List of giant squid specimens and sightings

Source: Wikipedia, the free encyclopedia.

Newfoundland (#30 on this list), draped over Reverend Moses Harvey's sponge bath, November or December 1873. Harvey wrote in his journal: "I knew that I had in my possession what all the savants in the world did not […] what the museums in the world did not contain […] A photograph could not lie and would silence the gainsayers".[1] The photograph includes contemporaneous annotations by zoologist Addison Emery Verrill, including a 1-foot scale bar (top left) and detailed marginal notes.[nb 1]

This list of giant squid specimens and sightings is a comprehensive timeline of recorded human encounters with members of the genus Architeuthis, popularly known as giant squid. It includes animals that were caught by fishermen, found washed ashore, recovered (in whole or in part) from sperm whales and other predatory species, as well as those reliably sighted at sea. The list also covers specimens incorrectly assigned to the genus Architeuthis in original descriptions or later publications.

Background

Twenty Thousand Leagues Under the Sea.[3]

History of discovery

Tales of giant squid have been common among mariners since ancient times, but the animals were long considered

The 19-foot (5.8 m) tentacle that Newfoundland fisherman Theophilus Picot hacked off a live animal (#29) on 26 October 1873[21]

The giant squid's existence was established beyond doubt only in the 1870s, with the appearance of an extraordinary number of complete specimens—both dead and alive—in Newfoundland waters (beginning with #21).[22] These were meticulously documented in a series of papers by Yale zoologist Addison Emery Verrill.[23][nb 3] Two of these Newfoundland specimens, both from 1873, were particularly significant as they were among the earliest to be photographed: first a single severed tentacle—hacked off a live animal as it "attacked" a fishing boat (#29)[24]—and weeks later an intact animal in two parts (#30).[nb 4] The head and limbs of this latter specimen were famously shown draped over the sponge bath of Moses Harvey, a local clergyman, essayist, and amateur naturalist.[29] Harvey secured and reported widely on both of these important specimens—as well as numerous others (most notably the Catalina specimen of 1877; #42)—and it was largely through his efforts that giant squid became known to North American and British zoologists.[30][nb 5] Recognition of Architeuthis as a real animal led to the reappraisal of earlier reports of gigantic tentacled sea creatures, with some of these subsequently being accepted as records of giant squid, the earliest stretching back to at least the 17th century.[32]

"I confess that until I saw and measured this enormous limb, I doubted the accuracy of some early observations which this specimen alone would suffice to prove worthy of confidence. The existence of gigantic cephalopods is no longer an open question. I, now, more than ever, appreciate the value of the adage: 'Truth is stranger than fiction.'"

British Museum (Natural History) that was examined by him in May 1873, from the concluding lines of his 1875 book The Octopus; or, the "devil-fish" of fiction and of fact.[33]

For a time in the late 19th century, almost every major specimen of which material was saved was described as a new species.[34] In all, some twenty species names were coined.[35] However, there is no widely agreed basis for distinguishing between the named species, and both morphological and genetic data point to the existence of a single, globally distributed species, which according to the principle of priority must be known by the earliest available name: Architeuthis dux.[36]

It is not known why giant squid become stranded on shore, but it may be because the distribution of deep, cold water where they live is temporarily altered. Marine biologist and Architeuthis specialist

#563).[43]

Though the total number of recorded giant squid specimens now runs

manned submersible—in July 2012.[47] Despite these recent advances and the growing number of both specimens and recordings of live animals, the species continues to occupy a unique place in the public imagination.[48] As Roper et al. (2015:83)
wrote: "Few events in the natural world stimulate more excitement and curiosity among scientists and laymen alike than the discovery of a specimen of Architeuthis."

Distribution patterns

).

The genus Architeuthis has a

South Atlantic off South Africa and Namibia; the northwestern Pacific off Japan (especially more recently[53]); and the southwestern Pacific around New Zealand[54] and Australia.[55]

The vast majority of specimens are of oceanic origin, including

550), but these records do not necessarily indicate that the Mediterranean falls within the natural range of the giant squid, as the specimens may have been transported there by inflowing Atlantic water.[42] Similarly, giant squid are unlikely to naturally occur in the North Sea owing to its shallow depth[42] (but see #107 and 113, the only known English strandings). They are generally absent from equatorial and high polar latitudes[57] (but see #215 and 249 from equatorial Atlantic waters, and specimens from northern Norway[58] or #102 from the edge of the Arctic Circle off western Greenland
).

Total number of specimens

According to Guerra et al. (2006), 592 confirmed giant squid specimens were known as of the end of 2004. Of these, 306 came from the Atlantic Ocean, 264 from the Pacific Ocean, 20 from the Indian Ocean, and 2 from the Mediterranean Sea. The figures for specimens collected in the Atlantic and Pacific Oceans further broke down as follows: 148 in the northeastern Atlantic, 126 in the northwestern Atlantic, 26 in the southeastern Atlantic, 6 in the southwestern Atlantic, 43 in the northeastern Pacific, 28 in the northwestern Pacific, 10 in the southeastern Pacific, and 183 in the southwestern Pacific.[59]

#563).[53] The giant squid nevertheless remains a rarely encountered animal, especially considering its wide distribution and large size,[60] with Richard Ellis writing that "each giant squid that washes up or is taken from the stomach of a sperm whale is still an occasion for a teuthological celebration".[61]

Giant squid at the surface with an approaching ship in the background, from a painting by Herbert B. Judy, 1905. Specimens found stranded or floating at the surface constitute almost 50% of all records from the Atlantic Ocean (see table).[62]
Records of giant squid specimens sorted by region and method of capture (from Guerra et al., 2011)
Region Number of specimens % of total Found stranded or floating (%) From fishing (%) From predators (%) Method of capture unknown (%)
NE Atlantic 152 22.5 49 31 15 5
NW Atlantic 148 21.9 61 30 1 8
SE Atlantic 60* 8.9 10 60 17 13
SW Atlantic 6 0.9 50 16 1 33
NE Pacific 43 6.4 7 56 30 7
NW Pacific 30* 4.4 30 35 30 5
SE Pacific 10 1.5 90 10 0 0
SW Pacific 183 27.0 12 41 42 5
Indian Ocean 33** 4.8 6 94 0 0
W Mediterranean 3 0.4 100 0 0 0
Equatorial/tropical 9 1.3 11 44 45 0
All regions 677 100.0
* Underestimates according to Guerra et al. (2011)
** Includes records from Durban, South Africa
"Wanted" poster issued by Frederick Aldrich on 24 August 1988. The flailing giant squid is from an illustration by Canadian wildlife artist Glen Loates, known for his naturalistic depictions of "Architeuthis in action",[63] which were based on collaborations with Aldrich and which Richard Ellis described as "certainly the most accurate and exciting depictions of the monster ever drawn".[64]

Procurement, preservation, and display

Preserved giant squid specimens are much sought after for both study and display.

Logy Bay, Newfoundland, assembled a substantial early collection of giant squid; as of 1971, it held 8 specimens, with the remains of 3 displayed together in a tank by the main entrance.[67]

national museums and 14 in private institutions; see table below). The purpose-built Museo del Calamar Gigante in Luarca, Spain, had by far the largest collection on public display (4 females and 1 male[68]), but many of the museum's 14 or so total specimens were destroyed during a storm on 2 February 2014.[69] At least 13 specimens were exhibited in Japan as of February 2017, of which 10 had been acquired since 2013.[70]

A number of fragmentary giant squid remains were displayed as part of "In Search of Giant Squid", a Smithsonian travelling exhibition curated by Clyde Roper that visited a dozen US museums and other educational institutions between September 2004 and August 2009.[71] The exhibition opened its national tour at Yale University's Peabody Museum of Natural History, which has maintained a strong association with the giant squid from the time of the Newfoundland strandings in the 1870s. Preparations for the Peabody exhibition, overseen by site curator Eric Lazo-Wasem, uncovered giant squid material in the museum's collections that was not previously known to be extant, including original specimens from Addison Emery Verrill's time.[72]

"Architeuthis is an elusive creature. Its occasional appearance on various beaches around the world has provided hardly more than a glimpse of its majestic and intimidating appearance, and hauling it out of the water in a trawl does it no justice either. Papier-mâché or fiberglass models have given us a sense of its size and shape, but they have not captured its mystery and vitality. The spirit of Architeuthis may well be uncapturable; at least no museum has even come close to this fabulous creature—the only living animal for which the term sea monster is truly applicable."

Richard Ellis, from the closing remarks of his 1997 article "The models of Architeuthis"[73]

In the late 19th century, the giant squid's popular appeal and desirability to museums—but scarcity of preserved specimens—spawned a long tradition of "life-sized" models that continues to the present day.

Logy Bay specimen of 1873 (#30) and on several Newfoundland specimens from the 1960s,[82] particularly the one found off Conche in October 1964 (#169).[83]

Real giant squid specimens have traditionally been preserved in either solutions of

wet specimen several years later. Additionally, a number of other specimens have been temporarily placed on public display in a frozen or chilled state (e.g. #564, 585, 592, 603, 662, 669, 675, 677, 679, and 680
).

Reported sizes

Trondheimsfjord, Norway, on 2 October 1954 (#136), being examined by Professors Erling Sivertsen and Svein Haftorn. This specimen measured 9.24 m in total length and had a mantle
length of 1.79 m.

Giant squid size—long a subject of both popular debate and academic inquiry

Thimble Tickle specimen" (#45) reported by Verrill (1880a:191) is often cited as the largest giant squid ever recorded,[nb 10] and the 55 ft 2 in (16.81 m) (or 57 ft [17.37 m]) specimen described by Kirk (1888) as Architeuthis longimanus (#62)—a strangely proportioned animal that has been much commented on—is sometimes cited as the longest.[89] It is now thought likely that such lengths were achieved by great lengthening of the two long feeding tentacles, analogous to stretching elastic bands, or resulted from inadequate measurement methods such as pacing.[90]

Based on a 40-year data set of more than 50

#480), as the "longest scientifically verified" and "largest recorded and well-preserved specimen in the contemporary, peer-reviewed literature". Charles G. M. Paxton performed a statistical analysis using literature records of giant squid specimens and concluded that "squid with a conservative TL of 20 m [66 ft] would seem likely based on current data",[93] but the study has been heavily criticised by experts in the field.[94]

Architeuthis dux, from McClain et al. (2015) (see also linear regressions). The 2,000 lb (910 kg) extreme outlier (#22) is from an estimate mentioned in Verrill (1880a:181) and is unlikely to be accurate; the next most massive individual in the data set was only 700 lb (320 kg) and 95% of specimens were below 250 kg (550 lb). Similarly, 95% of individuals had recorded mantle lengths below 3.26 m (10.7 ft) and total lengths below 15.26 m (50.1 ft).[95]

O'Shea & Bolstad (2008) give a maximum mantle length (ML) of 225 cm (7.38 ft) based on the examination of more than 130 specimens, as well as beaks recovered from sperm whales (which do not exceed the size of those found in the largest complete specimens), though there are recent scientific records of specimens that slightly exceed this size (such as #371, a 240 cm (7.9 ft) ML female captured off Tasmania, Australia; see also #647, with an estimated 2.15–3.06 m ML). Remeslo (2011) and Yukhov (2014:248) give a maximum mantle length of 260 cm (8.5 ft) for females from southern waters. Questionable records of up to 500 cm (16 ft) ML can be found in older literature.[96] Paxton (2016a) accepts a maximum recorded ML of 279 cm (9.15 ft), based on the Lyall Bay specimen (#47) reported by Kirk (1880:312), but this record has been called into question as the gladius of this specimen (which should approximate the mantle length) was said to be only 190 cm (6.2 ft) long.[94]

Including the head and arms but excluding the tentacles (standard length), the species very rarely exceeds 5 m (16 ft) according to O'Shea & Bolstad (2008). Paxton (2016a) considers 9.45 m (31.0 ft) to be the greatest reliably measured SL, based on a specimen (#46) reported by Verrill (1880a:192), and considers specimens of 10 m (33 ft) SL or more to be "very probable", but these conclusions have been criticised by giant squid experts.[94]

524). Giant squid are sexually size dimorphic, with the maximum weight for males estimated at 150 kg (330 lb),[92] though heavier specimens have occasionally been reported (see #412 for a 190 kg (420 lb) specimen). Similarly, Remeslo (2011) and Yukhov (2014:248) give maximum masses of 250–260 kg (550–570 lb) and 150 kg (330 lb) for females and males, respectively, based on records from southern latitudes. Roper & Jereb (2010:121) give a maximum weight of up to 500 kg (1,100 lb), and "possibly greater". Discredited weights of as much as a tonne (2,200 lb) or more are not uncommon in older literature (see e.g. #22, 114, and 117).[97]

The giant squid and the distantly related

dinner plate–sized" eyes (e.g. #37) are largely corroborated by modern measurements, with an accepted maximum diameter of at least 27 cm (11 in) and a 9 cm (3.5 in) pupil (based on #248).[98]

Species identifications

The taxonomy of the giant squid genus

mitogenomic analysis of Winkelmann et al. (2013) supports the existence of a single, globally distributed species (A. dux).[104] The same conclusion was reached by Förch (1998)
on the basis of morphological data.

The literature on giant squid has been further muddied by the frequent misattribution of various squid specimens to the genus Architeuthis, often based solely on their large size. In the academic literature alone, such misidentifications encompass at least the

#[9]; Ommastrephidae). This situation is further confused by the occasional usage of the common name 'giant squid' in reference to large squids of other genera.[106][nb 11]

List of giant squid

Sourcing and progenitors

Architeuthis, wrote its introduction.[108]

The present list generally follows "

the more recent specimens are supported by reports from the news media
, including newspapers and magazines, radio and television broadcasts, and online sources.

Earlier efforts to compile a list of all known giant squid encounters throughout history include those of marine writer and artist Richard Ellis.[109] Ellis's first list, published as an appendix to his 1994 work Monsters of the Sea, was probably the first such compilation to appear in print and was described in the book's table of contents as "the most complete and accurate list of the historical sightings and strandings of Architeuthis ever attempted".[110] Ellis's much-expanded second list, an appendix to his 1998 book The Search for the Giant Squid, comprised 166 entries spanning four and a half centuries, from 1545 to 1996.[111] Records which appear in Ellis's 1998 list but are not found in Sweeney & Roper's 2001 list have a citation to Ellis (1998a)—in the page range 257–265—in the 'Additional references' column of the main table.[nb 12]

In addition to these global specimen lists, a number of regional compilations have been published, including

scholarly papers unto themselves.[112]

Scope and inclusion criteria

The list includes records of giant squid (genus

Architeuthis) either supported by a physical specimen
(or parts thereof) or—in the absence of any saved material—where at least one of the following conditions is satisfied: the specimen was examined by an expert prior to disposal and thereby positively identified as a giant squid; a photograph or video recording of the specimen was taken, on the basis of which it was assigned to the genus Architeuthis by a recognised authority; or the record was accepted as being that of a giant squid by a contemporary expert or later authority for any other reason, such as the perceived credibility of the source or the verisimilitude of the account.

Nineteenth century engraving by W. A. Cranston of a giant squid attacking a boat (see #29). Only sightings deemed authentic by published experts are included in the list.

Purported sightings of giant squid lacking both

photo manipulation have been used to perpetrate hoaxes involving giant squid and these are occasionally circulated as records of actual news events, often accompanied by fictional background stories.[162] Such records are likewise excluded, as are speculative misidentifications with no scientific basis.[163]

The earliest surviving records of very large squid date to

Mediterranean are exceedingly rare). Basque and Portuguese cod fishermen observed what were likely giant squid carcasses in the waters of the Grand Banks of Newfoundland as early as the 16th century,[166] but conclusive evidence is similarly lacking. The earliest specimens identifiable as true giant squid are generally accepted to be ones from the early modern period in the 17th and 18th centuries,[167] and possibly as far back as the 16th century (#1).[168]

All developmental stages from hatchling to mature adult are included. In the literature there is a single anecdotal account of a giant squid "egg case",

sucker scars found on sperm whales
—falls outside the scope of this list.

Specimens misassigned to the genus Architeuthis in print publications or news reports are included, but are clearly highlighted as misidentifications.

List of specimens

Records are listed chronologically in ascending order and numbered accordingly. This numbering is not meant to be definitive but rather to provide a convenient means of referring to individual records. Specimens incorrectly assigned to the genus Architeuthis are counted separately, their numbers enclosed in square brackets, and are highlighted in pink ( ). Records that cover multiple whole specimens, or remains necessarily originating from multiple individuals (e.g. two lower beaks), have the '

from the 21st century) have the 'Nature of encounter
' cell highlighted in yellow ( ). Where a record falls into more than one of these categories, a combination of shadings is used. Where an image of a specimen is available, this is indicated by a camera symbol (📷) that links to the image.

Giant squid (Architeuthis dux), modified from an illustration by Addison Emery Verrill (Verrill, 1880a: pl. 20; based on #42), showing the exceptionally long feeding tentacles, which are often missing or damaged in recovered specimens. Some of the more extreme published giant squid measurements have been attributed to artificial lengthening of these tentacles.[97] Almost the entire bulk of the animal—that is, the mantle, head, and arms—takes up less than half of its total length; the absence of the tentacles, therefore, has a great effect on the animal's total length but very little on its mass.
  • Date – Date on which the specimen was first captured, found, or observed. Where this is unknown, the date on which the specimen was first reported is listed instead and noted as such. All times are local.
  • Location – Area where the specimen was encountered, including coordinates and depth information where available. Given as it appears in the cited reference(s), except where additional information is provided in square brackets. The quadrant of a major ocean in which the specimen was found is given in curly brackets (e.g. {NEA}; see Oceanic sectors).
  • Nature of encounter – Circumstances in which the specimen was recovered or observed. Given as they appear in the cited reference(s), although "washed ashore" encompasses all stranded animals.
  • Identification – Species- or
    vernacular name
    has been applied to the specimen (e.g. "giant squid" or a non-English equivalent), this is given instead.
  • Material cited – Original specimen material that was recovered or observed. "Entire" encompasses all more-or-less complete specimens. Names of anatomical features are retained from original sources (e.g. "jaws" may be given instead of the preferred "beak", or "body" instead of "mantle"). The specimen's state of preservation is also given, where known, and any missing parts enumerated (the tentacles, arm tips, reddish skin and eyes are the parts most often missing in stranded specimens, owing to their delicate nature and/or preferential targeting by scavengers).
  • Material saved – Material that was kept after examination and not discarded (if any). Information may be derived from outdated sources; material may no longer be extant even if stated as such.
  • Sex – Sex and sexual maturity of the specimen.
  • Size and measurements – Data relating to measurements and counts. Abbreviations used are based on standardised acronyms in
    arithmetic precision and original units preserved (metric conversions are shown alongside imperial measurements), though some of the more extreme lengths and weights found in older literature have since been discredited
    .
  • Repository – Institution in which the specimen material is deposited (based on cited sources; may not be current), including
    boldface
    . If an author has given a specimen a unique identifying number (e.g. Verrill specimen No. 28), this is included as well, whether or not the specimen is extant.
  • Main references – The most important sources, typically ones that provide extensive data and/or analysis on a particular specimen (often primary sources). Presented in author–date parenthetical referencing style, with page numbers included where applicable (those in square brackets refer either to unpaginated works or English translations of originally non-English works; see Full citations). Only the first page of relevant coverage is given, except where this is discontinuous. Any relevant figures ("figs.") and plates ("pls.") are enumerated.
  • Additional references – References of lesser importance or primacy, either because they provide less substantive information on a given record (often secondary sources), or else because they are not easily obtainable or possibly even extant (e.g. old newspaper articles, personal correspondence, and television broadcasts) but nonetheless mentioned in more readily accessible published works (see Full citations).
  • Notes – Miscellaneous information, often including persons and vessels involved in the specimen's recovery and subsequent handling, and any dissections, preservation work or scientific analyses carried out on the specimen. Where animals have been recorded while alive this is also noted. Material not referable to the genus Architeuthis, as well as specimens on public display, are both highlighted in bold (as "Non-architeuthid" and "On public display", respectively), though the latter information may no longer be current.

The total number of giant squid records listed across this page and successive lists is 714, though the number of individual animals covered is greater (the additional number exceeding 250) as some records encompass multiple specimens (indicated in grey). Additionally, 13 records relate to specimens misidentified as giant squid (indicated in pink).

# Date Location Nature of encounter Identification Material cited Material saved Sex Size and measurements Repository Main references Additional references      Notes     
1
(📷)
c. 1546[nb 15] Øresund, near Malmö, Denmark–Norway [since 1658 Malmö has been part of Sweden]
{NEA}
Found washed ashore; "caught live"[175] "sea monk"; Architeuthis monachus Steenstrup in Harting, 1860; Jenny Haniver made from a skate;[176] Squatina squatina (angelshark)[177] Entire? Undetermined ?WL: ≈3 m Hamer (1546:[1], fig.); Belon (1553:38, fig.); Rondelet (1554:492, fig.); Belon (1555:32, fig.); Lycosthenes (1557:609, fig.); Gessner (1558:519, fig.); Rondelet (1558:361, fig.); Sluperius (1572:"89", 105, fig.); Vedel (1575); Huitfeldt ([1595]:1545); Gessner (1604:438, fig.); Stephanius ([c. 1650]:344); Steenstrup (1855a:63, 3 figs.); Roeleveld & Knudsen (1980:293, 3 figs.); Ellis (1998a:60, fig.); Paxton & Holland (2005:39, fig. 1) records of Björn Jónsson á Skarðsá;[nb 16] Scheuchzer (1716:153); Holberg (1733:379); Lönnberg (1891:36); Nordgård (1928:71); Tambs-Lyche (1946:288); Carrington (1957:58, fig.); Muus (1959:170); Russell & Russell (1975:94); Strauss (1975:393, fig.); Aldrich (1980:55); Roeleveld (N.d.) Contemporaneously regarded as a "
sea bishop has also been interpreted as a giant squid carcass[180] or else a Jenny Haniver made from a skate.[181]
2 autumn 1639
Hunevandsyssel), Iceland[182]

{NEA}
Found washed ashore Architeuthis sp. Entire One arm BL+HL: ≈6 ft (1.8 m); AL: ≈3 ft (0.91 m); TL: ≈16–18 ft (4.9–5.5 m); BC: ≈3–4 ft (0.91–1.22 m) Thingøre monastery; "museum at Copenhagen" (ZMUC?)[183] Jónsson ([c. 1645]:238); Ólafsson (1772:716); Steenstrup (1849:950/[9]); Steenstrup (1898:425/[272]); Ellis (1998a:65) Packard (1873:87); Verrill (1875b:84); Robson (1933:691); Muus (1959:170); Berger (2009:260) Original Icelandic account is from the contemporaneous Annálar Björns á Skarðsá and has been translated into English.[nb 17] Crude drawing of animal mentioned by Eggert Ólafsson was lost with most of his books when his boat capsized off Iceland in 1768, leading to his death.[185] Identified by Japetus Steenstrup as decapod cephalopod in 1849.
3
(📷)
c. 15 October 1673 Dingle-I-cosh, Kerry, Ireland
{NEA}
Found floating at surface, in process of washing ashore, alive Dinoteuthis proboscideus More, 1875; Architeuthis monachus;[186] Ommastrephes (Architeuthis) monachus[187] Entire Two arms, buccal mass, and suckers taken to Dublin TL: ≈11 ft (3.4 m) + 9 ft (2.7 m); AL: ≈6–8 ft (1.8–2.4 m); "liver": 30 lb (14 kg) Undetermined [NMI?]; holotype of Dinoteuthis proboscideus More, 1875 [Anon.] (c. 1673); Hooke et al. (c. 1674:[1], fig.); More (1875a:4526); Verrill (1875c:214); Tryon (1879b:185); Ellis (1998a:66); Sueur-Hermel (2017:64) More (1875b:4571); Massy (1909:30); Ritchie (1918:137); Robson (1933:692); Rees (1950:40); Hardy (1956:285); Collins (1998:489) Found by James Steward. Original material relating to this specimen consists of: a broadsheet printed in
handbill;[190] and an eight-page booklet printed in London with a woodcut reproduction of the illustration in the broadsheet (both originating from a painting on canvas brought to London, as it was impossible to preserve the carcass).[191]
4 1680 Ulvangen Fjord, Alstadhoug parish, Norway
{NEA}
Not stated Entire? Pontoppidan (1753:344) Steenstrup (1857:184/[18]); Grieg (1933:19)
5 1770 Jutland, Denmark
{NEA}
Unknown Muss (1959) Ellis (1998a:257)
6 27 May 1785 Grand Banks, Newfoundland
{NWA}
Found floating at surface, dead Architeuthis sp. BL: 7 ft (2.1 m) Cartwright (1792:44); Thomas ([1795]:183); Aldrich (1991:457) Found during George Cartwright's sixth and final voyage to Newfoundland and Labrador. Spotted at 10 am surrounded by birds. Head broke off during retrieval. Described as "a large squid [...] when gutted, the body filled a pork barrel, and the whole of it would have filled a tierce".
7 November or December 1790 Arnarnaesvik, Modruvalle, Iceland
{NEA}
Found washed ashore Entire None; used for cod bait "longest tentacula": >3 fathoms (5.5 m); "body right from the head": 3.5 fathoms (6.4 m); "so thick that a fullgrown man could hardly embrace it with his arms" Steenstrup (1849:952/[11]); Steenstrup (1898:429/[276]); Ellis (1998a:68) February 1792 diary of Sveinn Pálsson (in library of Icelandic Literary Society, in Copenhagen); Verrill (1875b:84); Robson (1933:691) Called Kolkrabbe ('coal-crab') by local people. Identified by Japetus Steenstrup as decapod cephalopod in 1849.
8 1700s (reported 1795) Freshwater Bay, near mouth of St. John's harbour, Newfoundland
{NWA}
Unknown Architeuthis sp. Thomas ([1795]:183); Aldrich (1991:457)
9 1700s Grand Banks, Newfoundland
{NWA}
Unknown Architeuthis sp. Aldrich (1991:457)
10 1798 north coast of Denmark
{NEA}
Not stated "gigantic squid" Unknown "museum at Copenhagen" (ZMUC?) Packard (1873:87) Ellis (1998a:257)
11 9 January 1802 off Tasmania, Australia
{SWP}
Found at surface, alive ?Loligo ["vraisemblablement du genre Calmar [Loligo, Lamarck]"] "size of a barrel" ["grosseur d'un tonneau"]; AL: 1.9–2.2 m; AD: 18–21 cm Péron (1807:216) Quoy & Gaimard (1824:411); Ellis (1998a:257) Péron (1807:216) wrote: "it rolled with noise in the midst of the waves, and its long arms, stretched out on their surface, stirred like so many enormous reptiles" (translated from the French).
12 between 1817 and 1820 Atlantic Ocean, near equator
{?}
Found floating at surface "énorme calmar" Partial remains; "tentacles" ("tentacules") missing WT: 100 "
livres
" [estimate]; WT: 200 "livres" [estimate; if complete]
Quoy & Gaimard (1824:411) Packard (1873:88); Ellis (1998a:257) Found at surface in calm weather. Quoy & Gaimard (1824:411) opined: "it is easy to imagine that one of these terrible molluscs could readily remove a man from a fairly large boat, but not a medium-tonnage vessel, still less tilting this vessel and endangering it, as some would like to believe" (translated from the French).
13
(📷)
December 1853
North Jutland, coast of Skagerack, Denmark

{NEA}
Found washed ashore Architeuthis monachus Entire Jaws only; radula discarded after poor preservation; jaws cut out; portion used for bait; remainder buried after 2 days WT: 80–85 kg; jaw measurements Steenstrup (1898:423/[270]) ZMUC catalog no. CEP-133; holotype of Architeuthis monachus Steenstrup, 1857[192] Steenstrup (1855b:[14]); Harting (1860:11); Steenstrup (1898:415/[258], pl. 1 figs. 1–2); Kristensen & Knudsen (1983:222) Steenstrup (1857:[18]); Packard (1873:87); Gervais (1875:91); Verrill (1875b:84); Verrill (1880a:238, pl. 25 fig. 3); Verrill (1882c:51, pl. 12 fig. 3); Posselt (1890:144); Nordgård (1928:71) "Architeuthis monachus" Steenstrup = nomen nudum[193]
14
(📷)
5 November 1855 western Atlantic Ocean, near Bahamas (31°N 76°W / 31°N 76°W / 31; -76 (Giant squid specimen, 5 November 1855))
{NWA}
Not stated; presumably found floating at surface Architeuthis dux Steenstrup, 1857; Architeuthis titan[194] Various parts Gladius, mouthparts, part of arm, several suckers, and what may be hectocotylus[195] Male WL: 377 cm; AL: 1/2 whole length;[196] beak measurements; GL: 6 ft (1.8 m)[197] ZMUC catalog no. CEP-97 (or CEP-000097) and NHMD-77320 (multiple parts, each in its own glass vessel: gladius, mouthparts, part of arm, several suckers, and what may be hectocotylus);
Bergen Museum[198]
Steenstrup (1857:[18]); Steenstrup (1882:[160]); Steenstrup (1898:413, 450/[260, 298], pls. 3–4); Tryon (1879b:186, pl. 86 fig. 388); Kristensen & Knudsen (1983:222); Glaubrecht & Salcedo-Vargas (2000:273); [NHMD] (2019) Packard (1873:87); Verrill (1875b:84); Posselt (1890:144); Toll & Hess (1981:753) Obtained by Capt. Vilhelm Hygom.
Museum für Naturkunde in Berlin, which was incorporated into collection in 1883 according to catalogue entry. Preserved in 70% ethanol.[195]
15 December 1855 Aalbaekbugten, Denmark
{NEA}
Found washed ashore Architeuthis sp. Entire? Undetermined None Muus (1959:170) Posselt (1890:144)
16
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Unknown (reported 1860) Unknown
{?}
Not stated Architeuthis dux;[199] ?Ommastrephes hartingii;[200] Architeuthis hartingii (Verrill, 1875);[201] nomen nudum[202] Jaws, buccal mass, detached arm suckers Jaws, buccal mass, detached arm suckers ASD: 1.05 in (2.7 cm) Utrecht University Natural History Museum; holotype of Loligo hartingii Verrill, 1875. Harting specimen No. 1 Harting (1860:2, pl. 1); Kent (1874d:491); Verrill (1875b:85, fig. 28); Tryon (1879b:149, 184, pl. 60 figs. 194–195); Verrill (1880a:240, pl. 16 fig. 8, pl. 25 fig. 1); Verrill (1882c:52, pl. 12 figs. 1–1c); Pfeffer (1912:37) Dell (1970:27)
17 1860 or 1861 between Hillswick and Scalloway, Shetland, Scotland
{NEA}
Found washed ashore Architeuthis monachus Steenstrup, 1857; Architeuthis dux Steenstrup, 1857[203] Undetermined TL: 16 ft (4.9 m); AL: ≈8 ft (2.4 m); BL: ≈7 ft (2.1 m) Jeffreys (1869:124); Stephen (1944:263) More (1875b:4571); Pfeffer (1912:26); Rees (1950:40); Collins (1998:489)
18
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30 November ?1861 [=1860 Rees & Maul] about 20 miles (32 km) northeast of Teneriffe, Canary Islands
{NEA}
Found floating at surface Loligo bouyeri;[204] ?Ommastrephes bouyeri[200] Entire, decomposed None BL: 15–18 ft (4.6–5.5 m) None Bouyer (1861:1263); Crosse & Fischer (1862:135); Bouyer (1866:275, fig.); Kent (1874a:180); Verrill (1875b:86); Tryon (1879b:149, 184, pl. 59); Bourée (1912:113, fig. 108); Aldrich (1978:2); Ellis (1998a:5, 78); Heuvelmans (2003:185, figs. 95–96, 100) Frédol (1865:314, pl. 13); Figuier (1866:464, fig. 362); Frédol (1866:362); Mangin (1868:321); Meunier (1871:245); Kent (1874d:491); Gervais (1875:93); Lee (1883:38, fig. 8); Rees & Maul (1956:266); Carrington (1957:53, pl. 3b); Muntz (1995:19, fig. 11); Lagrange (2009:19) Observed only by officers of the French gunboat
Twenty Thousand Leagues Under the Sea.[3] Iconography discussed by Lagrange (2009)
.
19 1862 North Atlantic
{NEA/NWA}
Unknown Crosse & Fischer (1862) Ellis (1998a:258)
[1]
(📷)
Unknown; 1870? Cape Sable, Nova Scotia, Canada
{NWA}
Found washed ashore Architeuthis megaptera Verrill, 1878 [=Sthenoteuthis pteropus (Steenstrup, 1855)][205] Entire Entire BL: 14 in (36 cm); BL+HL: 19 in (48 cm); EL: 43 in (110 cm); TL: 22–24 in (56–61 cm); AL: 6.5–8.5 in (17–22 cm); FW: 13.5 in (34 cm); FL: 6 in (15 cm); extensive additional measurements NSMC catalog no. 1870-Z-2; YPM catalog nos. IZ 017932 Archived 3 August 2019 at the Wayback Machine (sucker) & IZ 017713; holotype of Architeuthis megaptera Verrill, 1878;[206] Verrill specimen No. 21 ("Cape Sable specimen") Verrill (1878:207); Tryon (1879b:187); Verrill (1880a:193, pl. 21); Verrill (1882c:17, pl. 16 figs. 1–9) Non-architeuthid. Collected by J.M. Jones.
20 September 1870 Waimarama, east coast of Wellington, New Zealand
{SWP}
Found washed ashore Entire Beak BL+HL: 10 ft 5 in (3.18 m); BC: 6 ft (1.8 m); AL: 5 ft 6 in (1.68 m) In Kirk's possession; Kirk specimen No. 1 Kirk (1880:310); Verrill (1881b:398) Meinertzhagen letter 27 June 1879 to Kirk; Pfeffer (1912:32); Dell (1952:98) Mr. Meinertzhagen sent beak, saved by third party (unidentified), to Kirk. Natives called specimen a "taniwha".
21 1870 (winter) Lamaline, Newfoundland
{NWA}
Found washed ashore Architeuthis monachus of Steenstrup Two specimens; entire? None?; used as fish bait[207] Two; EL: 40 ft (12 m) and EL: 47 ft (14 m) None?; Verrill specimen Nos. 8 & 9 ("Lamaline specimens") Murray (1874a:162); Verrill (1875a:36); Verrill (1880a:187); Verrill (1882c:11) Data from Mr. Harvey letter citing Rev. M. Gabriel's statement to Harvey.
22
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October 1871 Grand Banks, Newfoundland
{NWA}
Found floating at surface Architeuthis princeps Verrill, 1875 Entire; part used as bait Jaws obtained from Baird for examination by Verrill BL: ≈15 ft (4.6 m); BD: 19 in (48 cm); AL: ≈10 ft (3.0 m) [mutilated]; AD: 7 in (18 cm); AC: 22 in (56 cm); beak; BC: 4 ft 8 in (1.42 m); WT: 2,000 lb (910 kg) Jaws at NMNH[208] (no longer extant?[209]); lower jaw is syntype of Architeuthis princeps Verrill, 1875b; Verrill specimen No. 1 ("Grand Banks specimen" [1st]) Packard (1873:91); Verrill (1874a:158); Verrill (1874b:167); Verrill (1875b:79, fig. 27); Verrill (1880a:181, 210, pl. 18 fig. 3); Verrill (1882c:5, pl. 11 figs. 3–3a) Pfeffer (1912:20); Frost (1934:100) Taken by Capt. Campbell, Schooner B.D. Haskins.
23 1871 Wellington, New Zealand
{SWP}
?EL: 16 ft (4.9 m) Dell (1952) Ellis (1998a:258)
24 1872 (autumn or winter)
Coomb's Cove, Fortune Bay, Newfoundland

{NWA}
Found alive in shallow water, having been driven ashore in heavy sea Entire; "one long arm missing" (later changed to both present) BL: 10 ft (3.0 m); BD: 3–4 ft (0.91–1.22 m); TL: 42 ft (13 m); AL: ≈6 ft (1.8 m); AD: 9 in (23 cm); skin + flesh: 2.25 in (5.7 cm) thick; EL: 52 ft (16 m) Unknown; Verrill specimen No. 3 ("Coombs' Cove specimen") Verrill (1874a:159); Verrill (1874b:167); Verrill (1875a:35); Verrill (1880a:183); Verrill (1882c:7) Owen (1881:163); Frost (1934:101) Specimen had a reddish colour. Verrill's data taken from newspaper accounts and 15/VI/1873 T.R. Bennett letter to Prof. Baird. Verrill (1880a:186) states his No. 6 is same specimen as No. 3; this cannot be correct, since capture date for No. 6 is clearly stated as December 1874 by Verrill (1875c:213).[209]
25
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December 1872 Bonavista Bay, Newfoundland
{NWA}
Found washed ashore ?Architeuthis dux;[210] ?Architeuthis harveyi[201] Entire (damaged arms) Pair of jaws and two suckers TL: 32 ft (9.8 m); AL: ≈10 ft (3.0 m); BL: ≈14 ft (4.3 m) [estimate]; BC: 6 ft (1.8 m) NMNH; YPM catalog no. IZ 034835. Verrill specimen No. 4 ("Bonavista Bay specimen") (1875a:33); and possibly also Verrill specimen No. 11 ("Second Bonavista Bay specimen") (1875b:79) Verrill (1874a:160); Verrill (1874b:167); Verrill (1875a:33, fig. 11); Verrill (1875b:79); Verrill (1880a:184, 187, pl. 16 figs. 5–6, pl. 25 fig. 5); Verrill (1882c:8, 11, pl. 3 figs. 4–4a, pl. 4 figs. 1–1a) Pfeffer (1912:19); Frost (1934:101) Material from Rev. A. Munn, through Prof. Baird to Verrill.
26
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Unknown (reported 1873) North Atlantic Ocean
{NWA}
From sperm whale stomach Architeuthis princeps Verrill, 1875; Ommastrephes (Architeuthis) princeps[187] Upper and lower jaws Upper and lower jaws Beak measurements Presented by Capt. N.E. Atwood of Provincetown, Massachusetts to EI;[211] PASS;[201] syntype of Architeuthis princeps Verrill, 1875b; Verrill specimen No. 10 ("Sperm-whale specimen") Packard (1873:91, fig. 10); Verrill (1875a:22); Verrill (1875b:79, figs. 25–26); Tryon (1879b:185, pl. 85); Verrill (1880a:187, 210, pl. 18 figs. 1–2); Verrill (1882c:11, pl. 11 figs. 1–2) Frost (1934:101) First reported by Alpheus Spring Packard in February 1873. Verrill states Packard's illustration is inaccurate.
27
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Unknown (reported 1873) Unknown; possibly east coast of South America[212]
{SWA}?
Not stated Architeuthis monachus;[213] Plectoteuthis grandis Owen, 1881; Architeuthis sp.? (grandis);[214] nomen nudum[202] Sessile arm Arm AL: 9 ft (2.7 m); AC: 11 in (28 cm); ASD: ≤0.5 in (1.3 cm); total size and size of various missing parts estimated by Lee (1875:114) BMNH; holotype of Plectoteuthis grandis Owen, 1881 Kent (1874a:179); Kent (1874d:493); Lee (1875:113); Verrill (1875b:86); Owen (1881:156, pls. 34–35); Verrill (1881b:400); Verrill (1882b:72); Steenstrup (1882:[160]); Pfeffer (1912:37) Dell (1970:27) "No history relating to it has been preserved", but first examined by Henry Lee in May 1873, having been in BMNH collections for "long" time.[33] Bore c. 300 suckers.
28 1873 Yedo [Tokyo] fishmarket, Japan
{NWP}
Purchased Megateuthis martensii Hilgendorf, 1880; Nomen spurium[215] 'Entire', missing head, "abdominal sac", ends of tentacles and arms[216] Not specified ML: 186 cm; WL: 414 cm; HL: 41 cm; AL: 197 cm [longest]; ASD: 1.5 cm (with 37 cusps); EyD: 200 mm ZMB Moll. 34716 + 38980; holotype of Megateuthis martensii Hilgendorf, 1880 [34716a: eyeball, 200 mm diameter, dry; 34716b: pieces of arm and gladius, suckers; 34716c: larger piece of arm with suckers; 38980: four suckers from holotype arm piece] Hilgendorf (1880:67); Pfeffer (1912:31); Sasaki (1929:227); Glaubrecht & Salcedo-Vargas (2000:276) Owen (1881:163); Sasaki (1916:90) Second specimen from Tokyo fishmarket seen by Franz Martin Hilgendorf and used for description of gladius. Of other specimen, Hilgendorf saved "parts of an arm, the covering of the eye, and a fragment of the gladius" ("Theile eines Armes, die Hüllen des Auges, und ein Bruchstück des Schulpes").[217] Model of specimen placed in Exhibition of Fishery in Berlin.
29
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26 October 1873 off
{NWA}
Found floating at surface, alive Megaloteuthis harveyi Kent, 1874; Architeuthis monachus of Steenstrup;[218] ?Architeuthis harveyi[219] Entire One tentacle; one arm discarded (see
club measurement from Harvey letter;[220] BL: ≈10 ft (3.0 m); EL: ≈60 ft (18 m) [estimate]; AL: 6 ft (1.8 m); AD: 10 in (25 cm); TSC: ≈180; beak as large "as a six-gallon keg"; "tail" 10 ft (3.0 m) across[221]
YPM?; holotype of Megaloteuthis harveyi Kent, 1874; Verrill specimen No. 2 ("Conception Bay specimen") Harvey (1873a); Harvey (1873b); Harvey (1873c); Harvey (1874a:67, fig.); Murray (1874a:161); Murray (1874b:120); Verrill (1874a:159); Verrill (1874b:167); Kent (1874a:178, 182); Agassiz (1874:226); Kent (1874d:32); Buckland (1875:211); Verrill (1875a:34); Verrill (1875b:78); Verrill (1880a:181); Verrill (1881b:pl. 26 fig. 5); Verrill (1882b:74); Verrill (1882c:5, pl. 4 figs. 3–3a); Hatton & Harvey (1883:238); Harvey (1899:732, fig.); Ellis (1998a:81); Haslam (2017) "13 December Field"; [Anon.] (1873:2); Harvey (1873d:2); [Anon.] (1874:333); de La Blanchère (1874:197, fig.); Rathbun (1881:266, fig.); Owen (1881:161, pl. 33 fig. 2); Lee (1883:42, fig. 9); [Anon.] (1902b:6, fig.); Pfeffer (1912:19); Frost (1934:100); Aldrich (1991:457); Packham (1998); Dery (2013) Struck by Theophilus Picot from boat whereupon it "attacked" the boat; veracity of account has been questioned.
E. Annie Proulx.[231]
30
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25 November? 1873
Logy Bay (≈3 miles from St. John's), Newfoundland

{NWA}
In herring net ?Architeuthis monachus of Steenstrup;[210] Ommastrephes (Architeuthis) monachus;[200] Architeuthis harveyi (Kent, 1874)[201] Entire (badly mutilated, head severed, eyes missing, etc.) Miscellaneous parts obtained from Rev. M. Harvey (gladius and ?) (see
club
description; extensive description of reconstructed parts
YPM catalog nos. IZ 009634 (beak and limbs), IZ 017924 (radula), IZ 017925, IZ 017926 & IZ 034968[permanent dead link]. Verrill specimen No. 5 ("Logie Bay specimen") Harvey (1873d:2); Verrill (1874a:160); Verrill (1874b:167); Kent (1874a:181); Kent (1874d:32); Buckland (1875:212, 214); Verrill (1875a:22, figs. 1–6, 10); Verrill (1876:236); Tryon (1879b:184, pls. 83–84); Verrill (1880a:184, 197, pls. 13–15, pl. 16 figs. 1–4, pl. 16a); Verrill (1880b:295, pl. 13); Verrill (1882c:8, pls. 1–2, pl. 3 figs. 1–3, pl. 4 figs. 4–11, pl. 5 figs. 1–5); Hatton & Harvey (1883:240); Harvey (1899:735, fig.); Pfeffer (1912:18); Aldrich (1991:457, fig. 1A, B); Haslam (2017) Harvey in Morning Chronicle (newspaper) of St. John's; Maritime Monthly Magazine of St. John's, March 1874; several other newspapers; [Anon.] (1874:332); Lee (1883:43, fig. 10); [Anon.] (1902b:6, fig.); Frost (1934:101) Verrill's data from letter to Dr. Dawson from
shower curtain rod were subject of Preparing the Ghost (2014), a work of creative nonfiction by Matthew Gavin Frank.[234]
31 1874
Foldenfjord, Norway

{NEA}
Found washed ashore Architeuthis dux Entire None WL: ≈4 m Grieg (1933:19) Nordgård (1928:71)
32 10 May 1874 off Trincomalee, Sri Lanka (8°50′N 84°05′E / 8.833°N 84.083°E / 8.833; 84.083 (Supposed sinking of ship by giant squid, 10 May 1874))
{NIO}
Reportedly seen sinking ship Unknown The Times, 4 July 1874; Mystic Press, 31 July 1874; Lane (1957:205); Flynn & Weigall (1980); Ellis (1998a:198); Boyle (1999); Uragoda (2005:97) Welfare & Fairley (1980:74); Aldrich (1990a:5); Clarke (1992:72); Ellis (1998a:258)
Madras, which rescued five of the crew. Veracity of account has been questioned,[235] though taken seriously by Frederick Aldrich.[236] Fictionalised in Don C. Reed's 1995 novel The Kraken.[237]
33
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2 November 1874 on beach, St. Paul Island, Indian Ocean (38°43′S 77°32′E / 38.717°S 77.533°E / -38.717; 77.533 (Giant squid specimen, 2 November 1874))
{SIO}
Found washed ashore Architeuthis mouchezi Vélain (1875:1002) [nomen nudum]; Mouchezis sancti-pauli Vélain (1877:81); Ommastrephes mouchezi[200] Entire; found in advanced state of decay Tentacle(s?) and buccal mass EL: 7.15 m MNHN catalog nos. 3-2-658 & 3-2-659 (
tentacular clubs);[238]
holotype of Mouchezis sancti-pauli Vélain, 1877
Vélain (1875:1002); Vélain (1877:81 & 83, fig. 8); Vélain (1878:81 & 83, fig. 8); Tryon (1879b:184, pl. 82 fig. 378); Owen (1881:159); Pfeffer (1912:32) Gervais (1875:88); Verrill (1875c:213); Wright (1878:329) Recorded by geologist Charles Vélain during French astronomical mission to Île Saint-Paul to observe the transit of Venus. Specimen was photographed.[28]
34
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December 1874 Grand Bank, Fortune Bay, Newfoundland
{NWA}
Found washed ashore Architeuthis princeps Entire, except for tail (cut up for dog food) Jaws, one tentacular sucker EL: 42–43 ft (12.8–13.1 m); HL+BL: 12–13 ft (3.7–4.0 m); ?TL: 30 ft (9.1 m); TL: 26 ft (7.9 m); TC: 16 in (41 cm); BL: 10 ft (3.0 m); jaws YPM catalog nos. IZ 010272 (beak) & IZ 034836[permanent dead link]. Verrill specimen No. 6 and Verrill specimen No. 13 ("Fortune Bay specimen") Verrill (1875a:35); Verrill (1875c:213); Verrill (1880a:186, 188, 217, pl. 17 fig. 11); Verrill (1881b:445, pl. 54 fig. 1); Verrill (1882c:10, 12, pl. 7 fig. 1, pl. 9 fig. 11) Simms letter 27/X/1875 to Verrill; Frost (1934:102) Data from 10/XII/1873 letter from Mr. Harvey to unknown individual citing measurements taken by G. Simms; Pfeffer (1912:21). Measurements are given differently in different papers. Verrill (1880a:186) and Verrill (1882c:10) states his No. 6 is same specimen as No. 3; this cannot be correct, as capture date for No. 6 is clearly stated as December 1874 by Verrill (1875c:213).[239] Verrill (1880a:188, pl. 17) repeats record as his No. 13.
35 winter of 1874–1875 near
Harbor Grace, Newfoundland

{NWA}
Found washed ashore Destroyed None taken None; Verrill specimen No. 12 ("Harbor Grace specimen") Verrill (1875b:79); Verrill (1880a:188); Verrill (1882c:12) Frost (1934:102) "destroyed before its value became known, and no measurements are given"
36 Unknown (reported 1875) west St. Modent (on Labrador side), Strait of Belle Isle, Newfoundland
{NWA}
Found alive Architeuthis princeps or Architeuthis monachus of Steenstrup Entire None; cut up, salted, and barrelled for dog meat ?TL: 37 ft (11 m); BL+HL: 15 ft (4.6 m); EL: 52 ft (16 m); SD: ≈2 in (5.1 cm) None; Verrill specimen No. 7 ("Labrador specimen") Verrill (1875a:36); Verrill (1880a:186); Verrill (1882c:10) Dr. Honeyman article in Halifax newspaper; Frost (1934:101) Data from unidentified third party cited in Halifax newspaper article.
37
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25 April 1875 (or 26 April[240]) north-west of Boffin Island, Connemara, Ireland
{NEA}
Found immobile at surface; attacked and chased by fishermen; arms successively hacked off and eventually killed Architeuthis monachus; Architeuthis dux Steenstrup, 1857[241] Entire Beak and buccal mass, one arm ("much mutilated and decayed", missing horny rings), portions of both tentacles ("shrunk and distorted", missing horny rings on large central club suckers); head, eyes and second arm initially saved, but soon lost/destroyed TL: 30 ft (9.1 m) [fresh]; TL: 14/17 ft (4.3/5.2 m) [pickled]; CL: 2 ft 9 in (0.84 m) [shrunken]; CSD: nearly 1 in (2.5 cm); SSD: 320 in (0.38 cm); AL: 8 ft (2.4 m) [fresh]; AC: 15 in (38 cm) [fresh]; beak: ≈5+14 in (13 cm) × 3+12 in (8.9 cm); "trunk": "fully as long as the canoe"; EyD: ≈15 in (38 cm); WT: ≈6 st (38 kg) [head only]; additional sucker measurements NMI catalog no. 1995.16 (beak in spirit)[241] O'Connor (1875:4502); More (1875b:4569); More (1875c:123); Verrill (1875c:214); Massy (1909:30); Nunn & Holmes (2008) Galway Express 1875; Ritchie (1918:137); Massy (1928:32); Taylor (1932:3); Robson (1933:692); Rees (1950:40); Hardy (1956:285); Collins (1998:489) On public display. Caught by three-man
coarse fish. Found motionless at surface surrounded by gulls, becoming active upon being attacked by fishermen, swimming away "at a tremendous rate" and releasing ink. Progressively disabled with a knife (fishermen having no gaff or spare rope) as chased for 2 hours over 5 miles (8.0 km), before head eventually severed; heavy mantle allowed to sink. Specimen secured and preserved by Sergeant Thomas O'Connor of the Royal Irish Constabulary and forwarded by him to the museum of the Royal Dublin Society, Dublin (now the National Museum of Ireland – Natural History
).
38 October 1875 Grand Banks [of Newfoundland], Atlantic Ocean (chiefly 44°–44°30'N 49°30'–49°50'W)
{NWA}
Found floating at surface; "mostly entirely dead" but small minority "not quite dead, but entirely disabled" Architeuthis Multiple; mutilated by birds and fishes to varying degrees, especially limbs; No. 25 missing parts of arms; No. 26 with intact arms and tentacles None; cut up for cod bait No. 25: Filled ≈75 US gal (280 L) tub; WT: nearly 1,000 lb (450 kg) [estimate, complete]; No. 26: TL: 36 ft (11 m); Howard specimens: BL+HL?: mostly 10–15 ft (3.0–4.6 m) [excluding "arms"]; BD: ≈18 in (46 cm) [average]; AL: usually 3–4 ft (0.91–1.22 m) [incomplete]; AD: "about as large as a man's thigh" [at base]; Tragabigzanda specimens: BL+HL?: 8–12 ft (2.4–3.7 m) [excluding "arms"] None; included Verrill specimen No. 25 and Verrill specimen No. 26 Verrill (1881a:251); Verrill (1881b:396); Verrill (1882c:19) Frost (1934:103) An unusual number (≈25–30) of mostly dead giant squid found by
season
. Verrill conjectured that this mass mortality might have been due to an outbreak of disease or parasites, and/or related to their reproductive cycle.
39 c. 1876 Clifford Bay, Cape Campbell, New Zealand
{SWP}
Found washed ashore Entire Jaws[242] BL: 7 ft (2.1 m) [estimate]; EL: ≈20 ft (6.1 m) [estimate] Colonial Museum [NMNZ][242] Robson (1887:156); Kirk (1880) Pfeffer (1912:32); Dell (1952:98)
40 20 November 1876 Hammer Cove, southwest arm of Green Bay, Notre Dame Bay, Newfoundland
{NWA}
Found washed ashore Partial specimen; devoured by foxes and seabirds Piece of pen 16 in (41 cm) long WH: 18 in (46 cm); FW: 18 in (46 cm) In Harvey's possession; Verrill specimen No. 15 ("Hammer Cove specimen") Verrill (1880a:190); Verrill (1880b:284); Verrill (1882c:14) M. Harvey letter 25 August 1877 to Verrill; Frost (1934:102)
41 1877? Norway
{NEA}
Not stated Map location only Sivertsen (1955:11, fig. 4)
42
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24 September 1877
Trinity Bay, Newfoundland

{NWA}
Found washed ashore, alive Architeuthis princeps; Ommastrephes (Architeuthis) princeps[187] Entire; "nearly perfect specimen" Loose suckers (see Verrill, 1880a:220) HL+BL: 9.5 ft (2.9 m); BC: 7 ft (2.1 m); TL: 30 ft (9.1 m); AL: 11 ft (3.4 m) [longest, ventral]; AC: 17 in (43 cm) [ventral]; beak; FW: 2 ft 9 in (0.84 m) YPM catalog nos. IZ 017927, IZ 017928, IZ 017929 & IZ 017930. Verrill specimen No. 14 ("Catalina specimen") Harvey (1877); [Anon.] (1877a:266, 269, fig.); [Anon.] (1877b:867, fig.); [Anon.] (1877c:305, fig.); Verrill (1877:425); Tryon (1879b:185); Verrill (1880a:189, pl. 17 figs. 1–10, pls. 19–20); Verrill (1880b:295, pl. 12); Verrill (1882c:13, pl. 8, pl. 9 figs. 1–10, pl. 10) Owen (1881:163); Hatton & Harvey (1883:242); Pfeffer (1912:21); Frost (1934:102); Miner (1935:187, fig., 201); Ellis (1997a:31) Measured fresh by M. Harvey; examined preserved (poorly) by Verrill at
Ward's.[75] Described by Frederick Aldrich as "largest giant squid to be encountered in Newfoundland".[126]
43 October 1877
Trinity Bay, Newfoundland

{NWA}
Not stated "big squid" None None taken None; Verrill specimen No. 17 ("Trinity Bay specimen") Verrill (1880a:191); Verrill (1880b:285); Verrill (1882c:15) M. Harvey letter 17 November 1877 to Verrill citing reference to specimen by John Duffet; Frost (1934:102) Specimen cut up and used for manure.
44
(📷)
21 November 1877
Trinity Bay, Newfoundland

{NWA}
Found washed ashore, alive ?Architeuthis princeps Entire None; carried off by tide BL(+HL?): 11 ft (3.4 m); TL: 33 ft (10 m); AL: 13 ft (4.0 m) [estimate] None; Verrill specimen No. 16 ("Lance Cove specimen") Verrill (1880a:190); Verrill (1880b:285); Verrill (1882c:14) M. Harvey letter 27 November 1877 to Verrill citing measurements taken by John Duffet; Frost (1934:102) Found still alive, having "ploughed up a trench or furrow about 30 feet [9.1 m] long and of considerable depth by the stream of water that it ejected with great force from its siphon. When the tide receded it died."

(📷)
1878 (accessioned)
Catlins, New Zealand

{SWP}
Not stated Architeuthis sp. Entire? Beak BL: 7 ft (2.1 m);[243] ML: 1.6 m [estimate]; EL: ≈10 m [estimate][244]
Otago Museum
catalog no. IV119151
Lau (2021); [OM] (2021) Copedo (2022) On public display. Collected by Capt. Charles Hayward (
Otago Museum, according to museum records; rediscovered and publicised in 2021.[243] Placed on display in Animal Attic gallery.[245] Mantle and total length estimated by teuthologist Kat Bolstad based on beak measurements.[244]
45
(📷)
2 November 1878
{NWA}
Found aground offshore, alive; secured to tree with
grapnel
and rope; died as tide receded
?Architeuthis princeps Entire None; cut up for dog food BL+HL: 20 ft (6.1 m); TL: 35 ft (11 m)[nb 20] None; Verrill specimen No. 18 ("
Thimble Tickle specimen
")
Verrill (1880a:191); Verrill (1880b:285); Verrill (1882c:15); Ellis (1998a:6, 89, 107) M. Harvey letter 30 January 1879 to Discovered by fisherman Stephen Sherring and two others.
Giant Squid Interpretation Centre and "life-sized", 55-foot sculpture built near site of capture;[263] sculpture appeared on Canadian postage stamp issued in 2011[264] and has associated annual festival.[265]
46 2 December 1878
Three Arms, South Arm of Notre Dame Bay, Newfoundland

{NWA}
Found washed ashore ?Architeuthis princeps Entire, mutilated and with arms missing (only one arm "perfect") None; cut up for dog food BL+HL: 15 ft (4.6 m); BC: 12 ft (3.7 m); AL: 16 ft (4.9 m); AD: "thicker than a man's thigh" None; Verrill specimen No. 19 ("Three Arms specimen") Verrill (1880a:192); Verrill (1880b:286); Verrill (1882c:16) M. Harvey letter 30 January 1879 to Found dead by fisherman William Budgell after heavy gale. Considered by Paxton (2016a:83) as the "longest measured" standard length of any giant squid specimen.
47
(📷)
23 May 1879 Lyall Bay, Cook Strait, New Zealand
{SWP}
Found washed ashore Steenstrupia stockii Kirk, 1882 [=Architeuthis sp.?[266]] Entire, but somewhat mutilated; missing ends of tentacles Pen, beak, tongue, some suckers ML: 9 ft 2 in (2.79 m); BC: 7 ft 3 in (2.21 m); HL: 1 ft 11 in (0.58 m); BL+HL: 11 ft 1 in (3.38 m); HC: 4 ft (1.2 m); AL: 4 ft 3 in (1.30 m); AC: 11 in (28 cm); ASC: 36; TL: 6 ft 2 in (1.88 m) [incomplete]; FL: 24 in (61 cm); FW: 13 in (33 cm) (single); GL: 6 ft 3 in (1.91 m); GW: 11 in (28 cm); other measurements NMNZ catalog nos. M.125403 & M.125405;[267] holotype of Steenstrupia stockii Kirk, 1882. Kirk specimen No. 3 Kirk (1880:310); Verrill (1881b:398); Kirk (1882:286, pl. 36 figs. 2–4) Verrill (1882d:477); Kirk (1888:34); Pfeffer (1912:34); Suter (1913:1051); Dell (1952:98); Dell (1970:27); Stevens (1980:213, fig. 12.24); Stevens (1988:149, fig. 2); Judd (1996); Paxton (2016a:83); Greshko (2016) Measurements taken by T.W. Kirk. Has been called the "largest specimen recorded in the scientific literature" based on erroneous total length of "approximately 20 m",[268] itself based on claim by Roper & Boss (1982:104) relating to unspecified specimen "stranded on a beach in New Zealand in 1880 [sic]". Considered by Paxton (2016a:83) as the longest reliably measured mantle length of any giant squid specimen (less reliably that of #104), but measurement considered dubious by experts due to wide discrepancy with reported gladius length.[94][nb 23]
48 1879 off Nova Scotia, Canada (42°49′N 62°57′W / 42.817°N 62.950°W / 42.817; -62.950 (Giant squid specimen from lancetfish stomach, 1879))
{NWA}
From fish stomach, Alepidosaurus [sic] ferox ?Architeuthis megaptera Verrill, 1878; ?Architeuthis harveyi (Kent, 1874) Terminal part of tentacular arm Portion of arm 18 in (46 cm) long NMNH catalog no. 576962. Verrill specimen No. 20 ("Banquereau specimen" [after Banquereau Bank, a bank off Nova Scotia]) Verrill (1880a:193); Verrill (1880b:287); Verrill (1882c:16) Frost (1934:103) Lancetfish taken by Capt. J.W. Collins of schooner Marion on halibut trawl-line.
49
(📷)
September 1879 Olafsfjord, Iceland
{NEA}
Architeuthis Left tentacle TL: 7680+ mm; CL: 1010 mm; CSC: 268; TSC: 290; additional indices and counts ZMUC [specimen NA-7 of Roeleveld (2002)] Roeleveld (2002:727) Tentacle morphology examined by Roeleveld (2002).
50 October 1879 near Brigus, Conception Bay, Newfoundland
{NWA}
Found washed ashore Two arms with other mutilated parts Undetermined AL: 8 ft (2.4 m) None?; Verrill specimen No. 22 ("Brigus specimen") Verrill (1880a:194); Verrill (1880b:287); Verrill (1882c:17) Frost (1934:103) Found after storm. Information provided by Moses Harvey.
51 1 November 1879 James's Cove, Bonavista Bay, Newfoundland
{NWA}
Found at surface, alive Entire None; cut up by fishermen EL: 38 ft (12 m); BL: 9 ft (2.7 m); BC: ≈6 ft (1.8 m); TL: 29 ft (8.8 m) None; Verrill specimen No. 23 ("James's Cove specimen") Verrill (1880a:194); Verrill (1880b:287); Verrill (1882c:17) Morning Chronicle of St. John's 9 December 1879; Frost (1934:103) Found alive and driven ashore.
52 Unknown (reported 1880) near Boulder Bank, Nelson, New Zealand
{SWP}
Not stated; hook and line? Not indicated Undetermined 8 ft (2.4 m) long None?; Kirk specimen No. 4 Kirk (1880:310); Verrill (1881b:398) Newspaper article Caught by fishing party. No other data.
53 Unknown (reported 1880) near Flat Point, east coast, New Zealand
{SWP}
Not stated Not indicated Undetermined None None?; Kirk specimen No. 5 Kirk (1880:310); Verrill (1881b:398) Description sent to Mr. Beetham, M.H.R., by Mr. Moore Found by Mr. Moore. No other data.
54
(📷)
April 1880 Grand Banks, Newfoundland
{NWA}
Found dead at surface Architeuthis harveyi (Kent, 1874) Head, tentacles, and arms only Head, tentacles, and arms TL: 66 in (170 cm); ASC: 330; extensive measurements and counts YPM catalog no. 12600y. Verrill specimen No. 24 ("Grand Banks specimen" [2nd]) Verrill (1881b:259, pl. 26 figs. 1–4, pl. 38 figs. 3–7); Verrill (1882c:18, pl. 4 figs. 2–2a, pl. 5 figs. 6–8, pl. 6) Pfeffer (1912:19); Frost (1934:103) Found dead by Capt. O.A. Whitten of schooner Wm.H. Oakes. Arm and sucker regeneration documented by
#549
).
55
(📷)
6 June 1880
{SWP}
Found washed ashore Architeuthis verrilli Kirk, 1882 Entire Not specified ML: 7 ft 6 in (2.29 m); BC: 9 ft 2 in (2.79 m); TL: 25 ft (7.6 m); AL(I, II, IV): 9 ft (2.7 m); AC(I, II, IV): 15 in (38 cm); AL(III): 10 ft 5 in (3.18 m); AC(III): 21 in (53 cm); ASC(III): 71; HC: 4 ft 3 in (1.30 m); HL: 19 in (48 cm); FL: 30 in (76 cm); FW: 28 in (71 cm); EyD: 5 in (13 cm) by 4 in (10 cm) NMNZ; holotype of Architeuthis verrilli Kirk, 1882; specimen no longer extant[270] Kirk (1882:284, pl. 36 fig. 1) Verrill (1882d:477); Kirk (1888:35); Pfeffer (1912:33); Suter (1913:1052); Dell (1952:98); Dell (1970:27) Measurements taken by Kirk, except TL by James McColl. Beak and portions of gladius ("skeleton") taken by Italian fishermen and not recovered.
56 c. 1880 Kvænangen, Norway
{NEA}
Found washed ashore Architeuthis dux Steenstrup, 1857 Entire None None Grieg (1933:19) Sivertsen (1955:11)
57 c. 1880 Tønsvik, Tromsøysund, Norway
{NEA}
Found washed ashore Architeuthis dux Steenstrup, 1857 Entire None None Grieg (1933:19)
58 October 1880 Kilkee, County Clare, Ireland
{NEA}
Found washed ashore "octopus"; Architeuthis sp. O'Brien (1880:585); Ritchie (1918:137) Rees (1950:40); Collins (1998:489) Originally cited as an octopus.
59 first week of November 1881 on beach, Hennesey's Cove, Long Island, Placentia Bay, Newfoundland
{NWA}
Found washed ashore Architeuthis princeps? Entire; "much mutilated by crows and other birds" Not stated "very large"; BL+HL: 26 ft (7.9 m) [estimate] Verrill specimen No. 28 Verrill (1882c:221) M. Harvey letter 19 December 1881 to Verrill Found by Albert Butcher and George Wareham, "who cut a portion from the head", at uninhabited locality; Verrill considered their estimate of the specimen's length "probably too large". .
60
(📷)
10 November 1881
Portugal Cove, near St. John's, Newfoundland

{NWA}
Found floating dead near shore Architeuthis harveyi (Kent, 1874) Entire Entire (somewhat mutilated and poorly preserved) a) BL: 5.5 ft (1.7 m); HL: 1.25 ft (0.38 m); EL: 28 ft (8.5 m); BC: 4.5 ft (1.4 m) b) ML: 4.16 ft (1.27 m); BC: 4 ft (1.2 m); FL: 1.75 ft (0.53 m); FW: 8 in (20 cm) [single]; TL: 15 ft (4.6 m); CL: 2 ft (0.61 m); AL: 4.66 ft (1.42 m) [ventral, minus tip]; TC: 8.5 in (22 cm) [at base]; additional measurements E.M. Worth Museum (101 Bowery, NY, NY). Verrill specimen No. 27 [Anon.] (1881:821, fig.); Verrill (1881b:422); Verrill (1882a:71); Verrill (1882c:201, 219) Morris article in 25 November 1881 New York Herald; Hatton & Harvey (1883:242); Pfeffer (1912:19); Ellis (1997a:34) Obtained by Mr. Morris, photographed by E. Lyons (St. John's), shipped on ice by steamer Catima to New York, purchased and preserved by E.M. Worth. Measurements by a) Inspector Murphy (chief Board of Public Works) when iced; b) Verrill of fixed specimen. An 1881 specimen from Portugal Cove with a "body" reportedly 11 ft (3.4 m) long, mentioned in
Ward's), as Verrill saw the specimen shortly before he began modelling.[78]
61 30 June 1886 Cape Campbell, New Zealand
{SWP}
Found washed ashore Architeuthis kirkii Robson, 1887 Entire Beak and
club
ML: 8 ft 3 in (2.51 m); HL: 1 ft 9 in (0.53 m); AL: 6 ft 6 in (1.98 m); TL: 18 ft 10 in (5.74 m); EL: 28 ft 10 in (8.79 m); BC: ≈8 ft (2.4 m) [estimate] NMNZ catalog nos. M.125404 & ?M.125406;[267] holotype of Architeuthis kirkii Robson, 1887. Kirk specimen No. 2 Kirk (1879:310); Verrill (1881b:398); Robson (1887:156) C.H.[W.] Robson letter 19 June 1879 to T.W. Kirk; Pfeffer (1912:35); Suter (1913:1048); Dell (1952:98); Dell (1970:27) Found by Mr. C.H.[W.] Robson; beak given to Mr. A. Hamilton.
1886 Cupids and Hearts Content (one specimen from each), Newfoundland
{NWA}
Found washed ashore "giant squid" Two specimens; entire? None?; cut up for bait None given Earle (1977:53) Moses Harvey only learned of specimens after their destruction. Information sourced from clippings found in one of Harvey's scrapbooks preserved at Newfoundland Public Archives (PG/A/17).[207]
62
(📷)
"early" October 1887 Lyall Bay, New Zealand
{SWP}
Found washed ashore Architeuthis longimanus Kirk, 1888 Entire Beak and buccal-mass Female EL: 55 ft 2 in (16.81 m); ML: 71 in (180 cm); BC: 63 in (160 cm); extensive additional measurements and description[nb 24]
Dominion Museum [NMNZ];[271] holotype of Architeuthis longimanus Kirk, 1888. Specimen not found[270]
Kirk (1888:35, pls. 7–9); Pfeffer (1912:36) Suter (1913:1049); Dell (1952:98); Dell (1970:27); Wood (1982:191); Ellis (1998a:7, 92); O'Shea & Bolstad (2008); Dery (2013); Paxton (2016a:83) Strangely proportioned animal that has been much commented on; sometimes cited as the longest giant squid specimen ever recorded.[272][nb 24] Considered by Paxton (2016a:83) as candidate for "longest measured" total length of any giant squid specimen (together with #45, and less reliably #209). Found by Mr. Smith, local fisherman. Measurements taken by T.W. Kirk. Date found listed incorrectly in Dell (1952:98).[273]
63 27 August 1888 between Pico and St. George, Azores Islands (38°33′57″N 30°39′30″W / 38.56583°N 30.65833°W / 38.56583; -30.65833 (Giant squid specimen, 27 August 1888)) at 1266 m depth
{NEA}
By benthic trawl Architeuthis? sp.?[274] Large beak Undetermined None Joubin (1895:34)
64 September 1889 Løkberg farm, Mo i Rana, Norway
{NEA}
Found washed ashore Entire None BL: ≈5 ells (3.1 m); TL: 10–12 ells (6.3–7.5 m) [Anon.] (1890:190) Sivertsen (1955:11, fig. 4)
Bergen Museum notified of find by Lorentz Pettersen of Sjona, Helgeland. Failure to secure remains prompted museum to issue notice in June 1890 issue of Naturen seeking specimens in future (which would be first for a Norwegian museum) and offering to cover all associated transportation and packing costs in addition to regular compensation.[275]
1890 Island Cove, Newfoundland
{NWA}
Found washed ashore "giant squid" Entire? None?; cut up for bait? None given Earle (1977:53) Moses Harvey only learned of specimen after its destruction. Information sourced from clippings found in one of Harvey's scrapbooks preserved at Newfoundland Public Archives (PG/A/17).[207]
65 Unknown (reported 1892) Sao Miguel Island, Azores Islands
{NEA}
Found washed ashore Architeuthis princeps Entire?
tentacle club
Beak measurements Museum in Lisbon[276] Girard (1892:214, pls. 1–2) Pfeffer (1912:27); Robson (1933:692)
66 1892 Greenland
{NWA}
Not stated Architeuthis monachus Posselt (1898:279)
[2] Unknown (reported November 1894) Talcahuano, Chile
{SEP}
Unknown; collected and donated to ZMB by
Ludwig Plate
Ommastrephes gigas;
Dosidicus gigas[279]
Entire Entire, internal parts missing, preserved in alcohol; "exceptionally good condition" (Glaubrecht & Salcedo-Vargas, 2004:55) Female (adult) ML: 865 mm; MW: 230 mm; EL: 1740 mm; HL: 160 mm; HW: 190 mm; FL: 440 mm; FW: 600 mm; TL: 720 mm; CL: 225 mm; AL(I): 460 mm; AL(II): 450 mm; AL(III): 500 mm; AL(IV): 440 mm; LSD: 20 mm [tentacle]; LSD: 15 mm [arm II]; LSD: 14 mm [arm II]; EyD: 80 mm; Lens: 35 mm ZMB Moll. 49.804 Martens (1894); Glaubrecht & Salcedo-Vargas (2004:53, figs. 1a–f, 2a–g) Möbius (1898a:373); Möbius (1898b:135); [Anon.] (1899:38); [Anon.] (1902a:41); Kilias (1967:491, fig.); Wechsler (1999) Non-architeuthid. On public display. First noted by
Übersee-Museum Bremen with sperm whale
skull. Re-identified as Dosidicus gigas in June 1998 by Mario Alejandro Salcedo-Vargas. Internal parts apparently removed when specimen originally dissected by Martens or prepared for exhibition (1894–97).
67
(📷)
4 February 1895
Bay of Tateyama [Tokyo Bay], Province of Awa, Japan

{NWP}
In net Architeuthis japonica Pfeffer, 1912 Entire Undetermined Female ML: 720 mm; MW: 235 mm; GL: 640 mm; FL: 280 mm; FW: 200 mm; TL: 2910 mm; extensive additional measurements and description Undetermined; ?Zoological Institute, Science College, Tokyo; holotype of Architeuthis japonica Pfeffer, 1912 Mitsukuri & Ikeda (1895:39, pl. 10); Pfeffer (1912:27) Sasaki (1916:89) Caught in net after 2–3-day storm.
68
(📷)
18 July 1895 near Angra, Azores Islands (38°34'45"N, 29°37'W)
{NEA}
Caught at surface (from sperm whale vomit) using shrimp net Dubioteuthis physeteris Joubin, 1900 [=Architeuthis physeteris (Joubin, 1900)[281]] Mantle only Mantle Male ML: 460 mm; BD: 115 mm; FL: 220 mm; FW: 110 mm; GL: 390 mm MOM [station 588]; holotype of Dubioteuthis physeteris Joubin, 1900[282] Joubin (1900:102, pl. 15 figs. 8–10); Pfeffer (1912:24) Hardy (1956:288); Roper & Young (1972:220); Toll & Hess (1981:753)
[3]
(📷)
18 July 1895 near Angra, Azores Islands (38°34'45"N, 29°37'W)
{NEA}
Caught at surface (from sperm whale vomit) with shrimp net Architeuthis sp.?; non-architeuthid[276] Several jaws Undetermined None Joubin (1900:46, pl. 14 figs. 1–2) Pfeffer (1912:27); Clarke (1956:257) Non-architeuthid.
69
(📷)
10 April 1896
Hevnefjord, Norway

{NEA}
Found washed ashore Architeuthis dux Steenstrup, 1857 Entire Entire Female BL: 2.5 m; AL: 2.5 m; TL: 7.25 m VSM Storm (1897:99); Grieg (1933:19) Brinkmann (1916:178); Nordgård (1923:11); Nordgård (1928:71); Sivertsen (1955:11) Model completed in 1954 based on this specimen and #70; restored in 2010.[283]
70
(📷)
27 September 1896 [or 28 September[284]]
Hevnefjord, Norway

{NEA}
Found washed ashore Architeuthis dux Steenstrup, 1857 Entire Entire, posterior part missing Male TL: 1030+ mm; CL: 900 mm; CSC: 294; TSC: >298; LRL: 17.9 mm; URL: 16.2 mm; additional beak measurements, indices, and counts VSM; VSM 110a [specimen NA-18 of Roeleveld (2000) and Roeleveld (2002)] Storm (1897:99, fig. 20); Grieg (1933:19); Roeleveld (2000:185); Roeleveld (2002:727) Brinkmann (1916:178, fig. 2); Nordgård (1923:11); Nordgård (1928:71); Sivertsen (1955:11); Toll & Hess (1981:753) Beak morphometrics studied by Roeleveld (2000). Tentacle morphology examined by Roeleveld (2002). Model completed in 1954 based on this specimen and #69; restored in 2010.[283]
71 Unknown (reported 1898) Iceland
{NEA}
Not stated Architeuthis monachus Not specified Undetermined None Posselt (1898:279) Bardarson (1920:134)

Type specimens

The following table lists the nominal species-level

original combinations
.

Binomial name and author citation Systematic status Type locality Type specimen and type repository
Loligo bouyeri Crosse & Fischer, 1862:138 Architeuthid?[286] Canary Islands? (#18) Unresolved
Architeuthis clarkei Robson, 1933:682, text-figs. 1–7, pl. 1 Undetermined Scarborough Beach, Yorkshire, England (#107) BMNH Holotype 1933.1.30.5 + 1926.3.31.24 (radula and beak)[287]
Architeuthis dux Steenstrup, 1857:183
Nomen tantum
Architeuthis dux Steenstrup in Harting, 1860:11, pl. 1 fig. 1A Valid species[288] 31°N 76°W / 31°N 76°W / 31; -76 (Giant squid specimen, 5 November 1855) (Atlantic Ocean) (#14) ZMUC Holotype[192]
Plectoteuthis grandis Owen, 1881:156, pls. 34–35 Architeuthis sp.[289] Not indicated (#27) BMNH Holotype[290] [not traced by Lipiński et al. (2000)]
Architeuthis halpertius Aldrich, 1980:59 Nomen nudum[nb 25]
Loligo hartingii Verrill, 1875b:86, fig. 28 Valid species; Architeuthis hartingii[293] Not indicated (#16) University of Utrecht as Architeuthis dux, identification by Harting
Megaloteuthis harveyi Kent, 1874a:181 Architeuthis sp. Conception Bay, Newfoundland (#29) YPM Type 12600y[294]
Architeuthis japonica Pfeffer, 1912:27 Undetermined Tokyo Bay, Japan (#67) Undetermined; Holotype [=Mitsukuri & Ikeda (1895:39–50, pl. 10)]
Architeuthis kirkii Robson, 1887:155 Architeuthis stockii (Kirk, 1882)[295] Cape Campbell, New Zealand (#61) NMNZ Holotype M.125404 + ?M.125406[267]
Architeuthis longimanus Kirk, 1888:34, pls. 7–9 Architeuthis stockii (Kirk, 1882)[295] Lyall Bay, New Zealand (#62) NMNZ Holotype; specimen not located[270]
Megateuthis martensii Hilgendorf, 1880:67 Valid species; Architeuthis martensii
Yedo Japan fish market, Japan
(#28) ZMB Moll. 34716 + 38980
Architeuthis megaptera Verrill, 1878:207 Non-architeuthid; Sthenoteuthis pteropus (Steenstrup, 1855) Nova Scotia, Canada (#[1]) NSMC 1870–Z-2
Architeuthis? monachus Steenstrup, 1857:184
Nomen tantum
Architeuthis monachus Steenstrup in Harting, 1860:11 Architeuthis dux Steenstrup, 1857[203] Raabjerg Strand; Northwest coast of Jutland, Denmark[296] (#13) ZMUC Holotype[296]
Architeuthis mouchezi Vélain, 1875:1002 Nomen nudum; see Mouchezis sancti-pauli
Architeuthis nawaji Cadenat, 1935:513 Undetermined Île d'Yeu, Bay of Biscay, France (#110) Unresolved
Dubioteuthis physeteris Joubin, 1900:102, pl. 15 Valid species; Architeuthis physeteris[281] Azores (38°34'45"N 29°37'W); from sperm whale stomach (#68) MOM Holotype [station 588][282]
Architeuthis princeps Verrill, 1875a:22 Nomen nudum
Architeuthis princeps Verrill, 1875b:79, figs. 25–27 Undetermined a) Grand Banks, Newfoundland; b) North Atlantic (sperm whale stomach) (#22 and 26) NMNH? [not found in collections to date]; Syntypes (a) Verrill specimen No. 1, lower beak; b) Verrill specimen No. 10, upper and lower beak)
Dinoteuthis proboscideus More, 1875a:4527 Architeuthis sp.[289] Dingle, County Kerry, Ireland (#3) Unresolved
Mouchezis sancti-pauli Vélain, 1877:81, text-fig. 8 Valid species; Architeuthis sanctipauli on beach, St. Paul Island (38°43′S 77°32′E / 38.717°S 77.533°E / -38.717; 77.533 (Giant squid specimen, 2 November 1874)), South Indian Ocean (#33) MNHN Holotype 3-2-658 and 3-2-659 (tentacular clubs only)[297]
Steenstrupia stockii Kirk, 1882:286, pl. 36 figs. 2–4 Valid species; Architeuthis stockii[295] [Architeuthid per Pfeffer (1912:2)] Cook Strait, New Zealand (#47) NMNZ Holotype M.125405 + M.125403[267]
Architeuthis titan Steenstrup in Verrill, 1875b:84 [in Verrill (1881b:238, footnote)] Nomen nudum
Architeuthis verrilli Kirk, 1882:284, pl. 36 fig. 1 Species dubium[295] (#55) NMNZ Holotype; [see Förch (1998:89)]

Abbreviations

The following abbreviations are used in the List of giant squid table.

Oceanic sectors

Worldwide giant squid distribution based on recovered specimens

Oceanic sectors used in the main table follow Sweeney & Roper (2001): the Atlantic Ocean is divided into sectors at the equator and 30°W, the Pacific Ocean is divided at the equator and 180°, and the Indian Ocean is defined as the range 20°E to 115°E (the Arctic and Southern Oceans are not distinguished). An additional category has been created to accommodate the handful of specimens recorded from the Mediterranean Sea.

  • NEA, Northeast Atlantic Ocean
  • NWA, Northwest Atlantic Ocean
  • SEA, Southeast Atlantic Ocean
  • SWA, Southwest Atlantic Ocean
  • NEP, Northeast Pacific Ocean
  • NWP, Northwest Pacific Ocean
  • SEP, Southeast Pacific Ocean
  • SWP, Southwest Pacific Ocean
  • NIO, Northern Indian Ocean
  • SIO, Southern Indian Ocean
  • MED, Mediterranean Sea

Measurements

Measuring mantle width
Measuring beak dimensions
Taking sucker counts

Abbreviations used for measurements and counts follow Sweeney & Roper (2001) and are based on standardised acronyms in teuthology, primarily those defined by Roper & Voss (1983), with the exception of several found in older references. Following Sweeney & Roper (2001), the somewhat non-standard EL ("entire" length) and WL ("whole" length) are used in place of the more common TL (usually total length; here tentacle length) and SL (usually standard length; here spermatophore length), respectively.

  • AC, arm circumference (AC(I), AC(II), AC(III) and AC(IV) refer to measurements of specific arm pairs)
  • AD, arm diameter (AD(I), AD(II), AD(III) and AD(IV) refer to measurements of specific arm pairs)
  • AF, arm formula
  • AL, arm length (AL(I), AL(II), AL(III) and AL(IV) refer to measurements of specific arm pairs)
  • ASC, arm sucker count
  • ASD, arm sucker diameter
  • BAC, buccal apparatus circumference
  • BAL, buccal apparatus length
  • BC, body circumference (assumed to mean greatest circumference of mantle unless otherwise specified)
  • BD, body diameter (assumed to mean greatest diameter of mantle)
  • BL, body length (usually equivalent to mantle length, as head length is often given separately)
  • CaL, carpus length
  • CL, club length (usually refers to expanded portion at the apex of tentacle)
  • CSC, club sucker count
  • CSD, club sucker diameter (usually largest) [usually equivalent to LSD]
  • CW, club width
  • DC, dactylus club length
  • EC, egg count
  • ED, egg diameter
  • EL, "entire" length (end of tentacle(s), often stretched, to posterior tip of tail; in contrast to WL, measured from end of arms to posterior tip of tail)
  • EyD, eye diameter
  • EyOD, eye orbit diameter
  • FL, fin length
  • FuCL, funnel cartilage length
  • FuCW, funnel cartilage width
  • FuD,
    funnel
    opening diameter
  • FuL, funnel length
  • FW, fin width
  • GiL, gill length
  • GL, gladius (pen) length
  • GW, gladius (pen) width
  • G(W), daily growth rate (%)
  • HC, head circumference
  • HeL, hectocotylus length
  • HL, head length (most often base of arms to edge of mantle)
  • HW, head width
  • LAL, longest arm length
  • LRL, lower rostral length of beak
  • LSD, largest sucker diameter (on tentacle club) [usually equivalent to CSD]
  • MaL, manus length
  • ML, dorsal mantle length (used only where stated as such)
  • MT, mantle thickness
  • MW, maximum mantle width (used only where stated as such)
  • NGL, nidamental gland length
  • PL, penis length
  • RaL, radula length
  • RaW, radula width
  • RL, rachis length
  • RW, rachis width
  • SInc, number of statolith increments
  • SL, spermatophore length
  • SoA, spermatophores on arms
  • SSD, stalk sucker diameter
  • SSL, spermatophore sac length
  • TaL, tail length
  • TC, tentacle circumference (most often of tentacle stalk)
  • TCL, tentacle club length
  • TD, tentacle diameter (most often of tentacle stalk)
  • TL, tentacle length
  • TSC, tentacle sucker count (club and stalk combined)
  • TSD, tentacle sucker diameter (usually largest)
  • URL, upper rostral length of beak
  • VML, ventral mantle length
  • WL, "whole" length (end of arms, often damaged, to posterior tip of tail; in contrast to EL, measured from end of tentacles to posterior tip of tail)
  • WT, weight

Repositories

Institutional acronyms follow Sweeney & Roper (2001) and are primarily those defined by Leviton et al. (1985), Leviton & Gibbs (1988), and Sabaj (2016). Where the acronym is unknown, the full repository name is listed.

Specimen images

The following images relate to pre–20th century giant squid specimens and sightings. The number below each image corresponds to that given in the

date format
is used throughout).

Notes

Though fictional accounts often depict giant squid attacking boats (cf. #29), live animals found at the surface are almost invariably sick or dying, and no injuries resulting from such encounters have ever been documented.[222]
The surface encounter between a sperm whale and a giant squid supposedly witnessed by Frank Thomas Bullen, from his semi-autobiographical travel narrative The Cruise of the Cachalot, first published in 1898[317]
Pennsylvania Grit
of 13 December 1896
Paul Bartsch, longtime curator of molluscs at the National Museum of Natural History, believed that most reports of sea serpents were based on sightings of giant squid, particularly ones holding their tentacles above the water[318] (see also alternative image)
cetacean
in a state of arousal.
  1. ^ Verrill's marginal annotations read as follows: "Architeuthis monachus (No. 5) Logie Bay, N. Foundland about 18 natural size between 18 and 19. The tub is 38 12 inches [98 cm] in diameter and circular. Harvey (?) letter. Some of the suckers are broken off on the short arms. They alternate in two regular rows. On the club of the long arm there is a marginal row of small suckers on each side alternating with the large ones. One sucker gone on this long arm."[2]
  2. soup form on 3 March 1769, during James Cook's first voyage to the Pacific.[17] In 1783, Banks presented at the Royal Society an important early paper on ambergris by the London-based German physician Franz Xavier Schwediawer, in which was recounted an observation made by fishermen some ten years earlier of a sperm whale that had in its mouth an incomplete cephalopod tentacle nearly 27 ft (8.2 m) long.[18] To Schwediawer, this corroborated "the common saying of the fishermen, that the cuttle-fish is the largest fish of the ocean".[18] Schwediawer's footnote would be cited "uncritically" in subsequent works, such as Thomas Beale's influential The Natural History of the Sperm Whale of 1839.[17]
  3. ^ Ellis (1998a:86) described Verrill as someone with "an almost limitless capacity for work", who "began publishing papers on these specimens almost as fast as they came in". The full list of Verrill's publications on the Newfoundland strandings of 1870–1881 is as follows: Verrill 1874a, b, 1875a, b, c, 1876, 1877, 1878, 1880a, b, 1881a, b, 1882a, c.
  4. The Annals and Magazine of Natural History.[27] The giant squid found beached on Île Saint-Paul on 2 November 1874 (#33) was another early specimen to be photographed.[28] Perhaps the earliest of all was the beak of the October 1871 specimen (#22) from the Grand Banks of Newfoundland, whose photograph was taken some time after its discovery but already mentioned in February 1873 by Packard (1873:92)
    .
  5. ^ According to Harvey's entry in the Dictionary of Canadian Biography, written by giant squid expert Frederick Aldrich, "[a]t least ten per cent of his essays deal with [the giant] squid".[31]
  6. ^ Unconfirmed mass appearances of giant squid include the claim by Frederick Aldrich that a "school of 60 has been sighted off the coast of Newfoundland"[39] (cf. #38). Richard Ellis noted that Aldrich never repeated this claim in print, "so it is likely that he learned it was not accurately reported".[40] However, Aldrich had earlier given more details in an interview with The Muse, where he stated that in autumn 1964 a Japanese trawler operating around 100 miles (160 km) off St. John's "went through a school of sixty or seventy of these animals frolicking on the surface of the water".[41] Aldrich also told Clyde Roper that "Grand Banks fishermen have reported seeing hundreds of giant squid bodies floating on the surface".[42]
  7. ^
    Museum of Economic Fish Culture in South Kensington, London.[76] Its appearance was based on official papers and photographs Buckland received through the Colonial Office in December 1873 relating to both the first Portugal Cove specimen (#29) and the Logy Bay specimen (#30).[76] Buckland's cutout has been described as the "first attempt to show the public what a giant squid really looked like".[77]
  8. ^ See giant squid specimen #661, whose tank leaked around 1 tonne of formalin in February 2022, resulting in the month-long closure of the Ibaraki Nature Museum near Tokyo, Japan.
  9. Portugal Cove specimen of 1873 (#29)[87] and is close to that given for the total length of the Architeuthis longimanus holotype, which describer Thomas William Kirk variously cited as 55 ft 2 in (16.81 m) or 57 ft (17.4 m) and which the fisherman who found the specimen gave as 62 ft (18.9 m).[88]
  10. 524
    ).
  11. ^ In particular, the commercially important Humboldt squid (Dosidicus gigas) is sometimes called 'giant squid' (or its Spanish equivalent calamar gigante) and this, together with its large size, has led to confusion with Architeuthis (e.g. misidentification #[2]). In February 2002, the Manhattan-based seafood restaurant Esca offered what was claimed to be Architeuthis, but the ammonia-free flesh turned out to be that of the Humboldt squid.[107]
  12. ^ One record given by Ellis is omitted from the present list: Ellis (1998a:259) lists a specimen supposedly collected "[n]orth of Bahamas" in 1898, citing Steenstrup (1898), but this appears to stem from confusion with the type specimen of Architeuthis dux (#14), collected off the Bahamas in 1855.
  13. Portugal Cove, Newfoundland, said to have been spotted in 1817 by a local minister named Larveige and named after him.[143]
    These records are likewise excluded due to a lack of substantiating evidence.
  14. Historia de Gentibus Septentrionalibus of the same year (later reprinted by authors such as Gessner, Edward Topsell, Ulisse Aldrovandi, and John Jonston);[154] and various mythological and folkloric creatures such as the Greek Scylla,[155] the Japanese Akkorokamui[156] and umibōzu,[157] the Māori Te Wheke-a-Muturangi,[158] and the Caribbean Lusca.[156] Giant squid have even, on occasion, been proposed as an explanation for the Loch Ness Monster,[159] with the obvious problem that all living cephalopods are exclusively marine.[160] Some authors have cautioned against attributing "sea monster" sightings to giant squid and offered alternative explanations.[161]
  15. ^ Early sources variously give the date as 1545,[170] 1546,[171] December 1549,[172] or 1550.[173] Lycosthenes (1557) mentions three "sea monks" supposedly found in 1530, 1546, and 1549, the first in the Rhine and the latter two near Copenhagen (possibly referring to the same specimen). The earliest known source, a German woodcut of the creature dating from 1546[174]—according to which the event happened the same year—places an upper bound on the date, and Paxton & Holland (2005) consider 1546 to be the most likely year.
  16. ^ Mentioned by Steenstrup (1855a:78), presumably referring to Jónsson's encyclopedic Annálar Björns á Skarðsá, covering the history of Iceland from 1400 to 1645, which was eventually published as a dual Icelandic–Latin work in 1774–1775.[178]
  17. Cyclopterus lumpus]. No trace was seen of the head, except the one aperture, or two, which were found behind the tails or at a short distance from them. This very creature was observed by many trustworthy men, and one of the tails of the sea monster was brought home to Thingørekloster for examination."[184]
  18. ^ In a 2013 article about the giant squid, Mark Dery wrote: "contemporary teuthologists dismiss the "attack" as the death throes of a moribund animal, pointing out that virtually all giant squid encountered on the ocean's surface are dead or dying. "There is not a single corroborated story of a [giant] squid attacking a man, a boat, or a submersible", asserts Ellis."[222]
  19. ^ As part of his research for The Kraken, Reed travelled to Newfoundland and spoke to Picot's descendants and to Margueritte Aldrich, widow of giant squid expert Frederick Aldrich.[230]
  20. Thimble Tickle specimen" is sometimes mistakenly cited as 57 ft (17.4 m).[116] Over time, various other superlative measurements have been attributed to the specimen, including a mass of 2 tonnes[247] or exactly 4,480 lb (2,030 kg);[248] an eye diameter of 40 cm,[247] 18 in (46 cm),[249] or 9 in (23 cm);[250] and suckers 4 in (10 cm) across.[249] Wildly excessive mass estimates for the specimen have included:[251] 29.25 or 30 short tons (26.5 or 27.2 tonnes);[252] near 24 tonnes;[253] less than 8 tonnes;[254] and 2.8 or "more realistic[ally]" 2 tonnes.[255]
  21. ^ The identity of the two other fishermen in the boat is not given in Moses Harvey's original account,[246] but later sources identify Joseph Martin, the founder of Thimble Tickle (now known as Glovers Harbour), as one of the fishermen involved in the squid's capture.[256] George Marsh and Henry Rowsell—the founders of the nearby settlements of Winter House Cove and Lock's Harbour (Lockesporte), respectively—have also been suggested as participants.[257] The fishermen may have learned of Moses Harvey's interest in the giant squid when the latter visited Notre Dame Bay only a couple of months earlier, in August 1878, as part of a geological survey.[258] A CBC News report broadcast in 2004 features Maurice Martin, great-great-grandson of Joseph Martin, recounting the story of the squid's capture as told to him by his grandfather.[259]
  22. Newfoundland Banks may have a body length of 8 feet [2.4 m] and measure up to 40 feet [12 m] overall."[262]
  23. ^ Paxton countered that this was possibly attributable to shrinkage of the gladius prior to measurement, or to it having been severed in several places beforehand,[94] but Kirk wrote that it was measured "when first extracted" and only later "shr[a]nk considerably".[269]
  24. ^ a b c Kirk (1888:38) provides a table with a detailed breakdown of the specimen's various measurements. There is, however, a discrepancy between the total length of 684 in (17.37 m, or exactly 57 ft) given in the table—which agrees with the individual values of 71 in (1.80 m) for the mantle, 22 in (0.56 m) for the head, and 591 in (15.01 m) for the tentacles—and the total length of 55 ft 2 in (16.81 m) given by Kirk in the body of the article. Wood (1982:191) suggested that, due to the tentacles' highly retractile nature, the total length of 62 feet (18.9 m) originally reported by the fisherman "may have been correct at the time he found the squid", and that "[t]his probably also explains the discrepancy in Kirk's figures". Owing to its small mantle size, Wood (1982:191) estimated that "this specimen probably weighed less than 300 lb [140 kg]". O'Shea & Bolstad (2008) opined that the reported total length of 55 ft 2 in (16.81 m) "simply cannot be correct" and attributed it to either "imagination" or artificial lengthening of the tentacles. They added that a female giant squid with a mantle length of 71 in (180 cm) "measured post mortem and relaxed (by modern standards) today would have a total length of ≈32 feet [9.8 m]". Paxton (2016a:86) wrote that this specimen "clearly has the largest ratio of TL to ML [total length to mantle length] ever known in Architeuthis [...] which led [O'Shea & Bolstad, 2008] to suggest that the length was paced out and/or there was extensive post-mortem stretching. However, a re-reading of the original paper suggests that the specimen, although initially paced out, was actually measured, nevertheless the TL is at the edge of the 99.9% prediction interval range [...] and so it was certainly an unusual specimen."
  25. folklorist Herbert Halpert).[137] Aldrich restated this belief in "Monsters of the Deep", the second episode of the 1980 television series Arthur C. Clarke's Mysterious World: "I believe that the giant squid reach an approximate maximum size of something like a hundred and fifty feet [46 m]".[291] Richard Ellis, apparently unaware of Aldrich's similar statements in print, commented: "It is difficult to imagine why Aldrich would have made such an irresponsible statement, unless it had to do with being on camera".[272] Arthur C. Clarke himself wrote in 1992 that "the evidence suggests, they [giant squid] grow up to 150 feet in length".[292] The published literature includes many other such extreme size estimates based only on supposed giant squid suckers or scars thereof
    .

References

Short citations

  1. ^ Frank, 2014:246; Offord, 2016
  2. ^ Aldrich, 1991:459
  3. ^ a b c d Aldrich, 1978; Ellis, 1998a:79, 2005:143–144
  4. ^ Rees, 1949; Salvador & Tomotani, 2014; Hogenboom, 2014
  5. ^ Steenstrup, 1857:183; validated in Harting, 1860:11
  6. ^ Kent, 1874c:116–117; Paxton, 2018:56
  7. ^ Mitchill, 1813, 1815; see also response from Lee, 1819
  8. ^ Smith, 1842a:73, b:85; Earle, 1977:20
  9. ^ Steenstrup, 1849:952/[10]
  10. ^ Schwediawer, 1783:236; Ellis, 1998a:5, 2005:148
  11. ^ Ellis, 1998a:5, 133–134; King, 2019:142
  12. ^ Ellis, 1998a:133
  13. ^ see e.g. Earle, 1977:20–21
  14. ^ Heuvelmans, 1968
  15. ^ de Montfort, 1801; see Freeman, 2017:55
  16. ^ Paxton, 2018:56–57
  17. ^ a b Paxton, 2018:56
  18. ^ a b Schwediawer, 1783:236–237; Paxton, 2018:56
  19. ^ Steenstrup, 1857:184; validated in Harting, 1860:11
  20. ^ Bouyer, 1861
  21. ^ Harvey, 1874a
  22. ^ Lee, 1875:111; Earle, 1977; Muntz, 1995; McConvey, 2015b
  23. ^ Coe, 1929:36; G.E. Verrill, 1958:69
  24. ^ Murray, 1874b:121
  25. ^ a b c Offord, 2016; Keartes, 2016
  26. ^ Aldrich, 1991:458
  27. ^ Harvey, 1874a:68; Verrill, 1875a:34
  28. ^ a b c Wright, 1878:329
  29. ^ Aldrich, 1987:109; Frank, 2014
  30. ^ Hatton & Harvey, 1883:238–243; Aldrich, 1987:115, 1994
  31. ^ Aldrich, 1994
  32. ^ Ellis, 1994a:379, 1998a:257; Sweeney & Roper, 2001:[27]
  33. ^ a b c Lee, 1875:114
  34. ^ Nesis, 2001
  35. ^ a b Sweeney & Young, 2003
  36. ^ Aldrich, 1991:474; Förch, 1998:93; Winkelmann et al., 2013; Guerra et al., 2013
  37. ^ Downer, 1965:8; Aldrich, 1967a, 1968a
  38. ^ Stevens, 1988:158–160
  39. ^ Aldrich, 1967b
  40. ^ Ellis, 1998a:241
  41. ^ Downer, 1965:8
  42. ^ a b c Roper & Shea, 2013:111
  43. ^ Kubodera et al., 2016; see also Sakamoto, 2014
  44. ^ Ellis, 1998a:211
  45. ^ Oreskes, 2003:716, 2014:29
  46. ^ Baird, 2002; Kubodera, 2010
  47. ^ [NHK], 2013a, b; Dery, 2013
  48. ^ Hann, 2006; Guerra et al., 2011:1990; Walker, 2011
  49. ^ Okutani, 2015
  50. ^ Nesis, 2003; Coro et al., 2015
  51. ^ [NMNH], 2004
  52. ^ Guerra et al., 2006; Fernández-Pello García, N.d.
  53. ^ a b Kubodera et al., 2016
  54. ^ O'Shea, 1997, 1999
  55. ^ Roper & Shea, 2013:111; Roper, 2016
  56. ^ Roper et al., 2015
  57. ^ Roper & Jereb, 2010:121; Roper, 2016
  58. ^ see map in Sivertsen, 1955:11
  59. ^ Guerra et al., 2006
  60. ^ Turner, 1963:23
  61. ^ Ellis, 1994a:133
  62. ^ Guerra et al., 2011:1991
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  65. ^ Miller, 1983; Landman & Ellis, 1998; Landman et al., 1999; Roper & Jereb, 2010:123; Ablett, 2012
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  67. ^ [Anon.], 1971
  68. ^ Guerra & Segonzac, 2014:118
  69. ^ [Anon.], 2014a, b
  70. ^ Shimada et al., 2017:9
  71. ^ [NMNH], 2004; [SITES], 2004
  72. ^ [Anon.], 2004b; Miller, 2004
  73. ^ Ellis, 1997a:52
  74. ^ Emerton, 1883; Tratz, 1973; Ellis, 1997a, b, 1998a:214; Wechsler, 1999
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  76. ^ a b Buckland, 1875:211–216, 1886:400; Ellis, 1997a:31
  77. ^ Ellis, 1997a:31
  78. ^ a b Ellis, 1997a:34, 46
  79. ^ a b Ellis, 1997a:35
  80. ^ Emerton, 1883; Ellis, 1997a:35–36
  81. ^ Ellis, 1997a:44–46
  82. ^ [Anon.], 1966; Ellis, 1997a:34–35; [Anon.], 2004a
  83. ^ [Anon.], 1969
  84. ^ Drahl, 2008
  85. ^ [Anon.], 2010
  86. ^ Wood, 1982:188–193; Ellis, 1998a:6–7, b; Paxton, 2016a, b; Bittel, 2016; Romanov et al., 2017
  87. ^ Murray, 1874a:161, 1874b:121
  88. ^ Kirk, 1888:35, 38
  89. ^ Wood, 1982:189, 191; Ellis, 1998a:6–7; O'Shea & Bolstad, 2008; Paxton, 2016a
  90. ^ O'Shea & Bolstad, 2008; Dery, 2013; Roper & Shea, 2013:113; Hanlon & Messenger, 2018:267; see also Nesis, 2001
  91. ^ Cerullo & Roper, 2012:22
  92. ^ a b O'Shea, 2003b
  93. ^ Paxton, 2016a, b; [Anon.], 2016
  94. ^ a b c d e Greshko, 2016
  95. ^ McClain et al., 2015:Table 3
  96. ^ Roper & Jereb, 2010:121
  97. ^ a b O'Shea & Bolstad, 2008
  98. ^ a b Nilsson et al., 2012:683; Land & Nilsson, 2012:86
  99. ^ Robson, 1933:681
  100. ^ Ellis, 1998a:73; Norman, 2000:150; Roper & Jereb, 2010:121
  101. ^ Glaubrecht & Salcedo-Vargas, 2004:62
  102. ^ Roper & Jereb, 2010:123
  103. ^ see Roeleveld, 2002
  104. ^ [UCPH], 2013
  105. ^ see Ellis, 1998a; Salcedo-Vargas, 1999; Glaubrecht & Salcedo-Vargas, 2004
  106. ^ Robson, 1933:681; see e.g. Robson, 1929 by the same author
  107. ^ Ellis, 2005:198
  108. ^ a b Sweeney & Roper, 2001
  109. ^ Ellis, 1994a:379–384, 1998a:257–265
  110. ^ Ellis, 1994a:ix
  111. ^ [Anon.], 1999:109; Roper et al., 2015:78
  112. ^ e.g. Leite et al., 2016; Funaki, 2017; Romanov et al., 2017; Shimada et al., 2017; Guerra et al., 2018
  113. ^ Starkey, 1963; Welfare & Fairley, 1980:74; Bright, 1989:148; Ellis, 1998a:204; Paxton, 2016a:83; dramatised in Packham, 1998
  114. ^ Ellis, 1998a:245; Paxton, 2016a:83
  115. ^ Cousteau & Diolé, 1973:205; Ellis, 1998a:208, 2005:145
  116. ^ a b Paxton, 2016a:83
  117. ^ McDowall, 1998; Ellis, 1998a:248
  118. ^ Revkin, 2013
  119. ^ Hajicek, 2007, 2008; Cassell, 2007
  120. ^ Taylor, 1932:1; [Anon.], 1938a:21; originally reported in East Anglian Daily Times, 31 October 1896; see also Herrington, 1932:60
  121. ^ [Anon.], 1938a:21; [Anon.], 1938b:83; originally reported in East Anglian Daily Times, 22–25 October 1938
  122. ^ see Ellis, 1999
  123. ^ Heuvelmans, 1968:68; Berger, 2009:275
  124. ^ Hoff, 2003:84, 96
  125. ^ Aldrich, 1979:67; Welfare & Fairley, 1980:72, 74; Fitzgerald, 2018
  126. ^ a b Aldrich, 1979:67
  127. ^ Bullen, 1898:139; Ellis, 1998a:136–137, 180–181
  128. ^ Aldrich, 1977
  129. ^ [Anon.], 1988:4; Aldrich, 1990c
  130. ^ Lane, 1957:196; Gantès, 1979:38; Flynn & Weigall, 1980; Welfare & Fairley, 1980:71; Goss, 1985:39; Heuvelmans, 2003:252; Berger, 2009:261, 276; Dyer, 2021
  131. ^ LeBlond & Sibert, 1973:11, 32; Bright, 1989:140; Ellis, 1998a:202
  132. ^ a b Verrill, 1897:79; repeated by Bartsch, 1937:403
  133. ^ Pierce et al., 1995
  134. Chilean Blob); Evon, 2017
  135. ^ Ley, 1959:210
  136. ^ Dozier, 1976:72; Ellis, 1998a:7
  137. ^ a b c Aldrich, 1980:59
  138. ^ de Montfort, 1801
  139. ^ Freeman, 2017:55
  140. ^ a b [Anon.], 1933
  141. ^ Glaubrecht & Salcedo-Vargas, 2004:64
  142. ^ e.g. Labails, 2009:24
  143. ^ Bonnell, 1963:6
  144. ^ Gould, 1886; Gibson, 1887; Aldrich, 1983; Ellis, 1994a, 1998a:10–30
  145. ^ Lee, 1875, 1883; Buel, 1887:81; Bullen, 1898:139; Allan, 1955a, b; Heuvelmans, 1958; Aldrich & Sullivan, 1978; Garcin & Raynal, 2011; Salvador & Tomotani, 2014; Walker, 2016; Naish, 2017
  146. ^ Buckland, 1886:399; Oudemans, 1892; Bartsch, 1917:364, 1931:347, 1937:403; Aldrich & Sullivan, 1978; Aldrich, 1979:67; Ellis, 1994b
  147. ^ Lee, 1883:64; Ellis, 1998a:15; but see Paxton et al., 2005
  148. ^ [Anon.], 1849:264; Myklebust, 1946; Ellis, 1998a:20
  149. ^ Lee, 1883:79; Ellis, 1998a:20; Switek, 2011; but see Galbreath, 2015:42
  150. ^ Carrington, 1957:35
  151. ^ Carrington, 1957:36; Paxton, 2021:314–315
  152. ^ Stead, 1933; Ellis, 1998a:28; Dalton, 2021
  153. ^ Ellis, 1998a:13–14
  154. ^ Ellis, 1998a:12, 14–15
  155. ^ Ley, 1941; Aldrich & Sullivan, 1978; Aldrich, 1990b; Ellis, 1998a:10–11; Lotzof, 2021; Guerra, 2022
  156. ^ a b Lotzof, 2021
  157. ^ Greener, 2010
  158. ^ Kirk, 1880:310; Lotzof, 2021
  159. ^ e.g. [Anon.], 1934
  160. ^ Paxton & Shine, 2016:27, 32
  161. ^ Paxton, 2003; France, 2016a, b, 2017
  162. ^ Lee, 2014; Mikkelson, 2014; Evon, 2016b; see also Ellis, 1994a:163, and April Fools' Day joke of Carstens, 2016, taken seriously by [Anon.], 2020
  163. ^ e.g. Evon, 2016a, 2017; Howard, 2017
  164. ^ Gerhardt, 1966:171; Muntz, 1995; Ellis, 1998a:11–12; Guerra, 2022; see Aristotle, c. 350 BC; Pliny, AD 77–79
  165. ^ [Anon.], 2000
  166. ^ Roper et al., 2015:78
  167. ^ Ellis, 1994a:379
  168. ^ Ellis, 1998a:5, 257; Sweeney & Roper, 2001:[27]; Guerra et al., 2004a:428; but see Paxton & Holland, 2005
  169. ^ Gudger, 1953:199; Lane, 1957:129; Ellis, 1994a:144
  170. ^ Vedel, 1575
  171. ^ Hamer, 1546; Rondelet, 1554; Gessner, 1558
  172. ^ Holberg, 1733
  173. ^ Huitfeldt, [1595]; Stephanius, [c. 1650]
  174. ^ Hamer, 1546
  175. ^ Muus, 1959:170
  176. ^ Carrington, 1957:63
  177. ^ Paxton & Holland, 2005:39
  178. ^ Jónsson, [c. 1645]; see Paxton & Holland, 2005:40
  179. ^ Roeleveld, N.d.
  180. ^ Barber & Riches, 1971:26; Aldrich, 1980:57
  181. ^ Carrington, 1957:63; Russell & Russell, 1975:97
  182. ^ Steenstrup, 1849:950–951/[9]
  183. ^ Packard, 1873:87
  184. ^ Steenstrup, 1849:950–952/[9–10]; for the original Icelandic text, see there and Jónsson, [c. 1645]:238
  185. ^ Steenstrup, 1849:952/[10–11]
  186. ^ Verrill, 1875c:214
  187. ^ a b c Tryon, 1879b:185
  188. ^ Hooke et al., c. 1674
  189. ^ see More, 1875a
  190. ^ [Anon.], c. 1673
  191. ^ see Sueur-Hermel, 2017
  192. ^ a b c Kristensen & Knudsen, 1983:222
  193. ^ Robson, 1933:690
  194. ^ from unpublished manuscript by Steenstrup cited in Verrill, 1875b:84
  195. ^ a b c [NHMD], 2019
  196. ^ Steenstrup
  197. ^ Verrill, 1875b:84 citing Harting, 1860
  198. ^ Tambs-Lyche, 1946:288
  199. ^ Harting, 1860
  200. ^ a b c d e Tryon, 1879b:184
  201. ^ a b c d Verrill, 1880a
  202. ^ a b Dell, 1970:27
  203. ^ a b Stephen, 1962:154
  204. ^ Crosse & Fischer, 1862:138
  205. ^ Dunning, 1998:428
  206. ^ MacAlaster, 1977:14
  207. ^ a b c Earle, 1977:53
  208. ^ Verrill, 1874a:158
  209. ^ a b Sweeney & Roper, 2001:[9]
  210. ^ a b Verrill, 1874a
  211. ^ Verrill, 1875b
  212. ^ J.E. Gray cited in Lee, 1875:114; see also Kent, 1874a:179
  213. ^ Kent, 1874a:178
  214. ^ Verrill, 1881b:401
  215. ^ Pfeffer, 1912:31
  216. ^ Owen, 1881:163
  217. ^ Glaubrecht & Salcedo-Vargas, 2000:276
  218. ^ a b Verrill, 1875a:34
  219. ^ Verrill, 1880a:181
  220. ^ Verrill, 1875b:79
  221. ^ Buckland, 1875:212–214
  222. ^ a b Dery, 2013
  223. ^ Buckland, 1875:213
  224. ^ Maunder, 1991
  225. ^ Hatton & Harvey, 1883:239
  226. ^ Agassiz, 1874
  227. ^ Packham, 1998
  228. ^ a b Duncan, 1906:34; Ellis, 1998a:82
  229. ^ Reed, 1995; Ellis, 1998a:82, 187
  230. ^ Ellis, 1998a:187
  231. ^ Ellis, 1998a:186
  232. ^ [Anon.], 1966; Ellis, 1997a:35
  233. ^ [Anon.], 2004a
  234. ^ Frank, 2014
  235. ^ Ellis, 1998a:201
  236. ^ Aldrich, 1978; Aldrich, 1990a:5
  237. ^ Reed, 1995; Ellis, 1998a:187
  238. ^ Lu et al., 1995
  239. ^ Sweeney & Roper, 2001:[12]
  240. ^ O'Connor, 1875
  241. ^ a b Nunn & Holmes, 2008
  242. ^ a b Pfeffer, 1912:32
  243. ^ a b Lau, 2021
  244. ^ a b [OM], 2021
  245. ^ Copedo, 2022
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  247. ^ a b Ganeri, 1990:24; [Anon.], c. 2001
  248. ^ a b McWhirter & McWhirter, 1971:88
  249. ^ a b [Anon.], 1968
  250. ^ McWhirter & McWhirter, 1968:59, 1971:88
  251. ^ see Wood, 1982:190; Ellis, 1998a:107
  252. ^ MacGinitie & MacGinitie, 1949
  253. ^ Heuvelmans, 1958
  254. ^ MacGinitie & MacGinitie, 1968
  255. ^ Akimushkin, 1963; Wood, 1982:190
  256. ^ [Anon.], c. 2001; Harvey, 2004; Martin & Benoit, 2013; Marsh, 2016
  257. ^ Marsh, 2016
  258. ^ [Anon.], c. 2001
  259. ^ Harvey, 2004
  260. ^ McWhirter & McWhirter, 1968:59
  261. ^ e.g. Wood, 1982:189; Carwardine, 1995:240; Glenday, 2014:62; but see #121 for an exception
  262. ^ McWhirter & Page, 1956:101
  263. ^ Hickey, 2009
  264. ^ Hickey, 2010; [Anon.], N.d.
  265. ^ McConvey, 2015a
  266. ^ Verrill, 1882d:477
  267. ^ a b c d Marshall, 1996:45
  268. ^ Stevens, 1980:213, 1988:149–150
  269. ^ Kirk, 1880:312
  270. ^ a b c Marshall, 1996:46
  271. ^ see Dell, 1970:28
  272. ^ a b Ellis, 1998a:7
  273. ^ Sweeney & Roper, 2001:[87]
  274. ^ Joubin, 1895:34
  275. ^ [Anon.], 1890:190
  276. ^ a b Pfeffer, 1912:27
  277. ^ Martens, 1894
  278. ^ Kilias, 1967:491
  279. ^ Glaubrecht & Salcedo-Vargas, 2004
  280. ^ Kilias, 1993
  281. ^ a b Voss, 1956:136
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  283. ^ a b Eivindsen, 2011
  284. ^ Roeleveld, 2002:727
  285. ^ after Voss, 1998:104; Sweeney & Roper, 2001:[5]; Sweeney & Young, 2003; Roper et al., 2015:82; Sweeney, 2017
  286. ^ Gervais, 1875:93
  287. ^ Lipiński et al., 2000:106
  288. ^ Nesis, 1987:218
  289. ^ a b Pfeffer, 1912:2
  290. ^ Owen, 1881:156
  291. ^ Flynn & Weigall, 1980; Ellis, 1998a:6–7
  292. ^ Clarke, 1992:72
  293. ^ Verrill, 1880a:240
  294. ^ per unpublished notes by S.S. Berry at NMNH, cited by Sweeney, 2017:[194]
  295. ^ a b c d Förch, 1998:89
  296. ^ a b Kristensen & Knudsen, 1983:223
  297. ^ Lu et al., 1995:324
  298. ^ Paxton & Holland, 2005:41
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  300. ^ O'Shea, 2003d
  301. ^ Heuvelmans, 2003:fig. 100
  302. ^ Packard, 1873:94
  303. ^ Johnson & Scott, 1940; Ellis, 1998a:82
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  307. ^ Snell & Tucker, 2003:43
  308. ^ a b Ellis, 1997a:44
  309. ^ Ellis, 1997a:44; see also Snell & Tucker, 2003:97
  310. ^ Ellis, 1998c
  311. ^ Ellis, 1997a:41
  312. ^ Ellis, 1997a:42
  313. ^ Harvey, 2004; [Anon.], N.d.
  314. ^ Hickey, 2009, 2010
  315. ^ see e.g. Wood, 1982:189; Carwardine, 1995:240; Glenday, 2014:62
  316. ^ Eivindsen, 2011; Lervik, 2011
  317. ^ Bullen, 1898:ill. facing p. 143
  318. ^ Bartsch, 1917:364–366, 1931:347–349, 1937:403

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  • Imperadore, P. & G. Fiorito (2018). Cephalopod tissue regeneration: consolidating over a century of knowledge.

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