up seashell in Wiktionary, the free dictionary.

Wikimedia Commons has media related to Seashells.

Wikiquote has quotations related to Seashell.

• Hohlman Shell Collection, Florida Institute of Technology