List of giant squid specimens and sightings
This list of giant squid specimens and sightings is a comprehensive timeline of recorded human encounters with members of the genus Architeuthis, popularly known as giant squid. It includes animals that were caught by fishermen, found washed ashore, recovered (in whole or in part) from sperm whales and other predatory species, as well as those reliably sighted at sea. The list also covers specimens incorrectly assigned to the genus Architeuthis in original descriptions or later publications.
Background
History of discovery
Tales of giant squid have been common among mariners since ancient times, but the animals were long considered
The giant squid's existence was established beyond doubt only in the 1870s, with the appearance of an extraordinary number of complete specimens—both dead and alive—in Newfoundland waters (beginning with #21).[22] These were meticulously documented in a series of papers by Yale zoologist Addison Emery Verrill.[23][nb 3] Two of these Newfoundland specimens, both from 1873, were particularly significant as they were among the earliest to be photographed: first a single severed tentacle—hacked off a live animal as it "attacked" a fishing boat (#29)[24]—and weeks later an intact animal in two parts (#30).[nb 4] The head and limbs of this latter specimen were famously shown draped over the sponge bath of Moses Harvey, a local clergyman, essayist, and amateur naturalist.[29] Harvey secured and reported widely on both of these important specimens—as well as numerous others (most notably the Catalina specimen of 1877; #42)—and it was largely through his efforts that giant squid became known to North American and British zoologists.[30][nb 5] Recognition of Architeuthis as a real animal led to the reappraisal of earlier reports of gigantic tentacled sea creatures, with some of these subsequently being accepted as records of giant squid, the earliest stretching back to at least the 17th century.[32]
"I confess that until I saw and measured this enormous limb, I doubted the accuracy of some early observations which this specimen alone would suffice to prove worthy of confidence. The existence of gigantic cephalopods is no longer an open question. I, now, more than ever, appreciate the value of the adage: 'Truth is stranger than fiction.'"
—
For a time in the late 19th century, almost every major specimen of which material was saved was described as a new species.[34] In all, some twenty species names were coined.[35] However, there is no widely agreed basis for distinguishing between the named species, and both morphological and genetic data point to the existence of a single, globally distributed species, which according to the principle of priority must be known by the earliest available name: Architeuthis dux.[36]
It is not known why giant squid become stranded on shore, but it may be because the distribution of deep, cold water where they live is temporarily altered. Marine biologist and Architeuthis specialist
Though the total number of recorded giant squid specimens now runs
Distribution patterns
The genus Architeuthis has a
The vast majority of specimens are of oceanic origin, including
Total number of specimens
According to Guerra et al. (2006), 592 confirmed giant squid specimens were known as of the end of 2004. Of these, 306 came from the Atlantic Ocean, 264 from the Pacific Ocean, 20 from the Indian Ocean, and 2 from the Mediterranean Sea. The figures for specimens collected in the Atlantic and Pacific Oceans further broke down as follows: 148 in the northeastern Atlantic, 126 in the northwestern Atlantic, 26 in the southeastern Atlantic, 6 in the southwestern Atlantic, 43 in the northeastern Pacific, 28 in the northwestern Pacific, 10 in the southeastern Pacific, and 183 in the southwestern Pacific.[59]
Region | Number of specimens | % of total | Found stranded or floating (%) | From fishing (%) | From predators (%) | Method of capture unknown (%) |
---|---|---|---|---|---|---|
NE Atlantic | 152 | 22.5 | 49 | 31 | 15 | 5 |
NW Atlantic | 148 | 21.9 | 61 | 30 | 1 | 8 |
SE Atlantic | 60* | 8.9 | 10 | 60 | 17 | 13 |
SW Atlantic | 6 | 0.9 | 50 | 16 | 1 | 33 |
NE Pacific | 43 | 6.4 | 7 | 56 | 30 | 7 |
NW Pacific | 30* | 4.4 | 30 | 35 | 30 | 5 |
SE Pacific | 10 | 1.5 | 90 | 10 | 0 | 0 |
SW Pacific | 183 | 27.0 | 12 | 41 | 42 | 5 |
Indian Ocean | 33** | 4.8 | 6 | 94 | 0 | 0 |
W Mediterranean | 3 | 0.4 | 100 | 0 | 0 | 0 |
Equatorial/tropical | 9 | 1.3 | 11 | 44 | 45 | 0 |
All regions | 677 | 100.0 |
- * Underestimates according to Guerra et al. (2011)
- ** Includes records from Durban, South Africa
Procurement, preservation, and display
Preserved giant squid specimens are much sought after for both study and display.
A number of fragmentary giant squid remains were displayed as part of "In Search of Giant Squid", a Smithsonian travelling exhibition curated by Clyde Roper that visited a dozen US museums and other educational institutions between September 2004 and August 2009.[71] The exhibition opened its national tour at Yale University's Peabody Museum of Natural History, which has maintained a strong association with the giant squid from the time of the Newfoundland strandings in the 1870s. Preparations for the Peabody exhibition, overseen by site curator Eric Lazo-Wasem, uncovered giant squid material in the museum's collections that was not previously known to be extant, including original specimens from Addison Emery Verrill's time.[72]
"Architeuthis is an elusive creature. Its occasional appearance on various beaches around the world has provided hardly more than a glimpse of its majestic and intimidating appearance, and hauling it out of the water in a trawl does it no justice either. Papier-mâché or fiberglass models have given us a sense of its size and shape, but they have not captured its mystery and vitality. The spirit of Architeuthis may well be uncapturable; at least no museum has even come close to this fabulous creature—the only living animal for which the term sea monster is truly applicable."
—Richard Ellis, from the closing remarks of his 1997 article "The models of Architeuthis"[73]
In the late 19th century, the giant squid's popular appeal and desirability to museums—but scarcity of preserved specimens—spawned a long tradition of "life-sized" models that continues to the present day.
Real giant squid specimens have traditionally been preserved in either solutions of
Reported sizes
Giant squid size—long a subject of both popular debate and academic inquiry
Based on a 40-year data set of more than 50
O'Shea & Bolstad (2008) give a maximum mantle length (ML) of 225 cm (7.38 ft) based on the examination of more than 130 specimens, as well as beaks recovered from sperm whales (which do not exceed the size of those found in the largest complete specimens), though there are recent scientific records of specimens that slightly exceed this size (such as #371, a 240 cm (7.9 ft) ML female captured off Tasmania, Australia; see also #647, with an estimated 2.15–3.06 m ML). Remeslo (2011) and Yukhov (2014:248) give a maximum mantle length of 260 cm (8.5 ft) for females from southern waters. Questionable records of up to 500 cm (16 ft) ML can be found in older literature.[96] Paxton (2016a) accepts a maximum recorded ML of 279 cm (9.15 ft), based on the Lyall Bay specimen (#47) reported by Kirk (1880:312), but this record has been called into question as the gladius of this specimen (which should approximate the mantle length) was said to be only 190 cm (6.2 ft) long.[94]
Including the head and arms but excluding the tentacles (standard length), the species very rarely exceeds 5 m (16 ft) according to O'Shea & Bolstad (2008). Paxton (2016a) considers 9.45 m (31.0 ft) to be the greatest reliably measured SL, based on a specimen (#46) reported by Verrill (1880a:192), and considers specimens of 10 m (33 ft) SL or more to be "very probable", but these conclusions have been criticised by giant squid experts.[94]
The giant squid and the distantly related
Species identifications
"Undoubtedly several imperfectly distinguished forms have been included in the earlier anecdotal records of Architeuthis. Moreover, specimens of Architeuthis (the Giant Squid par excellence), the smaller
Ommatostrepheshave been indiscriminately described as 'Giant Squids.'"
—Guy Coburn Robson, from his description of Architeuthis clarkei, a species he erected in 1933 based on a carcass (#107) that washed ashore in Scarborough, England, earlier that year[99]
The taxonomy of the giant squid genus
The literature on giant squid has been further muddied by the frequent misattribution of various squid specimens to the genus Architeuthis, often based solely on their large size. In the academic literature alone, such misidentifications encompass at least the
List of giant squid
Sourcing and progenitors
The present list generally follows "
Earlier efforts to compile a list of all known giant squid encounters throughout history include those of marine writer and artist Richard Ellis.[109] Ellis's first list, published as an appendix to his 1994 work Monsters of the Sea, was probably the first such compilation to appear in print and was described in the book's table of contents as "the most complete and accurate list of the historical sightings and strandings of Architeuthis ever attempted".[110] Ellis's much-expanded second list, an appendix to his 1998 book The Search for the Giant Squid, comprised 166 entries spanning four and a half centuries, from 1545 to 1996.[111] Records which appear in Ellis's 1998 list but are not found in Sweeney & Roper's 2001 list have a citation to Ellis (1998a)—in the page range 257–265—in the 'Additional references' column of the main table.[nb 12]
In addition to these global specimen lists, a number of regional compilations have been published, including
Scope and inclusion criteria
The list includes records of giant squid (genus
Purported sightings of giant squid lacking both
The earliest surviving records of very large squid date to
All developmental stages from hatchling to mature adult are included. In the literature there is a single anecdotal account of a giant squid "egg case",
Specimens misassigned to the genus Architeuthis in print publications or news reports are included, but are clearly highlighted as misidentifications.
List of specimens
Records are listed chronologically in ascending order and numbered accordingly. This numbering is not meant to be definitive but rather to provide a convenient means of referring to individual records. Specimens incorrectly assigned to the genus Architeuthis are counted separately, their numbers enclosed in square brackets, and are highlighted in pink ( ). Records that cover multiple whole specimens, or remains necessarily originating from multiple individuals (e.g. two lower beaks), have the '
- Date – Date on which the specimen was first captured, found, or observed. Where this is unknown, the date on which the specimen was first reported is listed instead and noted as such. All times are local.
- Location – Area where the specimen was encountered, including coordinates and depth information where available. Given as it appears in the cited reference(s), except where additional information is provided in square brackets. The quadrant of a major ocean in which the specimen was found is given in curly brackets (e.g. {NEA}; see Oceanic sectors).
- Nature of encounter – Circumstances in which the specimen was recovered or observed. Given as they appear in the cited reference(s), although "washed ashore" encompasses all stranded animals.
- Identification – Species- or vernacular namehas been applied to the specimen (e.g. "giant squid" or a non-English equivalent), this is given instead.
- Material cited – Original specimen material that was recovered or observed. "Entire" encompasses all more-or-less complete specimens. Names of anatomical features are retained from original sources (e.g. "jaws" may be given instead of the preferred "beak", or "body" instead of "mantle"). The specimen's state of preservation is also given, where known, and any missing parts enumerated (the tentacles, arm tips, reddish skin and eyes are the parts most often missing in stranded specimens, owing to their delicate nature and/or preferential targeting by scavengers).
- Material saved – Material that was kept after examination and not discarded (if any). Information may be derived from outdated sources; material may no longer be extant even if stated as such.
- Sex – Sex and sexual maturity of the specimen.
- Size and measurements – Data relating to measurements and counts. Abbreviations used are based on standardised acronyms in arithmetic precision and original units preserved (metric conversions are shown alongside imperial measurements), though some of the more extreme lengths and weights found in older literature have since been discredited.
- Repository – Institution in which the specimen material is deposited (based on cited sources; may not be current), including boldface. If an author has given a specimen a unique identifying number (e.g. Verrill specimen No. 28), this is included as well, whether or not the specimen is extant.
- Main references – The most important sources, typically ones that provide extensive data and/or analysis on a particular specimen (often primary sources). Presented in author–date parenthetical referencing style, with page numbers included where applicable (those in square brackets refer either to unpaginated works or English translations of originally non-English works; see Full citations). Only the first page of relevant coverage is given, except where this is discontinuous. Any relevant figures ("figs.") and plates ("pls.") are enumerated.
- Additional references – References of lesser importance or primacy, either because they provide less substantive information on a given record (often secondary sources), or else because they are not easily obtainable or possibly even extant (e.g. old newspaper articles, personal correspondence, and television broadcasts) but nonetheless mentioned in more readily accessible published works (see Full citations).
- Notes – Miscellaneous information, often including persons and vessels involved in the specimen's recovery and subsequent handling, and any dissections, preservation work or scientific analyses carried out on the specimen. Where animals have been recorded while alive this is also noted. Material not referable to the genus Architeuthis, as well as specimens on public display, are both highlighted in bold (as "Non-architeuthid" and "On public display", respectively), though the latter information may no longer be current.
The total number of giant squid records listed across this page and successive lists is 714, though the number of individual animals covered is greater (the additional number exceeding 250) as some records encompass multiple specimens (indicated in grey). Additionally, 13 records relate to specimens misidentified as giant squid (indicated in pink).
# | Date | Location | Nature of encounter | Identification | Material cited | Material saved | Sex | Size and measurements | Repository | Main references | Additional references | Notes |
---|---|---|---|---|---|---|---|---|---|---|---|---|
1 (📷) |
c. 1546[nb 15] | Øresund, near Malmö, Denmark–Norway [since 1658 Malmö has been part of Sweden] {NEA} |
Found washed ashore; "caught live"[175] | "sea monk"; Architeuthis monachus Steenstrup in Harting, 1860; Jenny Haniver made from a skate;[176] Squatina squatina (angelshark)[177] | Entire? | Undetermined | ?WL: ≈3 m | Hamer (1546:[1], fig.); Belon (1553:38, fig.); Rondelet (1554:492, fig.); Belon (1555:32, fig.); Lycosthenes (1557:609, fig.); Gessner (1558:519, fig.); Rondelet (1558:361, fig.); Sluperius (1572:"89", 105, fig.); Vedel (1575); Huitfeldt ([1595]:1545); Gessner (1604:438, fig.); Stephanius ([c. 1650]:344); Steenstrup (1855a:63, 3 figs.); Roeleveld & Knudsen (1980:293, 3 figs.); Ellis (1998a:60, fig.); Paxton & Holland (2005:39, fig. 1) | records of Björn Jónsson á Skarðsá;[nb 16] Scheuchzer (1716:153); Holberg (1733:379); Lönnberg (1891:36); Nordgård (1928:71); Tambs-Lyche (1946:288); Carrington (1957:58, fig.); Muus (1959:170); Russell & Russell (1975:94); Strauss (1975:393, fig.); Aldrich (1980:55); Roeleveld (N.d.) | Contemporaneously regarded as a " sea bishop has also been interpreted as a giant squid carcass[180] or else a Jenny Haniver made from a skate.[181]
| ||
2 | autumn 1639 | {NEA} |
Found washed ashore | Architeuthis sp. | Entire | One arm | BL+HL: ≈6 ft (1.8 m); AL: ≈3 ft (0.91 m); TL: ≈16–18 ft (4.9–5.5 m); BC: ≈3–4 ft (0.91–1.22 m) | Thingøre monastery; "museum at Copenhagen" (ZMUC?)[183] | Jónsson ([c. 1645]:238); Ólafsson (1772:716); Steenstrup (1849:950/[9]); Steenstrup (1898:425/[272]); Ellis (1998a:65) | Packard (1873:87); Verrill (1875b:84); Robson (1933:691); Muus (1959:170); Berger (2009:260) | Original Icelandic account is from the contemporaneous Annálar Björns á Skarðsá and has been translated into English.[nb 17] Crude drawing of animal mentioned by Eggert Ólafsson was lost with most of his books when his boat capsized off Iceland in 1768, leading to his death.[185] Identified by Japetus Steenstrup as decapod cephalopod in 1849. | |
3 (📷) |
c. 15 October 1673 | Dingle-I-cosh, Kerry, Ireland {NEA} |
Found floating at surface, in process of washing ashore, alive | Dinoteuthis proboscideus More, 1875; Architeuthis monachus;[186] Ommastrephes (Architeuthis) monachus[187] | Entire | Two arms, buccal mass, and suckers taken to Dublin | TL: ≈11 ft (3.4 m) + 9 ft (2.7 m); AL: ≈6–8 ft (1.8–2.4 m); "liver": 30 lb (14 kg) | Undetermined [NMI?]; holotype of Dinoteuthis proboscideus More, 1875 | [Anon.] (c. 1673); Hooke et al. (c. 1674:[1], fig.); More (1875a:4526); Verrill (1875c:214); Tryon (1879b:185); Ellis (1998a:66); Sueur-Hermel (2017:64) | More (1875b:4571); Massy (1909:30); Ritchie (1918:137); Robson (1933:692); Rees (1950:40); Hardy (1956:285); Collins (1998:489) | Found by James Steward. Original material relating to this specimen consists of: a broadsheet printed in | |
4 | 1680 | Ulvangen Fjord, Alstadhoug parish, Norway {NEA} |
Not stated | Entire? | Pontoppidan (1753:344) | Steenstrup (1857:184/[18]); Grieg (1933:19) | ||||||
5 | 1770 | Jutland, Denmark {NEA} |
Unknown | Muss (1959) | Ellis (1998a:257) | |||||||
6 | 27 May 1785 | Grand Banks, Newfoundland {NWA} |
Found floating at surface, dead | Architeuthis sp. | BL: 7 ft (2.1 m) | Cartwright (1792:44); Thomas ([1795]:183); Aldrich (1991:457) | Found during George Cartwright's sixth and final voyage to Newfoundland and Labrador. Spotted at 10 am surrounded by birds. Head broke off during retrieval. Described as "a large squid [...] when gutted, the body filled a pork barrel, and the whole of it would have filled a tierce". | |||||
7 | November or December 1790 | Arnarnaesvik, Modruvalle, Iceland {NEA} |
Found washed ashore | Entire | None; used for cod bait | "longest tentacula": >3 fathoms (5.5 m); "body right from the head": 3.5 fathoms (6.4 m); "so thick that a fullgrown man could hardly embrace it with his arms" | Steenstrup (1849:952/[11]); Steenstrup (1898:429/[276]); Ellis (1998a:68) | February 1792 diary of Sveinn Pálsson (in library of Icelandic Literary Society, in Copenhagen); Verrill (1875b:84); Robson (1933:691) | Called Kolkrabbe ('coal-crab') by local people. Identified by Japetus Steenstrup as decapod cephalopod in 1849. | |||
8 | 1700s (reported 1795) | Freshwater Bay, near mouth of St. John's harbour, Newfoundland {NWA} |
Unknown | Architeuthis sp. | Thomas ([1795]:183); Aldrich (1991:457) | |||||||
9 | 1700s | Grand Banks, Newfoundland {NWA} |
Unknown | Architeuthis sp. | Aldrich (1991:457) | |||||||
10 | 1798 | north coast of Denmark {NEA} |
Not stated | "gigantic squid" | Unknown | "museum at Copenhagen" (ZMUC?) | Packard (1873:87) | Ellis (1998a:257) | ||||
11 | 9 January 1802 | off Tasmania, Australia {SWP} |
Found at surface, alive | ?Loligo ["vraisemblablement du genre Calmar [Loligo, Lamarck]"] | "size of a barrel" ["grosseur d'un tonneau"]; AL: 1.9–2.2 m; AD: 18–21 cm | Péron (1807:216) | Quoy & Gaimard (1824:411); Ellis (1998a:257) | Péron (1807:216) wrote: "it rolled with noise in the midst of the waves, and its long arms, stretched out on their surface, stirred like so many enormous reptiles" (translated from the French). | ||||
12 | between 1817 and 1820 | Atlantic Ocean, near equator {?} |
Found floating at surface | "énorme calmar" | Partial remains; "tentacles" ("tentacules") missing | WT: 100 " livres " [estimate]; WT: 200 "livres" [estimate; if complete] |
Quoy & Gaimard (1824:411) | Packard (1873:88); Ellis (1998a:257) | Found at surface in calm weather. Quoy & Gaimard (1824:411) opined: "it is easy to imagine that one of these terrible molluscs could readily remove a man from a fairly large boat, but not a medium-tonnage vessel, still less tilting this vessel and endangering it, as some would like to believe" (translated from the French). | |||
13 (📷) |
December 1853 | {NEA} |
Found washed ashore | Architeuthis monachus | Entire | Jaws only; radula discarded after poor preservation; jaws cut out; portion used for bait; remainder buried after 2 days | WT: 80–85 kg; jaw measurements Steenstrup (1898:423/[270]) | ZMUC catalog no. CEP-133; holotype of Architeuthis monachus Steenstrup, 1857[192] | Steenstrup (1855b:[14]); Harting (1860:11); Steenstrup (1898:415/[258], pl. 1 figs. 1–2); Kristensen & Knudsen (1983:222) | Steenstrup (1857:[18]); Packard (1873:87); Gervais (1875:91); Verrill (1875b:84); Verrill (1880a:238, pl. 25 fig. 3); Verrill (1882c:51, pl. 12 fig. 3); Posselt (1890:144); Nordgård (1928:71) | "Architeuthis monachus" Steenstrup = nomen nudum[193] | |
14 (📷) |
5 November 1855 | western Atlantic Ocean, near Bahamas (31°N 76°W / 31°N 76°W) {NWA} |
Not stated; presumably found floating at surface | Architeuthis dux Steenstrup, 1857; Architeuthis titan[194] | Various parts | Gladius, mouthparts, part of arm, several suckers, and what may be hectocotylus[195] | Male | WL: 377 cm; AL: 1/2 whole length;[196] beak measurements; GL: 6 ft (1.8 m)[197] | ZMUC catalog no. CEP-97 (or CEP-000097) and NHMD-77320 (multiple parts, each in its own glass vessel: gladius, mouthparts, part of arm, several suckers, and what may be hectocotylus); Bergen Museum[198] |
Steenstrup (1857:[18]); Steenstrup (1882:[160]); Steenstrup (1898:413, 450/[260, 298], pls. 3–4); Tryon (1879b:186, pl. 86 fig. 388); Kristensen & Knudsen (1983:222); Glaubrecht & Salcedo-Vargas (2000:273); [NHMD] (2019) | Packard (1873:87); Verrill (1875b:84); Posselt (1890:144); Toll & Hess (1981:753) | Obtained by Capt. Vilhelm Hygom. Museum für Naturkunde in Berlin, which was incorporated into collection in 1883 according to catalogue entry. Preserved in 70% ethanol.[195]
|
15 | December 1855 | Aalbaekbugten, Denmark {NEA} |
Found washed ashore | Architeuthis sp. | Entire? | Undetermined | None | Muus (1959:170) | Posselt (1890:144) | |||
16 (📷) |
Unknown (reported 1860) | Unknown {?} |
Not stated | Architeuthis dux;[199] ?Ommastrephes hartingii;[200] Architeuthis hartingii (Verrill, 1875);[201] nomen nudum[202] | Jaws, buccal mass, detached arm suckers | Jaws, buccal mass, detached arm suckers | ASD: 1.05 in (2.7 cm) | Utrecht University Natural History Museum; holotype of Loligo hartingii Verrill, 1875. Harting specimen No. 1 | Harting (1860:2, pl. 1); Kent (1874d:491); Verrill (1875b:85, fig. 28); Tryon (1879b:149, 184, pl. 60 figs. 194–195); Verrill (1880a:240, pl. 16 fig. 8, pl. 25 fig. 1); Verrill (1882c:52, pl. 12 figs. 1–1c); Pfeffer (1912:37) | Dell (1970:27) | ||
17 | 1860 or 1861 | between Hillswick and Scalloway, Shetland, Scotland {NEA} |
Found washed ashore | Architeuthis monachus Steenstrup, 1857; Architeuthis dux Steenstrup, 1857[203] | Undetermined | TL: 16 ft (4.9 m); AL: ≈8 ft (2.4 m); BL: ≈7 ft (2.1 m) | Jeffreys (1869:124); Stephen (1944:263) | More (1875b:4571); Pfeffer (1912:26); Rees (1950:40); Collins (1998:489) | ||||
18 (📷) |
30 November ?1861 [=1860 Rees & Maul] | about 20 miles (32 km) northeast of Teneriffe, Canary Islands {NEA} |
Found floating at surface | Loligo bouyeri;[204] ?Ommastrephes bouyeri[200] | Entire, decomposed | None | BL: 15–18 ft (4.6–5.5 m) | None | Bouyer (1861:1263); Crosse & Fischer (1862:135); Bouyer (1866:275, fig.); Kent (1874a:180); Verrill (1875b:86); Tryon (1879b:149, 184, pl. 59); Bourée (1912:113, fig. 108); Aldrich (1978:2); Ellis (1998a:5, 78); Heuvelmans (2003:185, figs. 95–96, 100) | Frédol (1865:314, pl. 13); Figuier (1866:464, fig. 362); Frédol (1866:362); Mangin (1868:321); Meunier (1871:245); Kent (1874d:491); Gervais (1875:93); Lee (1883:38, fig. 8); Rees & Maul (1956:266); Carrington (1957:53, pl. 3b); Muntz (1995:19, fig. 11); Lagrange (2009:19) | Observed only by officers of the French gunboat . | |
19 | 1862 | North Atlantic {NEA/NWA} |
Unknown | Crosse & Fischer (1862) | Ellis (1998a:258) | |||||||
[1] (📷) |
Unknown; 1870? | Cape Sable, Nova Scotia, Canada {NWA} |
Found washed ashore | Architeuthis megaptera Verrill, 1878 [=Sthenoteuthis pteropus (Steenstrup, 1855)][205] | Entire | Entire | BL: 14 in (36 cm); BL+HL: 19 in (48 cm); EL: 43 in (110 cm); TL: 22–24 in (56–61 cm); AL: 6.5–8.5 in (17–22 cm); FW: 13.5 in (34 cm); FL: 6 in (15 cm); extensive additional measurements | NSMC catalog no. 1870-Z-2; YPM catalog nos. IZ 017932 Archived 3 August 2019 at the Wayback Machine (sucker) & IZ 017713; holotype of Architeuthis megaptera Verrill, 1878;[206] Verrill specimen No. 21 ("Cape Sable specimen") | Verrill (1878:207); Tryon (1879b:187); Verrill (1880a:193, pl. 21); Verrill (1882c:17, pl. 16 figs. 1–9) | Non-architeuthid. Collected by J.M. Jones. | ||
20 | September 1870 | Waimarama, east coast of Wellington, New Zealand {SWP} |
Found washed ashore | Entire | Beak | BL+HL: 10 ft 5 in (3.18 m); BC: 6 ft (1.8 m); AL: 5 ft 6 in (1.68 m) | In Kirk's possession; Kirk specimen No. 1 | Kirk (1880:310); Verrill (1881b:398) | Meinertzhagen letter 27 June 1879 to Kirk; Pfeffer (1912:32); Dell (1952:98) | Mr. Meinertzhagen sent beak, saved by third party (unidentified), to Kirk. Natives called specimen a "taniwha". | ||
21 | 1870 (winter) | Lamaline, Newfoundland {NWA} |
Found washed ashore | Architeuthis monachus of Steenstrup | Two specimens; entire? | None?; used as fish bait[207] | Two; EL: 40 ft (12 m) and EL: 47 ft (14 m) | None?; Verrill specimen Nos. 8 & 9 ("Lamaline specimens") | Murray (1874a:162); Verrill (1875a:36); Verrill (1880a:187); Verrill (1882c:11) | Boston Traveller, November 1873; Harvey (1874a:69); Kent (1874a:182); Frost (1934:101); Earle (1977:52) |
Data from Mr. Harvey letter citing Rev. M. Gabriel's statement to Harvey. | |
22 (📷) |
October 1871 | Grand Banks, Newfoundland {NWA} |
Found floating at surface | Architeuthis princeps Verrill, 1875 | Entire; part used as bait | Jaws obtained from Baird for examination by Verrill | BL: ≈15 ft (4.6 m); BD: 19 in (48 cm); AL: ≈10 ft (3.0 m) [mutilated]; AD: 7 in (18 cm); AC: 22 in (56 cm); beak; BC: 4 ft 8 in (1.42 m); WT: 2,000 lb (910 kg) | Jaws at NMNH[208] (no longer extant?[209]); lower jaw is syntype of Architeuthis princeps Verrill, 1875b; Verrill specimen No. 1 ("Grand Banks specimen" [1st]) | Packard (1873:91); Verrill (1874a:158); Verrill (1874b:167); Verrill (1875b:79, fig. 27); Verrill (1880a:181, 210, pl. 18 fig. 3); Verrill (1882c:5, pl. 11 figs. 3–3a) | Pfeffer (1912:20); Frost (1934:100) | Taken by Capt. Campbell, Schooner B.D. Haskins. | |
23 | 1871 | Wellington, New Zealand {SWP} |
?EL: 16 ft (4.9 m) | Dell (1952) | Ellis (1998a:258) | |||||||
24 | 1872 (autumn or winter) | Coomb's Cove, Fortune Bay, Newfoundland {NWA} |
Found alive in shallow water, having been driven ashore in heavy sea | Entire; "one long arm missing" (later changed to both present) | BL: 10 ft (3.0 m); BD: 3–4 ft (0.91–1.22 m); TL: 42 ft (13 m); AL: ≈6 ft (1.8 m); AD: 9 in (23 cm); skin + flesh: 2.25 in (5.7 cm) thick; EL: 52 ft (16 m) | Unknown; Verrill specimen No. 3 ("Coombs' Cove specimen") | Verrill (1874a:159); Verrill (1874b:167); Verrill (1875a:35); Verrill (1880a:183); Verrill (1882c:7) | Owen (1881:163); Frost (1934:101) | Specimen had a reddish colour. Verrill's data taken from newspaper accounts and 15/VI/1873 T.R. Bennett letter to Prof. Baird. Verrill (1880a:186) states his No. 6 is same specimen as No. 3; this cannot be correct, since capture date for No. 6 is clearly stated as December 1874 by Verrill (1875c:213).[209] | |||
25 (📷) |
December 1872 | Bonavista Bay, Newfoundland {NWA} |
Found washed ashore | ?Architeuthis dux;[210] ?Architeuthis harveyi[201] | Entire (damaged arms) | Pair of jaws and two suckers | TL: 32 ft (9.8 m); AL: ≈10 ft (3.0 m); BL: ≈14 ft (4.3 m) [estimate]; BC: 6 ft (1.8 m) | NMNH; YPM catalog no. IZ 034835. Verrill specimen No. 4 ("Bonavista Bay specimen") (1875a:33); and possibly also Verrill specimen No. 11 ("Second Bonavista Bay specimen") (1875b:79) | Verrill (1874a:160); Verrill (1874b:167); Verrill (1875a:33, fig. 11); Verrill (1875b:79); Verrill (1880a:184, 187, pl. 16 figs. 5–6, pl. 25 fig. 5); Verrill (1882c:8, 11, pl. 3 figs. 4–4a, pl. 4 figs. 1–1a) | Pfeffer (1912:19); Frost (1934:101) | Material from Rev. A. Munn, through Prof. Baird to Verrill. | |
26 (📷) |
Unknown (reported 1873) | North Atlantic Ocean {NWA} |
From sperm whale stomach | Architeuthis princeps Verrill, 1875; Ommastrephes (Architeuthis) princeps[187] | Upper and lower jaws | Upper and lower jaws | Beak measurements | Presented by Capt. N.E. Atwood of Provincetown, Massachusetts to EI;[211] PASS;[201] syntype of Architeuthis princeps Verrill, 1875b; Verrill specimen No. 10 ("Sperm-whale specimen") | Packard (1873:91, fig. 10); Verrill (1875a:22); Verrill (1875b:79, figs. 25–26); Tryon (1879b:185, pl. 85); Verrill (1880a:187, 210, pl. 18 figs. 1–2); Verrill (1882c:11, pl. 11 figs. 1–2) | Frost (1934:101) | First reported by Alpheus Spring Packard in February 1873. Verrill states Packard's illustration is inaccurate. | |
27 (📷) |
Unknown (reported 1873) | Unknown; possibly east coast of South America[212] {SWA}? |
Not stated | Architeuthis monachus;[213] Plectoteuthis grandis Owen, 1881; Architeuthis sp.? (grandis);[214] nomen nudum[202] | Sessile arm | Arm | AL: 9 ft (2.7 m); AC: 11 in (28 cm); ASD: ≤0.5 in (1.3 cm); total size and size of various missing parts estimated by Lee (1875:114) | BMNH; holotype of Plectoteuthis grandis Owen, 1881 | Kent (1874a:179); Kent (1874d:493); Lee (1875:113); Verrill (1875b:86); Owen (1881:156, pls. 34–35); Verrill (1881b:400); Verrill (1882b:72); Steenstrup (1882:[160]); Pfeffer (1912:37) | Dell (1970:27) | "No history relating to it has been preserved", but first examined by Henry Lee in May 1873, having been in BMNH collections for "long" time.[33] Bore c. 300 suckers. | |
28 | 1873 | Yedo [Tokyo] fishmarket, Japan {NWP} |
Purchased | Megateuthis martensii Hilgendorf, 1880; Nomen spurium[215] | 'Entire', missing head, "abdominal sac", ends of tentacles and arms[216] | Not specified | ML: 186 cm; WL: 414 cm; HL: 41 cm; AL: 197 cm [longest]; ASD: 1.5 cm (with 37 cusps); EyD: 200 mm | ZMB Moll. 34716 + 38980; holotype of Megateuthis martensii Hilgendorf, 1880 [34716a: eyeball, 200 mm diameter, dry; 34716b: pieces of arm and gladius, suckers; 34716c: larger piece of arm with suckers; 38980: four suckers from holotype arm piece] | Hilgendorf (1880:67); Pfeffer (1912:31); Sasaki (1929:227); Glaubrecht & Salcedo-Vargas (2000:276) | Owen (1881:163); Sasaki (1916:90) | Second specimen from Tokyo fishmarket seen by Franz Martin Hilgendorf and used for description of gladius. Of other specimen, Hilgendorf saved "parts of an arm, the covering of the eye, and a fragment of the gladius" ("Theile eines Armes, die Hüllen des Auges, und ein Bruchstück des Schulpes").[217] Model of specimen placed in Exhibition of Fishery in Berlin. | |
29 (📷) |
26 October 1873 | off Portugal Cove, Conception Bay, Newfoundland {NWA} |
Found floating at surface, alive | Megaloteuthis harveyi Kent, 1874; Architeuthis monachus of Steenstrup;[218] ?Architeuthis harveyi[219] | Entire | One tentacle; one arm discarded | (see | YPM?; holotype of Megaloteuthis harveyi Kent, 1874; Verrill specimen No. 2 ("Conception Bay specimen") | Harvey (1873a); Harvey (1873b); Harvey (1873c); Harvey (1874a:67, fig.); Murray (1874a:161); Murray (1874b:120); Verrill (1874a:159); Verrill (1874b:167); Kent (1874a:178, 182); Agassiz (1874:226); Kent (1874d:32); Buckland (1875:211); Verrill (1875a:34); Verrill (1875b:78); Verrill (1880a:181); Verrill (1881b:pl. 26 fig. 5); Verrill (1882b:74); Verrill (1882c:5, pl. 4 figs. 3–3a); Hatton & Harvey (1883:238); Harvey (1899:732, fig.); Ellis (1998a:81); Haslam (2017) | "13 December Field"; [Anon.] (1873:2); Harvey (1873d:2); [Anon.] (1874:333); de La Blanchère (1874:197, fig.); Rathbun (1881:266, fig.); Owen (1881:161, pl. 33 fig. 2); Lee (1883:42, fig. 9); [Anon.] (1902b:6, fig.); Pfeffer (1912:19); Frost (1934:100); Aldrich (1991:457); Packham (1998); Dery (2013) | Struck by Theophilus Picot from boat whereupon it "attacked" the boat; veracity of account has been questioned. E. Annie Proulx.[231]
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30 (📷) |
25 November? 1873 | Logy Bay (≈3 miles from St. John's), Newfoundland {NWA} |
In herring net | ?Architeuthis monachus of Steenstrup;[210] Ommastrephes (Architeuthis) monachus;[200] Architeuthis harveyi (Kent, 1874)[201] | Entire (badly mutilated, head severed, eyes missing, etc.) | Miscellaneous parts obtained from Rev. M. Harvey (gladius and ?) | (see club description; extensive description of reconstructed parts |
YPM catalog nos. IZ 009634 (beak and limbs), IZ 017924 (radula), IZ 017925, IZ 017926 & IZ 034968[permanent dead link]. Verrill specimen No. 5 ("Logie Bay specimen") | Harvey (1873d:2); Verrill (1874a:160); Verrill (1874b:167); Kent (1874a:181); Kent (1874d:32); Buckland (1875:212, 214); Verrill (1875a:22, figs. 1–6, 10); Verrill (1876:236); Tryon (1879b:184, pls. 83–84); Verrill (1880a:184, 197, pls. 13–15, pl. 16 figs. 1–4, pl. 16a); Verrill (1880b:295, pl. 13); Verrill (1882c:8, pls. 1–2, pl. 3 figs. 1–3, pl. 4 figs. 4–11, pl. 5 figs. 1–5); Hatton & Harvey (1883:240); Harvey (1899:735, fig.); Pfeffer (1912:18); Aldrich (1991:457, fig. 1A, B); Haslam (2017) | Harvey in Morning Chronicle (newspaper) of St. John's; Maritime Monthly Magazine of St. John's, March 1874; several other newspapers; [Anon.] (1874:332); Lee (1883:43, fig. 10); [Anon.] (1902b:6, fig.); Frost (1934:101) | Verrill's data from letter to Dr. Dawson from shower curtain rod were subject of Preparing the Ghost (2014), a work of creative nonfiction by Matthew Gavin Frank.[234]
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31 | 1874 | Foldenfjord, Norway {NEA} |
Found washed ashore | Architeuthis dux | Entire | None | WL: ≈4 m | Grieg (1933:19) | Nordgård (1928:71) | |||
32 | 10 May 1874 | off Trincomalee, Sri Lanka (8°50′N 84°05′E / 8.833°N 84.083°E) {NIO} |
Reportedly seen sinking ship | Unknown | The Times, 4 July 1874; Mystic Press, 31 July 1874; Lane (1957:205); Flynn & Weigall (1980); Ellis (1998a:198); Boyle (1999); Uragoda (2005:97) | Welfare & Fairley (1980:74); Aldrich (1990a:5); Clarke (1992:72); Ellis (1998a:258) | Madras, which rescued five of the crew. Veracity of account has been questioned,[235] though taken seriously by Frederick Aldrich.[236] Fictionalised in Don C. Reed's 1995 novel The Kraken.[237]
| |||||
33 (📷) |
2 November 1874 | on beach, St. Paul Island, Indian Ocean (38°43′S 77°32′E / 38.717°S 77.533°E) {SIO} |
Found washed ashore | Architeuthis mouchezi Vélain (1875:1002) [nomen nudum]; Mouchezis sancti-pauli Vélain (1877:81); Ommastrephes mouchezi[200] | Entire; found in advanced state of decay | Tentacle(s?) and buccal mass | EL: 7.15 m | MNHN catalog nos. 3-2-658 & 3-2-659 ( tentacular clubs);[238] holotype of Mouchezis sancti-pauli Vélain, 1877 |
Vélain (1875:1002); Vélain (1877:81 & 83, fig. 8); Vélain (1878:81 & 83, fig. 8); Tryon (1879b:184, pl. 82 fig. 378); Owen (1881:159); Pfeffer (1912:32) | Gervais (1875:88); Verrill (1875c:213); Wright (1878:329) | Recorded by geologist Charles Vélain during French astronomical mission to Île Saint-Paul to observe the transit of Venus. Specimen was photographed.[28] | |
34 (📷) |
December 1874 | Grand Bank, Fortune Bay, Newfoundland {NWA} |
Found washed ashore | Architeuthis princeps | Entire, except for tail (cut up for dog food) | Jaws, one tentacular sucker | EL: 42–43 ft (12.8–13.1 m); HL+BL: 12–13 ft (3.7–4.0 m); ?TL: 30 ft (9.1 m); TL: 26 ft (7.9 m); TC: 16 in (41 cm); BL: 10 ft (3.0 m); jaws | YPM catalog nos. IZ 010272 (beak) & IZ 034836[permanent dead link]. Verrill specimen No. 6 and Verrill specimen No. 13 ("Fortune Bay specimen") | Verrill (1875a:35); Verrill (1875c:213); Verrill (1880a:186, 188, 217, pl. 17 fig. 11); Verrill (1881b:445, pl. 54 fig. 1); Verrill (1882c:10, 12, pl. 7 fig. 1, pl. 9 fig. 11) | Simms letter 27/X/1875 to Verrill; Frost (1934:102) | Data from 10/XII/1873 letter from Mr. Harvey to unknown individual citing measurements taken by G. Simms; Pfeffer (1912:21). Measurements are given differently in different papers. Verrill (1880a:186) and Verrill (1882c:10) states his No. 6 is same specimen as No. 3; this cannot be correct, as capture date for No. 6 is clearly stated as December 1874 by Verrill (1875c:213).[239] Verrill (1880a:188, pl. 17) repeats record as his No. 13. | |
35 | winter of 1874–1875 | near Harbor Grace, Newfoundland {NWA} |
Found washed ashore | Destroyed | None taken | None; Verrill specimen No. 12 ("Harbor Grace specimen") | Verrill (1875b:79); Verrill (1880a:188); Verrill (1882c:12) | Frost (1934:102) | "destroyed before its value became known, and no measurements are given" | |||
36 | Unknown (reported 1875) | west St. Modent (on Labrador side), Strait of Belle Isle, Newfoundland {NWA} |
Found alive | Architeuthis princeps or Architeuthis monachus of Steenstrup | Entire | None; cut up, salted, and barrelled for dog meat | ?TL: 37 ft (11 m); BL+HL: 15 ft (4.6 m); EL: 52 ft (16 m); SD: ≈2 in (5.1 cm) | None; Verrill specimen No. 7 ("Labrador specimen") | Verrill (1875a:36); Verrill (1880a:186); Verrill (1882c:10) | Dr. Honeyman article in Halifax newspaper; Frost (1934:101) | Data from unidentified third party cited in Halifax newspaper article. | |
37 (📷) |
25 April 1875 (or 26 April[240]) | north-west of Boffin Island, Connemara, Ireland {NEA} |
Found immobile at surface; attacked and chased by fishermen; arms successively hacked off and eventually killed | Architeuthis monachus; Architeuthis dux Steenstrup, 1857[241] | Entire | Beak and buccal mass, one arm ("much mutilated and decayed", missing horny rings), portions of both tentacles ("shrunk and distorted", missing horny rings on large central club suckers); head, eyes and second arm initially saved, but soon lost/destroyed | TL: 30 ft (9.1 m) [fresh]; TL: 14/17 ft (4.3/5.2 m) [pickled]; CL: 2 ft 9 in (0.84 m) [shrunken]; CSD: nearly 1 in (2.5 cm); SSD: 3⁄20 in (0.38 cm); AL: 8 ft (2.4 m) [fresh]; AC: 15 in (38 cm) [fresh]; beak: ≈5+1⁄4 in (13 cm) × 3+1⁄2 in (8.9 cm); "trunk": "fully as long as the canoe"; EyD: ≈15 in (38 cm); WT: ≈6 st (38 kg) [head only]; additional sucker measurements | NMI catalog no. 1995.16 (beak in spirit)[241] | O'Connor (1875:4502); More (1875b:4569); More (1875c:123); Verrill (1875c:214); Massy (1909:30); Nunn & Holmes (2008) | Galway Express 1875; Ritchie (1918:137); Massy (1928:32); Taylor (1932:3); Robson (1933:692); Rees (1950:40); Hardy (1956:285); Collins (1998:489) | On public display. Caught by three-man coarse fish. Found motionless at surface surrounded by gulls, becoming active upon being attacked by fishermen, swimming away "at a tremendous rate" and releasing ink. Progressively disabled with a knife (fishermen having no gaff or spare rope) as chased for 2 hours over 5 miles (8.0 km), before head eventually severed; heavy mantle allowed to sink. Specimen secured and preserved by Sergeant Thomas O'Connor of the Royal Irish Constabulary and forwarded by him to the museum of the Royal Dublin Society, Dublin (now the National Museum of Ireland – Natural History ).
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38 | October 1875 | Grand Banks [of Newfoundland], Atlantic Ocean (chiefly 44°–44°30'N 49°30'–49°50'W) {NWA} |
Found floating at surface; "mostly entirely dead" but small minority "not quite dead, but entirely disabled" | Architeuthis | Multiple; mutilated by birds and fishes to varying degrees, especially limbs; No. 25 missing parts of arms; No. 26 with intact arms and tentacles | None; cut up for cod bait | No. 25: Filled ≈75 US gal (280 L) tub; WT: nearly 1,000 lb (450 kg) [estimate, complete]; No. 26: TL: 36 ft (11 m); Howard specimens: BL+HL?: mostly 10–15 ft (3.0–4.6 m) [excluding "arms"]; BD: ≈18 in (46 cm) [average]; AL: usually 3–4 ft (0.91–1.22 m) [incomplete]; AD: "about as large as a man's thigh" [at base]; Tragabigzanda specimens: BL+HL?: 8–12 ft (2.4–3.7 m) [excluding "arms"] | None; included Verrill specimen No. 25 and Verrill specimen No. 26 | Verrill (1881a:251); Verrill (1881b:396); Verrill (1882c:19) | Frost (1934:103) | An unusual number (≈25–30) of mostly dead giant squid found by season . Verrill conjectured that this mass mortality might have been due to an outbreak of disease or parasites, and/or related to their reproductive cycle.
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39 | c. 1876 | Clifford Bay, Cape Campbell, New Zealand {SWP} |
Found washed ashore | Entire | Jaws[242] | BL: 7 ft (2.1 m) [estimate]; EL: ≈20 ft (6.1 m) [estimate] | Colonial Museum [NMNZ][242] | Robson (1887:156); Kirk (1880) | Pfeffer (1912:32); Dell (1952:98) | |||
40 | 20 November 1876 | Hammer Cove, southwest arm of Green Bay, Notre Dame Bay, Newfoundland {NWA} |
Found washed ashore | Partial specimen; devoured by foxes and seabirds | Piece of pen 16 in (41 cm) long | WH: 18 in (46 cm); FW: 18 in (46 cm) | In Harvey's possession; Verrill specimen No. 15 ("Hammer Cove specimen") | Verrill (1880a:190); Verrill (1880b:284); Verrill (1882c:14) | M. Harvey letter 25 August 1877 to Verrill; Frost (1934:102) | |||
41 | 1877? | Norway {NEA} |
Not stated | Map location only | Sivertsen (1955:11, fig. 4) | |||||||
42 (📷) |
24 September 1877 | Trinity Bay, Newfoundland {NWA} |
Found washed ashore, alive | Architeuthis princeps; Ommastrephes (Architeuthis) princeps[187] | Entire; "nearly perfect specimen" | Loose suckers | (see Verrill, 1880a:220) HL+BL: 9.5 ft (2.9 m); BC: 7 ft (2.1 m); TL: 30 ft (9.1 m); AL: 11 ft (3.4 m) [longest, ventral]; AC: 17 in (43 cm) [ventral]; beak; FW: 2 ft 9 in (0.84 m) | YPM catalog nos. IZ 017927, IZ 017928, IZ 017929 & IZ 017930. Verrill specimen No. 14 ("Catalina specimen") | Harvey (1877); [Anon.] (1877a:266, 269, fig.); [Anon.] (1877b:867, fig.); [Anon.] (1877c:305, fig.); Verrill (1877:425); Tryon (1879b:185); Verrill (1880a:189, pl. 17 figs. 1–10, pls. 19–20); Verrill (1880b:295, pl. 12); Verrill (1882c:13, pl. 8, pl. 9 figs. 1–10, pl. 10) | Owen (1881:163); Hatton & Harvey (1883:242); Pfeffer (1912:21); Frost (1934:102); Miner (1935:187, fig., 201); Ellis (1997a:31) | Measured fresh by M. Harvey; examined preserved (poorly) by Verrill at Ward's.[75] Described by Frederick Aldrich as "largest giant squid to be encountered in Newfoundland".[126]
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43 | October 1877 | Trinity Bay, Newfoundland {NWA} |
Not stated | "big squid" | None | None taken | None; Verrill specimen No. 17 ("Trinity Bay specimen") | Verrill (1880a:191); Verrill (1880b:285); Verrill (1882c:15) | M. Harvey letter 17 November 1877 to Verrill citing reference to specimen by John Duffet; Frost (1934:102) | Specimen cut up and used for manure. | ||
44 (📷) |
21 November 1877 | Trinity Bay, Newfoundland {NWA} |
Found washed ashore, alive | ?Architeuthis princeps | Entire | None; carried off by tide | BL(+HL?): 11 ft (3.4 m); TL: 33 ft (10 m); AL: 13 ft (4.0 m) [estimate] | None; Verrill specimen No. 16 ("Lance Cove specimen") | Verrill (1880a:190); Verrill (1880b:285); Verrill (1882c:14) | M. Harvey letter 27 November 1877 to Verrill citing measurements taken by John Duffet; Frost (1934:102) | Found still alive, having "ploughed up a trench or furrow about 30 feet [9.1 m] long and of considerable depth by the stream of water that it ejected with great force from its siphon. When the tide receded it died." | |
(📷) |
1878 (accessioned) | Catlins, New Zealand {SWP} |
Not stated | Architeuthis sp. | Entire? | Beak | BL: 7 ft (2.1 m);[243] ML: 1.6 m [estimate]; EL: ≈10 m [estimate][244] | Otago Museum catalog no. IV119151 |
Lau (2021); [OM] (2021) | Copedo (2022) | On public display. Collected by Capt. Charles Hayward ( | |
45 (📷) |
2 November 1878 | {NWA} |
Found aground offshore, alive; secured to tree with grapnel and rope; died as tide receded |
?Architeuthis princeps | Entire | None; cut up for dog food | BL+HL: 20 ft (6.1 m); TL: 35 ft (11 m)[nb 20] | None; Verrill specimen No. 18 (" Thimble Tickle specimen ") |
Verrill (1880a:191); Verrill (1880b:285); Verrill (1882c:15); Ellis (1998a:6, 89, 107) | M. Harvey letter 30 January 1879 to | Discovered by fisherman Stephen Sherring and two others. | |
46 | 2 December 1878 | Three Arms, South Arm of Notre Dame Bay, Newfoundland {NWA} |
Found washed ashore | ?Architeuthis princeps | Entire, mutilated and with arms missing (only one arm "perfect") | None; cut up for dog food | BL+HL: 15 ft (4.6 m); BC: 12 ft (3.7 m); AL: 16 ft (4.9 m); AD: "thicker than a man's thigh" | None; Verrill specimen No. 19 ("Three Arms specimen") | Verrill (1880a:192); Verrill (1880b:286); Verrill (1882c:16) | M. Harvey letter 30 January 1879 to | Found dead by fisherman William Budgell after heavy gale. Considered by Paxton (2016a:83) as the "longest measured" standard length of any giant squid specimen. | |
47 (📷) |
23 May 1879 | Lyall Bay, Cook Strait, New Zealand {SWP} |
Found washed ashore | Steenstrupia stockii Kirk, 1882 [=Architeuthis sp.?[266]] | Entire, but somewhat mutilated; missing ends of tentacles | Pen, beak, tongue, some suckers | ML: 9 ft 2 in (2.79 m); BC: 7 ft 3 in (2.21 m); HL: 1 ft 11 in (0.58 m); BL+HL: 11 ft 1 in (3.38 m); HC: 4 ft (1.2 m); AL: 4 ft 3 in (1.30 m); AC: 11 in (28 cm); ASC: 36; TL: 6 ft 2 in (1.88 m) [incomplete]; FL: 24 in (61 cm); FW: 13 in (33 cm) (single); GL: 6 ft 3 in (1.91 m); GW: 11 in (28 cm); other measurements | NMNZ catalog nos. M.125403 & M.125405;[267] holotype of Steenstrupia stockii Kirk, 1882. Kirk specimen No. 3 | Kirk (1880:310); Verrill (1881b:398); Kirk (1882:286, pl. 36 figs. 2–4) | Verrill (1882d:477); Kirk (1888:34); Pfeffer (1912:34); Suter (1913:1051); Dell (1952:98); Dell (1970:27); Stevens (1980:213, fig. 12.24); Stevens (1988:149, fig. 2); Judd (1996); Paxton (2016a:83); Greshko (2016) | Measurements taken by T.W. Kirk. Has been called the "largest specimen recorded in the scientific literature" based on erroneous total length of "approximately 20 m",[268] itself based on claim by Roper & Boss (1982:104) relating to unspecified specimen "stranded on a beach in New Zealand in 1880 [sic]". Considered by Paxton (2016a:83) as the longest reliably measured mantle length of any giant squid specimen (less reliably that of #104), but measurement considered dubious by experts due to wide discrepancy with reported gladius length.[94][nb 23] | |
48 | 1879 | off Nova Scotia, Canada (42°49′N 62°57′W / 42.817°N 62.950°W) {NWA} |
From fish stomach, Alepidosaurus [sic] ferox | ?Architeuthis megaptera Verrill, 1878; ?Architeuthis harveyi (Kent, 1874) | Terminal part of tentacular arm | Portion of arm | 18 in (46 cm) long | NMNH catalog no. 576962. Verrill specimen No. 20 ("Banquereau specimen" [after Banquereau Bank, a bank off Nova Scotia]) | Verrill (1880a:193); Verrill (1880b:287); Verrill (1882c:16) | Frost (1934:103) | Lancetfish taken by Capt. J.W. Collins of schooner Marion on halibut trawl-line. | |
49 (📷) |
September 1879 | Olafsfjord, Iceland {NEA} |
Architeuthis | Left tentacle | TL: 7680+ mm; CL: 1010 mm; CSC: 268; TSC: 290; additional indices and counts | ZMUC [specimen NA-7 of Roeleveld (2002)] | Roeleveld (2002:727) | Tentacle morphology examined by Roeleveld (2002). | ||||
50 | October 1879 | near Brigus, Conception Bay, Newfoundland {NWA} |
Found washed ashore | Two arms with other mutilated parts | Undetermined | AL: 8 ft (2.4 m) | None?; Verrill specimen No. 22 ("Brigus specimen") | Verrill (1880a:194); Verrill (1880b:287); Verrill (1882c:17) | Frost (1934:103) | Found after storm. Information provided by Moses Harvey. | ||
51 | 1 November 1879 | James's Cove, Bonavista Bay, Newfoundland {NWA} |
Found at surface, alive | Entire | None; cut up by fishermen | EL: 38 ft (12 m); BL: 9 ft (2.7 m); BC: ≈6 ft (1.8 m); TL: 29 ft (8.8 m) | None; Verrill specimen No. 23 ("James's Cove specimen") | Verrill (1880a:194); Verrill (1880b:287); Verrill (1882c:17) | Morning Chronicle of St. John's 9 December 1879; Frost (1934:103) | Found alive and driven ashore. | ||
52 | Unknown (reported 1880) | near Boulder Bank, Nelson, New Zealand {SWP} |
Not stated; hook and line? | Not indicated | Undetermined | 8 ft (2.4 m) long | None?; Kirk specimen No. 4 | Kirk (1880:310); Verrill (1881b:398) | Newspaper article | Caught by fishing party. No other data. | ||
53 | Unknown (reported 1880) | near Flat Point, east coast, New Zealand {SWP} |
Not stated | Not indicated | Undetermined | None | None?; Kirk specimen No. 5 | Kirk (1880:310); Verrill (1881b:398) | Description sent to Mr. Beetham, M.H.R., by Mr. Moore | Found by Mr. Moore. No other data. | ||
54 (📷) |
April 1880 | Grand Banks, Newfoundland {NWA} |
Found dead at surface | Architeuthis harveyi (Kent, 1874) | Head, tentacles, and arms only | Head, tentacles, and arms | TL: 66 in (170 cm); ASC: 330; extensive measurements and counts | YPM catalog no. 12600y. Verrill specimen No. 24 ("Grand Banks specimen" [2nd]) | Verrill (1881b:259, pl. 26 figs. 1–4, pl. 38 figs. 3–7); Verrill (1882c:18, pl. 4 figs. 2–2a, pl. 5 figs. 6–8, pl. 6) | Pfeffer (1912:19); Frost (1934:103) | Found dead by Capt. O.A. Whitten of schooner Wm.H. Oakes. Arm and sucker regeneration documented by #549 ).
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55 (📷) |
6 June 1880 | Island Bay, Cook Strait, New Zealand {SWP} |
Found washed ashore | Architeuthis verrilli Kirk, 1882 | Entire | Not specified | ML: 7 ft 6 in (2.29 m); BC: 9 ft 2 in (2.79 m); TL: 25 ft (7.6 m); AL(I, II, IV): 9 ft (2.7 m); AC(I, II, IV): 15 in (38 cm); AL(III): 10 ft 5 in (3.18 m); AC(III): 21 in (53 cm); ASC(III): 71; HC: 4 ft 3 in (1.30 m); HL: 19 in (48 cm); FL: 30 in (76 cm); FW: 28 in (71 cm); EyD: 5 in (13 cm) by 4 in (10 cm) | NMNZ; holotype of Architeuthis verrilli Kirk, 1882; specimen no longer extant[270] | Kirk (1882:284, pl. 36 fig. 1) | Verrill (1882d:477); Kirk (1888:35); Pfeffer (1912:33); Suter (1913:1052); Dell (1952:98); Dell (1970:27) | Measurements taken by Kirk, except TL by James McColl. Beak and portions of gladius ("skeleton") taken by Italian fishermen and not recovered. | |
56 | c. 1880 | Kvænangen, Norway {NEA} |
Found washed ashore | Architeuthis dux Steenstrup, 1857 | Entire | None | None | Grieg (1933:19) | Sivertsen (1955:11) | |||
57 | c. 1880 | Tønsvik, Tromsøysund, Norway {NEA} |
Found washed ashore | Architeuthis dux Steenstrup, 1857 | Entire | None | None | Grieg (1933:19) | ||||
58 | October 1880 | Kilkee, County Clare, Ireland {NEA} |
Found washed ashore | "octopus"; Architeuthis sp. | O'Brien (1880:585); Ritchie (1918:137) | Rees (1950:40); Collins (1998:489) | Originally cited as an octopus. | |||||
59 | first week of November 1881 | on beach, Hennesey's Cove, Long Island, Placentia Bay, Newfoundland {NWA} |
Found washed ashore | Architeuthis princeps? | Entire; "much mutilated by crows and other birds" | Not stated | "very large"; BL+HL: 26 ft (7.9 m) [estimate] | Verrill specimen No. 28 | Verrill (1882c:221) | M. Harvey letter 19 December 1881 to Verrill | Found by Albert Butcher and George Wareham, "who cut a portion from the head", at uninhabited locality; Verrill considered their estimate of the specimen's length "probably too large". Harbour Buffet, Placentia Bay. Only mentioned in Verrill (1882c:221); overlooked by Ellis (1994a:379–384), Ellis (1998a:257–265), and Sweeney & Roper (2001) .
| |
60 (📷) |
10 November 1881 | Portugal Cove, near St. John's, Newfoundland {NWA} |
Found floating dead near shore | Architeuthis harveyi (Kent, 1874) | Entire | Entire (somewhat mutilated and poorly preserved) | a) BL: 5.5 ft (1.7 m); HL: 1.25 ft (0.38 m); EL: 28 ft (8.5 m); BC: 4.5 ft (1.4 m) b) ML: 4.16 ft (1.27 m); BC: 4 ft (1.2 m); FL: 1.75 ft (0.53 m); FW: 8 in (20 cm) [single]; TL: 15 ft (4.6 m); CL: 2 ft (0.61 m); AL: 4.66 ft (1.42 m) [ventral, minus tip]; TC: 8.5 in (22 cm) [at base]; additional measurements | E.M. Worth Museum (101 Bowery, NY, NY). Verrill specimen No. 27 | [Anon.] (1881:821, fig.); Verrill (1881b:422); Verrill (1882a:71); Verrill (1882c:201, 219) | Morris article in 25 November 1881 New York Herald; Hatton & Harvey (1883:242); Pfeffer (1912:19); Ellis (1997a:34) | Obtained by Mr. Morris, photographed by E. Lyons (St. John's), shipped on ice by steamer Catima to New York, purchased and preserved by E.M. Worth. Measurements by a) Inspector Murphy (chief Board of Public Works) when iced; b) Verrill of fixed specimen. An 1881 specimen from Portugal Cove with a "body" reportedly 11 ft (3.4 m) long, mentioned in Ward's), as Verrill saw the specimen shortly before he began modelling.[78]
| |
61 | 30 June 1886 | Cape Campbell, New Zealand {SWP} |
Found washed ashore | Architeuthis kirkii Robson, 1887 | Entire | Beak and club |
ML: 8 ft 3 in (2.51 m); HL: 1 ft 9 in (0.53 m); AL: 6 ft 6 in (1.98 m); TL: 18 ft 10 in (5.74 m); EL: 28 ft 10 in (8.79 m); BC: ≈8 ft (2.4 m) [estimate] | NMNZ catalog nos. M.125404 & ?M.125406;[267] holotype of Architeuthis kirkii Robson, 1887. Kirk specimen No. 2 | Kirk (1879:310); Verrill (1881b:398); Robson (1887:156) | C.H.[W.] Robson letter 19 June 1879 to T.W. Kirk; Pfeffer (1912:35); Suter (1913:1048); Dell (1952:98); Dell (1970:27) | Found by Mr. C.H.[W.] Robson; beak given to Mr. A. Hamilton. | |
1886 | Cupids and Hearts Content (one specimen from each), Newfoundland {NWA} |
Found washed ashore | "giant squid" | Two specimens; entire? | None?; cut up for bait | None given | Earle (1977:53) | Moses Harvey only learned of specimens after their destruction. Information sourced from clippings found in one of Harvey's scrapbooks preserved at Newfoundland Public Archives (PG/A/17).[207] | ||||
62 (📷) |
"early" October 1887 | Lyall Bay, New Zealand {SWP} |
Found washed ashore | Architeuthis longimanus Kirk, 1888 | Entire | Beak and buccal-mass | Female | EL: 55 ft 2 in (16.81 m); ML: 71 in (180 cm); BC: 63 in (160 cm); extensive additional measurements and description[nb 24] | Kirk (1888:35, pls. 7–9); Pfeffer (1912:36) | Suter (1913:1049); Dell (1952:98); Dell (1970:27); Wood (1982:191); Ellis (1998a:7, 92); O'Shea & Bolstad (2008); Dery (2013); Paxton (2016a:83) | Strangely proportioned animal that has been much commented on; sometimes cited as the longest giant squid specimen ever recorded.[272][nb 24] Considered by Paxton (2016a:83) as candidate for "longest measured" total length of any giant squid specimen (together with #45, and less reliably #209). Found by Mr. Smith, local fisherman. Measurements taken by T.W. Kirk. Date found listed incorrectly in Dell (1952:98).[273] | |
63 | 27 August 1888 | between Pico and St. George, Azores Islands (38°33′57″N 30°39′30″W / 38.56583°N 30.65833°W) at 1266 m depth {NEA} |
By benthic trawl | Architeuthis? sp.?[274] | Large beak | Undetermined | None | Joubin (1895:34) | ||||
64 | September 1889 | Løkberg farm, Mo i Rana, Norway {NEA} |
Found washed ashore | Entire | None | BL: ≈5 ells (3.1 m); TL: 10–12 ells (6.3–7.5 m) | [Anon.] (1890:190) | Sivertsen (1955:11, fig. 4) | Bergen Museum notified of find by Lorentz Pettersen of Sjona, Helgeland. Failure to secure remains prompted museum to issue notice in June 1890 issue of Naturen seeking specimens in future (which would be first for a Norwegian museum) and offering to cover all associated transportation and packing costs in addition to regular compensation.[275]
| |||
1890 | Island Cove, Newfoundland {NWA} |
Found washed ashore | "giant squid" | Entire? | None?; cut up for bait? | None given | Earle (1977:53) | Moses Harvey only learned of specimen after its destruction. Information sourced from clippings found in one of Harvey's scrapbooks preserved at Newfoundland Public Archives (PG/A/17).[207] | ||||
65 | Unknown (reported 1892) | Sao Miguel Island, Azores Islands {NEA} |
Found washed ashore | Architeuthis princeps | Entire? | tentacle club |
Beak measurements | Museum in Lisbon[276] | Girard (1892:214, pls. 1–2) | Pfeffer (1912:27); Robson (1933:692) | ||
66 | 1892 | Greenland {NWA} |
Not stated | Architeuthis monachus | Posselt (1898:279) | |||||||
[2] | Unknown (reported November 1894) | Talcahuano, Chile {SEP} |
Unknown; collected and donated to ZMB by Ludwig Plate |
Ommastrephes gigas; Dosidicus gigas[279] |
Entire | Entire, internal parts missing, preserved in alcohol; "exceptionally good condition" (Glaubrecht & Salcedo-Vargas, 2004:55) | Female (adult) | ML: 865 mm; MW: 230 mm; EL: 1740 mm; HL: 160 mm; HW: 190 mm; FL: 440 mm; FW: 600 mm; TL: 720 mm; CL: 225 mm; AL(I): 460 mm; AL(II): 450 mm; AL(III): 500 mm; AL(IV): 440 mm; LSD: 20 mm [tentacle]; LSD: 15 mm [arm II]; LSD: 14 mm [arm II]; EyD: 80 mm; Lens: 35 mm | ZMB Moll. 49.804 | Martens (1894); Glaubrecht & Salcedo-Vargas (2004:53, figs. 1a–f, 2a–g) | Möbius (1898a:373); Möbius (1898b:135); [Anon.] (1899:38); [Anon.] (1902a:41); Kilias (1967:491, fig.); Wechsler (1999) | Non-architeuthid. On public display. First noted by Übersee-Museum Bremen with sperm whale skull. Re-identified as Dosidicus gigas in June 1998 by Mario Alejandro Salcedo-Vargas. Internal parts apparently removed when specimen originally dissected by Martens or prepared for exhibition (1894–97).
|
67 (📷) |
4 February 1895 | {NWP} |
In net | Architeuthis japonica Pfeffer, 1912 | Entire | Undetermined | Female | ML: 720 mm; MW: 235 mm; GL: 640 mm; FL: 280 mm; FW: 200 mm; TL: 2910 mm; extensive additional measurements and description | Undetermined; ?Zoological Institute, Science College, Tokyo; holotype of Architeuthis japonica Pfeffer, 1912 | Mitsukuri & Ikeda (1895:39, pl. 10); Pfeffer (1912:27) | Sasaki (1916:89) | Caught in net after 2–3-day storm. |
68 (📷) |
18 July 1895 | near Angra, Azores Islands (38°34'45"N, 29°37'W) {NEA} |
Caught at surface (from sperm whale vomit) using shrimp net | Dubioteuthis physeteris Joubin, 1900 [=Architeuthis physeteris (Joubin, 1900)[281]] | Mantle only | Mantle | Male | ML: 460 mm; BD: 115 mm; FL: 220 mm; FW: 110 mm; GL: 390 mm | MOM [station 588]; holotype of Dubioteuthis physeteris Joubin, 1900[282] | Joubin (1900:102, pl. 15 figs. 8–10); Pfeffer (1912:24) | Hardy (1956:288); Roper & Young (1972:220); Toll & Hess (1981:753) | |
[3] (📷) |
18 July 1895 | near Angra, Azores Islands (38°34'45"N, 29°37'W) {NEA} |
Caught at surface (from sperm whale vomit) with shrimp net | Architeuthis sp.?; non-architeuthid[276] | Several jaws | Undetermined | None | Joubin (1900:46, pl. 14 figs. 1–2) | Pfeffer (1912:27); Clarke (1956:257) | Non-architeuthid. | ||
69 (📷) |
10 April 1896 | Hevnefjord, Norway {NEA} |
Found washed ashore | Architeuthis dux Steenstrup, 1857 | Entire | Entire | Female | BL: 2.5 m; AL: 2.5 m; TL: 7.25 m | VSM | Storm (1897:99); Grieg (1933:19) | Brinkmann (1916:178); Nordgård (1923:11); Nordgård (1928:71); Sivertsen (1955:11) | Model completed in 1954 based on this specimen and #70; restored in 2010.[283] |
70 (📷) |
27 September 1896 [or 28 September[284]] | Hevnefjord, Norway {NEA} |
Found washed ashore | Architeuthis dux Steenstrup, 1857 | Entire | Entire, posterior part missing | Male | TL: 1030+ mm; CL: 900 mm; CSC: 294; TSC: >298; LRL: 17.9 mm; URL: 16.2 mm; additional beak measurements, indices, and counts | VSM; VSM 110a [specimen NA-18 of Roeleveld (2000) and Roeleveld (2002)] | Storm (1897:99, fig. 20); Grieg (1933:19); Roeleveld (2000:185); Roeleveld (2002:727) | Brinkmann (1916:178, fig. 2); Nordgård (1923:11); Nordgård (1928:71); Sivertsen (1955:11); Toll & Hess (1981:753) | Beak morphometrics studied by Roeleveld (2000). Tentacle morphology examined by Roeleveld (2002). Model completed in 1954 based on this specimen and #69; restored in 2010.[283] |
71 | Unknown (reported 1898) | Iceland {NEA} |
Not stated | Architeuthis monachus | Not specified | Undetermined | None | Posselt (1898:279) | Bardarson (1920:134) |
Type specimens
The following table lists the nominal species-level
Binomial name and author citation | Systematic status | Type locality | Type specimen and type repository |
---|---|---|---|
Loligo bouyeri Crosse & Fischer, 1862:138 | Architeuthid?[286] | Canary Islands? | (#18) Unresolved |
Architeuthis clarkei Robson, 1933:682, text-figs. 1–7, pl. 1 | Undetermined | Scarborough Beach, Yorkshire, England | (#107) BMNH Holotype 1933.1.30.5 + 1926.3.31.24 (radula and beak)[287] |
Architeuthis dux Steenstrup, 1857:183 | Nomen tantum |
||
Architeuthis dux Steenstrup in Harting, 1860:11, pl. 1 fig. 1A | Valid species[288] | 31°N 76°W / 31°N 76°W (Atlantic Ocean) | (#14) ZMUC Holotype[192] |
Plectoteuthis grandis Owen, 1881:156, pls. 34–35 | Architeuthis sp.[289] | Not indicated | (#27) BMNH Holotype[290] [not traced by Lipiński et al. (2000)] |
Architeuthis halpertius Aldrich, 1980:59 | Nomen nudum[nb 25] | ||
Loligo hartingii Verrill, 1875b:86, fig. 28 | Valid species; Architeuthis hartingii[293] | Not indicated | (#16) University of Utrecht as Architeuthis dux, identification by Harting |
Megaloteuthis harveyi Kent, 1874a:181 | Architeuthis sp. | Conception Bay, Newfoundland | (#29) YPM Type 12600y[294] |
Architeuthis japonica Pfeffer, 1912:27 | Undetermined | Tokyo Bay, Japan | (#67) Undetermined; Holotype [=Mitsukuri & Ikeda (1895:39–50, pl. 10)] |
Architeuthis kirkii Robson, 1887:155 | Architeuthis stockii (Kirk, 1882)[295] | Cape Campbell, New Zealand | (#61) NMNZ Holotype M.125404 + ?M.125406[267] |
Architeuthis longimanus Kirk, 1888:34, pls. 7–9 | Architeuthis stockii (Kirk, 1882)[295] | Lyall Bay, New Zealand | (#62) NMNZ Holotype; specimen not located[270] |
Megateuthis martensii Hilgendorf, 1880:67 | Valid species; Architeuthis martensii | Yedo Japan fish market, Japan |
(#28) ZMB Moll. 34716 + 38980 |
Architeuthis megaptera Verrill, 1878:207 | Non-architeuthid; Sthenoteuthis pteropus (Steenstrup, 1855) | Nova Scotia, Canada | (#[1]) NSMC 1870–Z-2 |
Architeuthis? monachus Steenstrup, 1857:184 | Nomen tantum |
||
Architeuthis monachus Steenstrup in Harting, 1860:11 | Architeuthis dux Steenstrup, 1857[203] | Raabjerg Strand; Northwest coast of Jutland, Denmark[296] | (#13) ZMUC Holotype[296] |
Architeuthis mouchezi Vélain, 1875:1002 | Nomen nudum; see Mouchezis sancti-pauli | ||
Architeuthis nawaji Cadenat, 1935:513 | Undetermined | Île d'Yeu, Bay of Biscay, France | (#110) Unresolved |
Dubioteuthis physeteris Joubin, 1900:102, pl. 15 | Valid species; Architeuthis physeteris[281] | Azores (38°34'45"N 29°37'W); from sperm whale stomach | (#68) MOM Holotype [station 588][282] |
Architeuthis princeps Verrill, 1875a:22 | Nomen nudum | ||
Architeuthis princeps Verrill, 1875b:79, figs. 25–27 | Undetermined | a) Grand Banks, Newfoundland; b) North Atlantic (sperm whale stomach) | (#22 and 26) NMNH? [not found in collections to date]; Syntypes (a) Verrill specimen No. 1, lower beak; b) Verrill specimen No. 10, upper and lower beak) |
Dinoteuthis proboscideus More, 1875a:4527 | Architeuthis sp.[289] | Dingle, County Kerry, Ireland | (#3) Unresolved |
Mouchezis sancti-pauli Vélain, 1877:81, text-fig. 8 | Valid species; Architeuthis sanctipauli | on beach, St. Paul Island (38°43′S 77°32′E / 38.717°S 77.533°E), South Indian Ocean | (#33) MNHN Holotype 3-2-658 and 3-2-659 (tentacular clubs only)[297] |
Steenstrupia stockii Kirk, 1882:286, pl. 36 figs. 2–4 | Valid species; Architeuthis stockii[295] [Architeuthid per Pfeffer (1912:2)] | Cook Strait, New Zealand | (#47) NMNZ Holotype M.125405 + M.125403[267] |
Architeuthis titan Steenstrup in Verrill, 1875b:84 [in Verrill (1881b:238, footnote)] | Nomen nudum | ||
Architeuthis verrilli Kirk, 1882:284, pl. 36 fig. 1 | Species dubium[295] | Island Bay, Cook Strait, New Zealand |
(#55) NMNZ Holotype; [see Förch (1998:89)] |
Abbreviations
The following abbreviations are used in the List of giant squid table.
Oceanic sectors
Oceanic sectors used in the main table follow Sweeney & Roper (2001): the Atlantic Ocean is divided into sectors at the equator and 30°W, the Pacific Ocean is divided at the equator and 180°, and the Indian Ocean is defined as the range 20°E to 115°E (the Arctic and Southern Oceans are not distinguished). An additional category has been created to accommodate the handful of specimens recorded from the Mediterranean Sea.
- NEA, Northeast Atlantic Ocean
- NWA, Northwest Atlantic Ocean
- SEA, Southeast Atlantic Ocean
- SWA, Southwest Atlantic Ocean
- NEP, Northeast Pacific Ocean
- NWP, Northwest Pacific Ocean
- SEP, Southeast Pacific Ocean
- SWP, Southwest Pacific Ocean
- NIO, Northern Indian Ocean
- SIO, Southern Indian Ocean
- MED, Mediterranean Sea
Measurements
Abbreviations used for measurements and counts follow Sweeney & Roper (2001) and are based on standardised acronyms in teuthology, primarily those defined by Roper & Voss (1983), with the exception of several found in older references. Following Sweeney & Roper (2001), the somewhat non-standard EL ("entire" length) and WL ("whole" length) are used in place of the more common TL (usually total length; here tentacle length) and SL (usually standard length; here spermatophore length), respectively.
- AC, arm circumference (AC(I), AC(II), AC(III) and AC(IV) refer to measurements of specific arm pairs)
- AD, arm diameter (AD(I), AD(II), AD(III) and AD(IV) refer to measurements of specific arm pairs)
- AF, arm formula
- AL, arm length (AL(I), AL(II), AL(III) and AL(IV) refer to measurements of specific arm pairs)
- ASC, arm sucker count
- ASD, arm sucker diameter
- BAC, buccal apparatus circumference
- BAL, buccal apparatus length
- BC, body circumference (assumed to mean greatest circumference of mantle unless otherwise specified)
- BD, body diameter (assumed to mean greatest diameter of mantle)
- BL, body length (usually equivalent to mantle length, as head length is often given separately)
- CaL, carpus length
- CL, club length (usually refers to expanded portion at the apex of tentacle)
- CSC, club sucker count
- CSD, club sucker diameter (usually largest) [usually equivalent to LSD]
- CW, club width
- DC, dactylus club length
- EC, egg count
- ED, egg diameter
- EL, "entire" length (end of tentacle(s), often stretched, to posterior tip of tail; in contrast to WL, measured from end of arms to posterior tip of tail)
- EyD, eye diameter
- EyOD, eye orbit diameter
- FL, fin length
- FuCL, funnel cartilage length
- FuCW, funnel cartilage width
- FuD, funnelopening diameter
- FuL, funnel length
- FW, fin width
- GiL, gill length
- GL, gladius (pen) length
- GW, gladius (pen) width
- G(W), daily growth rate (%)
- HC, head circumference
- HeL, hectocotylus length
- HL, head length (most often base of arms to edge of mantle)
- HW, head width
- LAL, longest arm length
- LRL, lower rostral length of beak
- LSD, largest sucker diameter (on tentacle club) [usually equivalent to CSD]
- MaL, manus length
- ML, dorsal mantle length (used only where stated as such)
- MT, mantle thickness
- MW, maximum mantle width (used only where stated as such)
- NGL, nidamental gland length
- PL, penis length
- RaL, radula length
- RaW, radula width
- RL, rachis length
- RW, rachis width
- SInc, number of statolith increments
- SL, spermatophore length
- SoA, spermatophores on arms
- SSD, stalk sucker diameter
- SSL, spermatophore sac length
- TaL, tail length
- TC, tentacle circumference (most often of tentacle stalk)
- TCL, tentacle club length
- TD, tentacle diameter (most often of tentacle stalk)
- TL, tentacle length
- TSC, tentacle sucker count (club and stalk combined)
- TSD, tentacle sucker diameter (usually largest)
- URL, upper rostral length of beak
- VML, ventral mantle length
- WL, "whole" length (end of arms, often damaged, to posterior tip of tail; in contrast to EL, measured from end of tentacles to posterior tip of tail)
- WT, weight
Repositories
Institutional acronyms follow Sweeney & Roper (2001) and are primarily those defined by Leviton et al. (1985), Leviton & Gibbs (1988), and Sabaj (2016). Where the acronym is unknown, the full repository name is listed.
- AMNH, American Museum of Natural History, New York City, New York, United States
- AMS, Australian Museum, Sydney, New South Wales, Australia
- BAMZ, Bermuda Aquarium, Museum and Zoo, Flatts Village, Bermuda
- BMNH, Natural History Museum, Cromwell Road, London, England (formerly British Museum (Natural History))
- CEPESMA, Museo-Aula del Mar, Coordinadora para el Estudio y la Protección de las Especies Marinas, Luarca, Spain
- EI, Essex Institute, Salem, Massachusetts, United States
- FOSJ, Department of Fisheries and Oceans, St. John's, Newfoundland, Canada
- ICM, Instituto de Ciencias del Mar, Barcelona, Spain
- MCNOPM, Museo de Ciencias Naturales de Puerto Madryn (Museum of Natural Sciences and Oceanography), Puerto Madryn, Argentina
- MHNLR, Muséum national d'histoire naturelle, La Rochelle, France
- MHNN, Muséum national d'histoire naturelle(Musee Barla), Nice, France
- MMF, Museu Municipal do Funchal, Funchal, Madeira
- MNHN, Muséum national d'histoire naturelle, Paris, France
- MOM, Musée Océanographique, Monaco
- MUDB, Department of Biology, Memorial University, Newfoundland, Canada
- NIWA, National Institute of Water and Atmospheric Research, Wellington, New Zealand
- NMI, National Museum of Ireland – Natural History, Dublin, Ireland
- NMML, National Marine Mammal Laboratory, Alaska Fisheries Science Center, Seattle, Washington, United States
- NMNH, National Museum of Natural History, Smithsonian Institution, Washington, District of Columbia, United States
- NMNZ, Museum of New Zealand Te Papa Tongarewa, Wellington, New Zealand (formerly Colonial Museum; Dominion Museum)
- NMSJ, Newfoundland Museum, St. John's, Newfoundland, Canada
- NMSZ, National Museum of Scotland, Zoology Department, Edinburgh, Scotland (formerly Royal Museum of Scotland; formerly Royal Scottish Museum, Edinburgh)
- NMV, Museum Victoria, Melbourne, Victoria, Australia (formerly National Museum of Victoria)
- NSMC, Nova Scotia Museum, Halifax, Canada
- PASS, Peabody Academy of Science, Salem, Massachusetts, United States (now in Peabody Museum of Salem?)
- RMNH, Naturalis Biodiversity Center, Leiden, Netherlands
- RSMAS, Rosenstiel School of Marine, Atmospheric, and Earth Science, Miami, Florida, United States (= UMML, University of Miami Marine Lab)
- SAM, Iziko South African Museum, Cape Town, South Africa
- SAMA, South Australian Museum, North Terrace, Adelaide, Australia
- SBMNH, Santa Barbara Museum of Natural History, Santa Barbara, California, United States
- SMNH, Swedish Museum of Natural History, Stockholm, Sweden
- TMAG, Tasmanian Museum and Art Gallery, Hobart, Tasmania, Australia
- VSM, NTNU Museum of Natural History and Archaeology, Trondheim, Norway (formerly Det Kgl. Norske Videnskabers Selskab Museet)
- YPM, Peabody Museum of Natural History, Yale University, New Haven, Connecticut, United States
- ZMB, Zoologisches Museum, Museum für Naturkunde der Humboldt University of Berlin, Berlin, Germany
- ZMMGU, Zoological Museum, Lomonosov Moscow State University, Moscow, Russia
- ZMUB, Universitetet i Bergen, Bergen, Norway
- ZMUC, Kobenhavns Universitet, Zoologisk Museum, Copenhagen, Denmark
Specimen images
The following images relate to pre–20th century giant squid specimens and sightings. The number below each image corresponds to that given in the
-
#1 (c. 1546)
Japetus Steenstrup's comparison of a common squid (centre; its tentacles in an anatomically implausible position) with two 16th century drawings of the "sea monk of the Øresund"—Guillaume Rondelet's (1554:492, fig.) on the left and Pierre Belon's (1553:39, fig.) on the right (Steenstrup, 1855a:83, fig.). It has been noted that Steenstrup's reproductions differ somewhat from the 16th century originals.[298] -
#1 (c. 1546)
Three further 16th century depictions of the "sea monk of the Øresund", possibly a giant squid or an angelshark (Squatina squatina), from the works of (left to right): Stefan Hamer (1546:[1], fig.), Conrad Lycosthenes (1557:609, fig.), and Johannes Sluperius (1572:105, fig.), as collected in Paxton & Holland (2005:41, fig. 1). -
Watercolour of the sea monk by an anonymous artist, from the so-called Gessner Albums kept at the University of Amsterdam. These illustrations were collected by Felix Platter to serve as the basis for the woodcuts in Conrad Gessner's encyclopedic Historia animalium, the first modern zoological work to attempt to describe all known animal species (see Gessner, 1558:519, fig.for sea monk depiction published therein).
-
sea bishop (the latter also thought by some to be based on a giant squid[299]), issued in 1669 and based on Conrad Gessner's originals from 1558 (themselves based on Rondelet's).
-
cranchiid than it is to an architeuthid".[300]
-
#13 (?/12/1853)
Upper and lower beaks of the type specimen of Architeuthis monachus (centre) and assorted smaller squid species: Gonatus fabricii (top), Sthenoteuthis pteropus (middle), and Loligo forbesii (bottom) (Steenstrup, 1898:pl. 1) -
#13 (?/12/1853)
Holotype of Architeuthis monachus at the Zoological Museum in Copenhagen, as it appeared in 2015. It consists of a dried beak, the sole part of the animal that was preserved. -
#13 (?/12/1853)
Another view of the same specimen, showing areas where damage has been patched up with tape and repaired with sutures (see also original label and assorted beak fragments) -
#14 (5/11/1855)
Arm fragment and associated suckers of the type specimen of Architeuthis dux, collected off the Bahamas on 5 November 1855 (Steenstrup, 1898:pl. 3) -
#14 (5/11/1855)
Four jars containing soft tissue remains of the Architeuthis dux holotype, from the collections of the Zoological Museum in Copenhagen (as they appeared in 2013) -
#14 (5/11/1855)
Closeup of one of the jars containing the Architeuthis dux type material (see alternative view) -
#14 (5/11/1855)
Arm fragment of the Architeuthis dux holotype (see also detail of one of the suckers) -
#16 (≤1860)
Beak with associated buccal musculature, radula, and loose suckers of the type specimen of Loligo (later Architeuthis) hartingii, the provenance of which is unknown (Harting, 1860:pl. 1) -
#18 (30/11/1861)
The French corvette Alecton attempts to capture a giant squid off Tenerife on 30 November 1861. Reproduction of the original watercolour by officers of the Alecton, from Bourée (1912:115, fig. 108). -
#18 (30/11/1861)
An 1865 illustration of the Alecton incident by P. Lackerbauer, clearly based on the officers' watercolour. It appeared in the first edition of Alfred Moquin-Tandon's Le Monde de la Mer, which he published under the pseudonym "Alfred Frédol" (Frédol, 1865:314, pl. 13). -
#18 (30/11/1861)
Another depiction of the encounter, by Édouard Riou (credited in caption) and A. Etherington (signed), based on a sketch by ensign E. Rodolphe, an officer on the Alecton. This engraving appeared in Bouyer (1866:276, fig.) and subsequently featured in other publications, including an 1867 issue of the Dutch travel magazine De Aarde en haar Volken (from which the present image was extracted). -
#18 (30/11/1861)
This much-reproduced image appeared in Louis Figuier's La vie et les mœurs des animaux (Figuier, 1866:467, fig. 362; shown here) and Henry Lee's Sea Monsters Unmasked (Lee, 1883:39, fig. 8), among others, but Muntz (1995:21) wrote that its original source was uncertain. -
#18 (30/11/1861)
A wood engraving of the Alecton encounter published in 1868, the squid clearly based on the one from the more famous image that had earlier appeared in Figuier (1866) -
Twenty Thousand Leagues Under the Sea (1871), whose depiction of the giant squid was undoubtedly inspired by the Alecton encounter.[3] Bernard Heuvelmans's The Kraken and the Colossal Octopus incorrectly described this illustration as "[t]he Alecton squid after Arthur Mangin, 1864".[301]
-
#18 (30/11/1861)
An illustration of the Alecton encounter from Les Animaux Excentriques by Henri Coupin , first published in 1903, based on the original from Bouyer (here given as "Rouyer"). -
#[1] (1870?)
Anatomical details of the squid that washed ashore in Nova Scotia, Canada, around 1870. It was originally described as Architeuthis megaptera and later identified as Sthenoteuthis pteropus. Shown are the mantle (fig. 1), teeth of the radula and palate (figs. 2–7), and arm and tentacle suckers (figs. 8–9) (Verrill, 1880a:pl. 21). -
#[1] (1870?)
Preserved sucker (oral view) of the Architeuthis megaptera type specimen from Nova Scotia. Deposited at Yale's Peabody Museum of Natural History. -
#[1] (1870?)
Sucker of the same specimen in lateral view. Photographed in 2006 at the Peabody Museum of Natural History. -
J. H. Emerton) and upper and lower beaks of Verrill specimen No. 10 from 1873 or earlier (#26; top and bottom—drawn by A. E. Verrill) (Verrill, 1880a:pl. 18). A front view of the former is additionally shown in Verrill (1882c:pl. 11 fig. 3a).
-
Doryteuthis pealeii; fig. 7) and giant squid specimen #16, redrawn from Harting (1860:pl. 1 fig. 4)(fig. 8).
-
#25 (?/12/1872) and others
Verrill's sketches of the two halves of the lower mandible of specimen No. 4 from 1872 (#25), drawn from photographs (figs. 5–5a), as well as beaks and other remains redrawn from earlier sources: #16 from Harting (1860:pl. 1 figs. 1, 8, 8') (figs. 1–1b); Steenstrup's A. dux from Harting (1860:pl. 1 fig. 1A) (fig. 2); #13 after what was eventually published as Steenstrup (1898:pl. 1 fig. 2) (fig. 3); and "Enoploteuthis hartingii" from Harting (1860:pl. 2 fig. 15, pl. 3 figs. 23–24) (figs. 4–4b) (Verrill, 1880a:pl. 25) -
#26 (≤1873)
Upper beak extracted from a sperm whale stomach, from Packard (1873:93, fig. 10). A. E. Verrill considered the illustration inaccurate and published his own version. The much smaller beak (drawn to scale), possibly from the same whale, may belong to Gonatus fabricii, as suggested by Steenstrup.[302] -
#27 (≤1873)
The type specimen of Plectoteuthis grandis, consisting of a single arm (Owen, 1881:pl. 34). The provenance of this material is uncertain, but it may originate from the east coast of South America.[33] -
#29 (26/10/1873)
Victor Nehlig's 1881 impression of fisherman Theophilus Picot's encounter with a giant squid off Portugal Cove, Newfoundland, on 26 October 1873 (Rathbun, 1881:267, fig.; compare similar illustration by W. A. Cranston, another by Charles Livingston Bull, and this cover by Quinto Cenni) -
The Anniston Hot Blast ([Anon.], 1902b:6, fig.).
-
#29 (26/10/1873)
The 19-foot (5.8 m) tentacle of the first Portugal Cove specimen, hacked off a live animal on 26 October 1873 (Owen, 1881:pl. 33 fig. 2; see very similar but horizontally flipped woodcut from Harvey, 1874a:68, fig.) -
#29 (26/10/1873)
The Adventure of the Giant Squid, painted by N. C. Wyeth c. 1939 to illustrate Norman Duncan's The Adventures of Billy Topsail (1906)[228] as republished in Anthology of Children's Literature (1940).[303] The story includes a fictionalised encounter with a giant squid based on the widely reported first Portugal Cove specimen. -
The Anniston Hot Blast on 12 February 1902 ([Anon.], 1902b:6, fig.).
-
Logy Bay, photographed in Moses Harvey's home (the caudal fin is visible on the right). This original photograph appears not to have been published prior to Aldrich (1991:458, fig. 1B).
-
#30 (25/11?/1873)
The Logy Bay giant squid draped over Reverend Moses Harvey's sponge bath, November or December 1873. The original photographs (as opposed to line drawings thereof) were apparently not published until much later; the present one is taken from the April 1899 issue of The Wide World Magazine (Harvey, 1899:737, fig.), where it was captioned "Dr. Harvey's immense octopus [sic]". -
#30 (25/11?/1873)
Another very similar (though non-identical) photograph of the head and arms of the Logy Bay giant squid. This one appeared in Frank Walter Lane's Kingdom of the Octopus (Lane, 1957:pl. 44), where it was described as "the best-known photograph of a giant squid". -
#30 (25/11?/1873)
Line drawings taken from two photographs of the Logy Bay specimen (Verrill, 1880a:pl. 13). Note that the upper illustration is based on a slightly different frame to the preceding photo (as evidenced by the contrasting arrangement of arm tips on the lower left, which are closer to those seen in this version). -
A. E. Verrill's reconstruction of "Architeuthis Harveyi", the Logy Bay giant squid (Verrill, 1880a:pl. 14)
-
#30 (25/11?/1873)
Two-part beak and gladius fragments of the Logy Bay specimen (Verrill, 1880a:pl. 15) -
#30 (25/11?/1873)
Suckers of the arms and tentacles and closeups of the radula and its associated teeth (Verrill, 1880a:pl. 16a) -
#30 (25/11?/1873)
Extant remains of the Logy Bay specimen (preserved in 70% alcohol) as they appeared in 2015, at Yale's Peabody Museum of Natural History. -
#30 (25/11?/1873)
Limbs of the Logy Bay specimen in 2015, at the Peabody Museum of Natural History. -
#30 (25/11?/1873)
Radula as it appeared in 2006, at the Peabody Museum of Natural History (see also reverse side). -
A. E. Verrill in 1883 and was the world's first.[79]
-
#33 (2/11/1874)
The "calmar gigantesque" that washed ashore on Île Saint-Paul on 2 November 1874, as documented by geologist Charles Vélain during a French astronomical mission to Île Saint-Paul to observe the transit of Venus (Vélain, 1877:81, fig. 8). The illustration is based on a photograph,[28] one of the first ever taken of a giant squid. -
#33 (2/11/1874)
Simplified illustration from George Tryon's Manual of Conchology (Tryon, 1879b:pl. 82 fig. 378), highlighting the strange morphology, on which Tryon opined: "The truncated arms and the gradually attenuating body, with very narrow fins extending along each side for half its length, are noticeable peculiarities, as is also the web connecting the arms; it is questionable, however, how much the engraving may be relied upon".[200] -
J. H. Emerton's reconstruction of Architeuthis princeps, based chiefly on the Fortune Bay specimen, also known as Verrill specimen No. 13 (Verrill, 1881b:pl. 54 fig. 1). Identified as a drawing of the Catalina specimen (#42) in Richard Ellis's The Search for the Giant Squid.[304]
-
#34 (?/12/1874)
Beak of the Fortune Bay specimen (preserved in 70% alcohol) as it appeared in 2007. Deposited at the Peabody Museum of Natural History. -
#34 (?/12/1874) and #42 (24/9/1877)
Details of the horny ring of a large tentacular sucker of Verrill specimen No. 13 from 1874 (#34; fig. 11) and assorted arm and tentacular suckers of Verrill specimen No. 14 from 1877 (#42; remaining figures) (Verrill, 1880a:pl. 17) -
#37 (25/4/1875)
Beak and buccal mass of giant squid caught on 25 April 1875, off Inishbofin, Connemara, Ireland, as it appeared at the National Museum of Ireland – Natural History in 2019. -
Trinity Bay, Newfoundland, on 24 September 1877, from the 27 October 1877 issue of Canadian Illustrated News ([Anon.], 1877a:269, fig.)
-
#42 (24/9/1877)
Illustration of the "devil-fish" from Catalina, from the 3 November 1877 issue of Harper's Weekly ([Anon.], 1877b:868, fig.) -
The Penny Illustrated Paper and Illustrated Times ([Anon.], 1877c:305, fig.)
-
#42 (24/9/1877)
A further illustration of the Catalina specimen, showing the animal after it had died (see related figure), from the 9 June 1883 issue of the French popular science magazine La Nature (Oustalet, 1883:17, fig. 1; also reproduced in Heuvelmans, 2003:fig. 103). -
#42 (24/9/1877)
Drawing by A. E. Verrill of the Catalina specimen, here incorrectly identified as the Fortune Bay specimen (#34); see original illustration (Verrill, 1880a:pl. 20) -
#42 (24/9/1877)
Ventral views of the supposedly anteriorly-free caudal fins of the Catalina specimen (top: after several days in alcohol; bottom: after several months in same), now known to be a preservation artefact[305] (Verrill, 1880a:pl. 19) -
A. E. Verrill for the Smithsonian and based on the Catalina specimen, is seen hanging in the background.[306]
-
#42 (24/9/1877)
Another illustration of the American court at the International Fisheries Exhibition of 1883, with a giant squid model based on the Catalina specimen hanging overhead, from the fourth of a series of papers about the exhibition published in Science by American ichthyologist George Brown Goode (Goode, 1883:614, fig.). -
J. H. Emerton's giant squid and giant octopus models are suspended from the ceiling surrounded by fishing nets (see also engraving of same).
-
#42 (24/9/1877)
Hanging models of a giant squid (based on the Catalina specimen[306]) and a giant Pacific octopus, designed for the Tennessee Centennial and International Exposition of 1897, as they appeared in the Lower Invertebrates exhibit in the West Wing of the Smithsonian Institution Building c. 1901 (see also image from c. 1903). -
#42 (24/9/1877)
The original Star-Spangled Banner flag undergoing restoration work in 1914 in the West Wing of the Smithsonian Institution Building, under the hanging giant squid model, one of the few exhibits not removed for the occasion (see also photograph showing seamstresses at work under the guidance of Amelia Fowler) -
Ward's Natural Science Establishment of Rochester, New York, it was installed a year later.[308] It was destroyed during the Blitz in 1940;[309] the second and current model dates from 1974.[308]
-
Ward's models of the Catalina giant squid and a giant Pacific octopus, suspended from the ceiling of the American Museum of Natural History's Shell and Coral Hall, early 20th century. The squid was later moved to the Hall of Ocean Life, the Invertebrate Hall, and, following refurbishment, the "Wall of Life" in the Hall of Biodiversity (see 2015 image).[310] Both models were acquired in 1895 for a total of US$750 (equivalent to $27,468 in 2023).[311]
-
taxidermist and artist Wendy Christensen-Senk repainted it a "more passionate reddish maroon" from its previous "battleship gray".[312]
-
?#44 (21/11/1877)
Giant squid beached on the Newfoundland coast, apparently on 22 November 1877 (Heydebrand und der Lasa, 1887:72, fig.; closest is #44 from 21 November, from Smith's Sound). Two figures standing over the specimen, which are present in the original, have been removed in this version. Note the similarity to this illustration of the Catalina specimen. -
Thimble Tickle specimen" found on 2 November 1878, which reportedly measured 55 feet (17 m) in total length, from Charles Frederick Holder's Marvels of Animal Life (Holder, 1885:198, pl. 25). No photographs or exact measurements exist as the specimen was cut up for dog food soon after its discovery.
-
Giant Squid Interpretation Site, the sculpture is located close to the site of the original specimen's capture and has featured on a Canada Post stamp.[314]
-
Newfoundland, where the giant specimen of 1878 was found. It has long been regarded by Guinness World Records and its previous incarnations as the largest giant squid ever recorded.[315]
-
type description (Kirk, 1882:pl. 36 figs. 2–4)
-
#49 (?/9/1879)
Soft tissue remains of the 1879 specimen from Olafsfjord, Iceland, as they appeared in 2012 at the Zoological Museum in Copenhagen. The material is labelled as Architeuthis monachus, a taxon now considered synonymous with A. dux. -
M. Thomas P. Gilbertpointing to the preserved left tentacle of the Olafsfjord giant squid of 1879
-
#49 (?/9/1879)
Closeup of the same specimen, showing the enlarged suckers of the tentacular club. The morphology of this tentacle was studied by Roeleveld (2002). -
oesophagus, tentacular club, and arm fragments of the Grand Banks specimen found dead at the surface in April 1880 (#54), the first in which arm and sucker regeneration was documented (Verrill, 1881b:pl. 26). A single sucker (fig. 5) of the first Portugal Cove specimen (#29) is also shown.
-
#54 (?/4/1880)
Selected details of the Grand Banks specimen: 50th sucker of one of arms II (fig. 3), statolith in front and lateral aspects (fig. 4), and various views of the radular teeth (figs. 5–7) (Verrill, 1881b:pl. 38 figs. 3–7; see uncropped version) -
formal description of the latter (Kirk, 1882:pl. 36 fig. 1)
-
#60 (10/11/1881)
Illustration of the giant squid that was found in Portugal Cove, Newfoundland, on 10 November 1881 (Verrill's penultimate specimen—No. 27). Based on a photograph by E. Lyons of St. John's, it was published in Harper's Weekly for 10 December ([Anon.], 1881:821, fig.). -
type specimen in lateral aspect (Kirk, 1888:pl. 7). Note the extreme length of the feeding tentacles relative to the mantle and arms, which has been questioned by some authors.[nb 24]
-
#62 (?/10/1887)
Illustration of the A. longimanus specimen by T. W. Kirk, showing the posterior end of the mantle (including caudal fins), details of arm and tentacular suckers, and transverse sections of arms I–IV (Kirk, 1888:pl. 8) -
#62 (?/10/1887)
T. W. Kirk's final plate of A. longimanus, showing the two-part beak (both in situ and following extraction) and a single row of radular teeth (Kirk, 1888:pl. 9) -
#67 (4/2/1895)
Type specimen of Architeuthis japonica, caught in a net in Tokyo Bay on 4 February 1895 after a 2–3 day storm (Mitsukuri & Ikeda, 1895:pl. 10) -
#68 (18/7/1895)
Mantle measuring 46 cm originally recovered from sperm whale vomit, from a 1900 work by Louis Joubin (Joubin, 1900:pl. 15 fig. 8). This specimen is the holotype of Dubioteuthis physeteris. -
#68 (18/7/1895)
Gladius (internal shell remnant) measuring 39 cm extracted from the same specimen (Joubin, 1900:pl. 15 fig. 10) -
#68 (18/7/1895)
Detail of the dorsal anterior mantle of the Dubioteuthis physeteris holotype (Joubin, 1900:pl. 15 fig. 9) -
#[3] (18/7/1895)
Two-part beak of an undetermined squid (Joubin, 1900:pl. 14 figs. 1–2), identified as possibly Architeuthis by Joubin (1900) and as non-architeuthid by Pfeffer (1912) -
#69 (10/4/1896) and others
View from the other end of the corridor, showing the tentacles of the three more-or-less complete specimens and various other assorted giant squid parts (see also tentacular club being examined) -
#69 (10/4/1896)
Tentacular suckers of the largest specimen, examined in 2011, when the box containing the museum's six giant squid was reopened for maintenance, registration in the database, and to add new preservatives.[316] -
#69 (10/4/1896)
Beak of the same specimen, as it appeared during the 2011 inspection -
#69 (10/4/1896) or #70 (27/9/1896)
One of the two giant specimens from Kyrksæterøra, stretched out for measurement -
NTNU Museum of Natural History and Archaeology in 2011; lower-resolution versions originally appeared in Storm (1897:100, fig. 20) and Brinkmann (1916:179, fig. 2).
Notes
- ^ Verrill's marginal annotations read as follows: "Architeuthis monachus (No. 5) Logie Bay, N. Foundland about 1⁄8 natural size between 1⁄8 and 1⁄9. The tub is 38 1⁄2 inches [98 cm] in diameter and circular. Harvey (?) letter. Some of the suckers are broken off on the short arms. They alternate in two regular rows. On the club of the long arm there is a marginal row of small suckers on each side alternating with the large ones. One sucker gone on this long arm."[2]
- soup form on 3 March 1769, during James Cook's first voyage to the Pacific.[17] In 1783, Banks presented at the Royal Society an important early paper on ambergris by the London-based German physician Franz Xavier Schwediawer, in which was recounted an observation made by fishermen some ten years earlier of a sperm whale that had in its mouth an incomplete cephalopod tentacle nearly 27 ft (8.2 m) long.[18] To Schwediawer, this corroborated "the common saying of the fishermen, that the cuttle-fish is the largest fish of the ocean".[18] Schwediawer's footnote would be cited "uncritically" in subsequent works, such as Thomas Beale's influential The Natural History of the Sperm Whale of 1839.[17]
- ^ Ellis (1998a:86) described Verrill as someone with "an almost limitless capacity for work", who "began publishing papers on these specimens almost as fast as they came in". The full list of Verrill's publications on the Newfoundland strandings of 1870–1881 is as follows: Verrill 1874a, b, 1875a, b, c, 1876, 1877, 1878, 1880a, b, 1881a, b, 1882a, c.
- The Annals and Magazine of Natural History.[27] The giant squid found beached on Île Saint-Paul on 2 November 1874 (#33) was another early specimen to be photographed.[28] Perhaps the earliest of all was the beak of the October 1871 specimen (#22) from the Grand Banks of Newfoundland, whose photograph was taken some time after its discovery but already mentioned in February 1873 by Packard (1873:92).
- ^ According to Harvey's entry in the Dictionary of Canadian Biography, written by giant squid expert Frederick Aldrich, "[a]t least ten per cent of his essays deal with [the giant] squid".[31]
- ^ Unconfirmed mass appearances of giant squid include the claim by Frederick Aldrich that a "school of 60 has been sighted off the coast of Newfoundland"[39] (cf. #38). Richard Ellis noted that Aldrich never repeated this claim in print, "so it is likely that he learned it was not accurately reported".[40] However, Aldrich had earlier given more details in an interview with The Muse, where he stated that in autumn 1964 a Japanese trawler operating around 100 miles (160 km) off St. John's "went through a school of sixty or seventy of these animals frolicking on the surface of the water".[41] Aldrich also told Clyde Roper that "Grand Banks fishermen have reported seeing hundreds of giant squid bodies floating on the surface".[42]
- ^ Museum of Economic Fish Culture in South Kensington, London.[76] Its appearance was based on official papers and photographs Buckland received through the Colonial Office in December 1873 relating to both the first Portugal Cove specimen (#29) and the Logy Bay specimen (#30).[76] Buckland's cutout has been described as the "first attempt to show the public what a giant squid really looked like".[77]
- ^ See giant squid specimen #661, whose tank leaked around 1 tonne of formalin in February 2022, resulting in the month-long closure of the Ibaraki Nature Museum near Tokyo, Japan.
- Portugal Cove specimen of 1873 (#29)[87] and is close to that given for the total length of the Architeuthis longimanus holotype, which describer Thomas William Kirk variously cited as 55 ft 2 in (16.81 m) or 57 ft (17.4 m) and which the fisherman who found the specimen gave as 62 ft (18.9 m).[88]
- 524).
- ^ In particular, the commercially important Humboldt squid (Dosidicus gigas) is sometimes called 'giant squid' (or its Spanish equivalent calamar gigante) and this, together with its large size, has led to confusion with Architeuthis (e.g. misidentification #[2]). In February 2002, the Manhattan-based seafood restaurant Esca offered what was claimed to be Architeuthis, but the ammonia-free flesh turned out to be that of the Humboldt squid.[107]
- ^ One record given by Ellis is omitted from the present list: Ellis (1998a:259) lists a specimen supposedly collected "[n]orth of Bahamas" in 1898, citing Steenstrup (1898), but this appears to stem from confusion with the type specimen of Architeuthis dux (#14), collected off the Bahamas in 1855.
- Portugal Cove, Newfoundland, said to have been spotted in 1817 by a local minister named Larveige and named after him.[143]These records are likewise excluded due to a lack of substantiating evidence.
- Historia de Gentibus Septentrionalibus of the same year (later reprinted by authors such as Gessner, Edward Topsell, Ulisse Aldrovandi, and John Jonston);[154] and various mythological and folkloric creatures such as the Greek Scylla,[155] the Japanese Akkorokamui[156] and umibōzu,[157] the Māori Te Wheke-a-Muturangi,[158] and the Caribbean Lusca.[156] Giant squid have even, on occasion, been proposed as an explanation for the Loch Ness Monster,[159] with the obvious problem that all living cephalopods are exclusively marine.[160] Some authors have cautioned against attributing "sea monster" sightings to giant squid and offered alternative explanations.[161]
- ^ Early sources variously give the date as 1545,[170] 1546,[171] December 1549,[172] or 1550.[173] Lycosthenes (1557) mentions three "sea monks" supposedly found in 1530, 1546, and 1549, the first in the Rhine and the latter two near Copenhagen (possibly referring to the same specimen). The earliest known source, a German woodcut of the creature dating from 1546[174]—according to which the event happened the same year—places an upper bound on the date, and Paxton & Holland (2005) consider 1546 to be the most likely year.
- ^ Mentioned by Steenstrup (1855a:78), presumably referring to Jónsson's encyclopedic Annálar Björns á Skarðsá, covering the history of Iceland from 1400 to 1645, which was eventually published as a dual Icelandic–Latin work in 1774–1775.[178]
- Cyclopterus lumpus]. No trace was seen of the head, except the one aperture, or two, which were found behind the tails or at a short distance from them. This very creature was observed by many trustworthy men, and one of the tails of the sea monster was brought home to Thingørekloster for examination."[184]
- ^ In a 2013 article about the giant squid, Mark Dery wrote: "contemporary teuthologists dismiss the "attack" as the death throes of a moribund animal, pointing out that virtually all giant squid encountered on the ocean's surface are dead or dying. "There is not a single corroborated story of a [giant] squid attacking a man, a boat, or a submersible", asserts Ellis."[222]
- ^ As part of his research for The Kraken, Reed travelled to Newfoundland and spoke to Picot's descendants and to Margueritte Aldrich, widow of giant squid expert Frederick Aldrich.[230]
- Thimble Tickle specimen" is sometimes mistakenly cited as 57 ft (17.4 m).[116] Over time, various other superlative measurements have been attributed to the specimen, including a mass of 2 tonnes[247] or exactly 4,480 lb (2,030 kg);[248] an eye diameter of 40 cm,[247] 18 in (46 cm),[249] or 9 in (23 cm);[250] and suckers 4 in (10 cm) across.[249] Wildly excessive mass estimates for the specimen have included:[251] 29.25 or 30 short tons (26.5 or 27.2 tonnes);[252] near 24 tonnes;[253] less than 8 tonnes;[254] and 2.8 or "more realistic[ally]" 2 tonnes.[255]
- ^ The identity of the two other fishermen in the boat is not given in Moses Harvey's original account,[246] but later sources identify Joseph Martin, the founder of Thimble Tickle (now known as Glovers Harbour), as one of the fishermen involved in the squid's capture.[256] George Marsh and Henry Rowsell—the founders of the nearby settlements of Winter House Cove and Lock's Harbour (Lockesporte), respectively—have also been suggested as participants.[257] The fishermen may have learned of Moses Harvey's interest in the giant squid when the latter visited Notre Dame Bay only a couple of months earlier, in August 1878, as part of a geological survey.[258] A CBC News report broadcast in 2004 features Maurice Martin, great-great-grandson of Joseph Martin, recounting the story of the squid's capture as told to him by his grandfather.[259]
- Newfoundland Banks may have a body length of 8 feet [2.4 m] and measure up to 40 feet [12 m] overall."[262]
- ^ Paxton countered that this was possibly attributable to shrinkage of the gladius prior to measurement, or to it having been severed in several places beforehand,[94] but Kirk wrote that it was measured "when first extracted" and only later "shr[a]nk considerably".[269]
- ^ a b c Kirk (1888:38) provides a table with a detailed breakdown of the specimen's various measurements. There is, however, a discrepancy between the total length of 684 in (17.37 m, or exactly 57 ft) given in the table—which agrees with the individual values of 71 in (1.80 m) for the mantle, 22 in (0.56 m) for the head, and 591 in (15.01 m) for the tentacles—and the total length of 55 ft 2 in (16.81 m) given by Kirk in the body of the article. Wood (1982:191) suggested that, due to the tentacles' highly retractile nature, the total length of 62 feet (18.9 m) originally reported by the fisherman "may have been correct at the time he found the squid", and that "[t]his probably also explains the discrepancy in Kirk's figures". Owing to its small mantle size, Wood (1982:191) estimated that "this specimen probably weighed less than 300 lb [140 kg]". O'Shea & Bolstad (2008) opined that the reported total length of 55 ft 2 in (16.81 m) "simply cannot be correct" and attributed it to either "imagination" or artificial lengthening of the tentacles. They added that a female giant squid with a mantle length of 71 in (180 cm) "measured post mortem and relaxed (by modern standards) today would have a total length of ≈32 feet [9.8 m]". Paxton (2016a:86) wrote that this specimen "clearly has the largest ratio of TL to ML [total length to mantle length] ever known in Architeuthis [...] which led [O'Shea & Bolstad, 2008] to suggest that the length was paced out and/or there was extensive post-mortem stretching. However, a re-reading of the original paper suggests that the specimen, although initially paced out, was actually measured, nevertheless the TL is at the edge of the 99.9% prediction interval range [...] and so it was certainly an unusual specimen."
- folklorist Herbert Halpert).[137] Aldrich restated this belief in "Monsters of the Deep", the second episode of the 1980 television series Arthur C. Clarke's Mysterious World: "I believe that the giant squid reach an approximate maximum size of something like a hundred and fifty feet [46 m]".[291] Richard Ellis, apparently unaware of Aldrich's similar statements in print, commented: "It is difficult to imagine why Aldrich would have made such an irresponsible statement, unless it had to do with being on camera".[272] Arthur C. Clarke himself wrote in 1992 that "the evidence suggests, they [giant squid] grow up to 150 feet in length".[292] The published literature includes many other such extreme size estimates based only on supposed giant squid suckers or scars thereof.
References
Short citations
- ^ Frank, 2014:246; Offord, 2016
- ^ Aldrich, 1991:459
- ^ a b c d Aldrich, 1978; Ellis, 1998a:79, 2005:143–144
- ^ Rees, 1949; Salvador & Tomotani, 2014; Hogenboom, 2014
- ^ Steenstrup, 1857:183; validated in Harting, 1860:11
- ^ Kent, 1874c:116–117; Paxton, 2018:56
- ^ Mitchill, 1813, 1815; see also response from Lee, 1819
- ^ Smith, 1842a:73, b:85; Earle, 1977:20
- ^ Steenstrup, 1849:952/[10]
- ^ Schwediawer, 1783:236; Ellis, 1998a:5, 2005:148
- ^ Ellis, 1998a:5, 133–134; King, 2019:142
- ^ Ellis, 1998a:133
- ^ see e.g. Earle, 1977:20–21
- ^ Heuvelmans, 1968
- ^ de Montfort, 1801; see Freeman, 2017:55
- ^ Paxton, 2018:56–57
- ^ a b Paxton, 2018:56
- ^ a b Schwediawer, 1783:236–237; Paxton, 2018:56
- ^ Steenstrup, 1857:184; validated in Harting, 1860:11
- ^ Bouyer, 1861
- ^ Harvey, 1874a
- ^ Lee, 1875:111; Earle, 1977; Muntz, 1995; McConvey, 2015b
- ^ Coe, 1929:36; G.E. Verrill, 1958:69
- ^ Murray, 1874b:121
- ^ a b c Offord, 2016; Keartes, 2016
- ^ Aldrich, 1991:458
- ^ Harvey, 1874a:68; Verrill, 1875a:34
- ^ a b c Wright, 1878:329
- ^ Aldrich, 1987:109; Frank, 2014
- ^ Hatton & Harvey, 1883:238–243; Aldrich, 1987:115, 1994
- ^ Aldrich, 1994
- ^ Ellis, 1994a:379, 1998a:257; Sweeney & Roper, 2001:[27]
- ^ a b c Lee, 1875:114
- ^ Nesis, 2001
- ^ a b Sweeney & Young, 2003
- ^ Aldrich, 1991:474; Förch, 1998:93; Winkelmann et al., 2013; Guerra et al., 2013
- ^ Downer, 1965:8; Aldrich, 1967a, 1968a
- ^ Stevens, 1988:158–160
- ^ Aldrich, 1967b
- ^ Ellis, 1998a:241
- ^ Downer, 1965:8
- ^ a b c Roper & Shea, 2013:111
- ^ Kubodera et al., 2016; see also Sakamoto, 2014
- ^ Ellis, 1998a:211
- ^ Oreskes, 2003:716, 2014:29
- ^ Baird, 2002; Kubodera, 2010
- ^ [NHK], 2013a, b; Dery, 2013
- ^ Hann, 2006; Guerra et al., 2011:1990; Walker, 2011
- ^ Okutani, 2015
- ^ Nesis, 2003; Coro et al., 2015
- ^ [NMNH], 2004
- ^ Guerra et al., 2006; Fernández-Pello García, N.d.
- ^ a b Kubodera et al., 2016
- ^ O'Shea, 1997, 1999
- ^ Roper & Shea, 2013:111; Roper, 2016
- ^ Roper et al., 2015
- ^ Roper & Jereb, 2010:121; Roper, 2016
- ^ see map in Sivertsen, 1955:11
- ^ Guerra et al., 2006
- ^ Turner, 1963:23
- ^ Ellis, 1994a:133
- ^ Guerra et al., 2011:1991
- ^ Ellis, 1998a:242; see Loates, 1989
- ^ Ellis, 1998a:242–243
- ^ Miller, 1983; Landman & Ellis, 1998; Landman et al., 1999; Roper & Jereb, 2010:123; Ablett, 2012
- ^ [Anon.], 1988:4, 1989:7; Ellis, 1998a:46; Lien, 2000:278; Grann, 2004; see also [NTA], 1966 and identically worded magazine appeal in Aldrich, 1968b
- ^ [Anon.], 1971
- ^ Guerra & Segonzac, 2014:118
- ^ [Anon.], 2014a, b
- ^ Shimada et al., 2017:9
- ^ [NMNH], 2004; [SITES], 2004
- ^ [Anon.], 2004b; Miller, 2004
- ^ Ellis, 1997a:52
- ^ Emerton, 1883; Tratz, 1973; Ellis, 1997a, b, 1998a:214; Wechsler, 1999
- ^ a b Ellis, 1997a:31, 48
- ^ a b Buckland, 1875:211–216, 1886:400; Ellis, 1997a:31
- ^ Ellis, 1997a:31
- ^ a b Ellis, 1997a:34, 46
- ^ a b Ellis, 1997a:35
- ^ Emerton, 1883; Ellis, 1997a:35–36
- ^ Ellis, 1997a:44–46
- ^ [Anon.], 1966; Ellis, 1997a:34–35; [Anon.], 2004a
- ^ [Anon.], 1969
- ^ Drahl, 2008
- ^ [Anon.], 2010
- ^ Wood, 1982:188–193; Ellis, 1998a:6–7, b; Paxton, 2016a, b; Bittel, 2016; Romanov et al., 2017
- ^ Murray, 1874a:161, 1874b:121
- ^ Kirk, 1888:35, 38
- ^ Wood, 1982:189, 191; Ellis, 1998a:6–7; O'Shea & Bolstad, 2008; Paxton, 2016a
- ^ O'Shea & Bolstad, 2008; Dery, 2013; Roper & Shea, 2013:113; Hanlon & Messenger, 2018:267; see also Nesis, 2001
- ^ Cerullo & Roper, 2012:22
- ^ a b O'Shea, 2003b
- ^ Paxton, 2016a, b; [Anon.], 2016
- ^ a b c d e Greshko, 2016
- ^ McClain et al., 2015:Table 3
- ^ Roper & Jereb, 2010:121
- ^ a b O'Shea & Bolstad, 2008
- ^ a b Nilsson et al., 2012:683; Land & Nilsson, 2012:86
- ^ Robson, 1933:681
- ^ Ellis, 1998a:73; Norman, 2000:150; Roper & Jereb, 2010:121
- ^ Glaubrecht & Salcedo-Vargas, 2004:62
- ^ Roper & Jereb, 2010:123
- ^ see Roeleveld, 2002
- ^ [UCPH], 2013
- ^ see Ellis, 1998a; Salcedo-Vargas, 1999; Glaubrecht & Salcedo-Vargas, 2004
- ^ Robson, 1933:681; see e.g. Robson, 1929 by the same author
- ^ Ellis, 2005:198
- ^ a b Sweeney & Roper, 2001
- ^ Ellis, 1994a:379–384, 1998a:257–265
- ^ Ellis, 1994a:ix
- ^ [Anon.], 1999:109; Roper et al., 2015:78
- ^ e.g. Leite et al., 2016; Funaki, 2017; Romanov et al., 2017; Shimada et al., 2017; Guerra et al., 2018
- ^ Starkey, 1963; Welfare & Fairley, 1980:74; Bright, 1989:148; Ellis, 1998a:204; Paxton, 2016a:83; dramatised in Packham, 1998
- ^ Ellis, 1998a:245; Paxton, 2016a:83
- ^ Cousteau & Diolé, 1973:205; Ellis, 1998a:208, 2005:145
- ^ a b Paxton, 2016a:83
- ^ McDowall, 1998; Ellis, 1998a:248
- ^ Revkin, 2013
- ^ Hajicek, 2007, 2008; Cassell, 2007
- ^ Taylor, 1932:1; [Anon.], 1938a:21; originally reported in East Anglian Daily Times, 31 October 1896; see also Herrington, 1932:60
- ^ [Anon.], 1938a:21; [Anon.], 1938b:83; originally reported in East Anglian Daily Times, 22–25 October 1938
- ^ see Ellis, 1999
- ^ Heuvelmans, 1968:68; Berger, 2009:275
- ^ Hoff, 2003:84, 96
- ^ Aldrich, 1979:67; Welfare & Fairley, 1980:72, 74; Fitzgerald, 2018
- ^ a b Aldrich, 1979:67
- ^ Bullen, 1898:139; Ellis, 1998a:136–137, 180–181
- ^ Aldrich, 1977
- ^ [Anon.], 1988:4; Aldrich, 1990c
- ^ Lane, 1957:196; Gantès, 1979:38; Flynn & Weigall, 1980; Welfare & Fairley, 1980:71; Goss, 1985:39; Heuvelmans, 2003:252; Berger, 2009:261, 276; Dyer, 2021
- ^ LeBlond & Sibert, 1973:11, 32; Bright, 1989:140; Ellis, 1998a:202
- ^ a b Verrill, 1897:79; repeated by Bartsch, 1937:403
- ^ Pierce et al., 1995
- Chilean Blob); Evon, 2017
- ^ Ley, 1959:210
- ^ Dozier, 1976:72; Ellis, 1998a:7
- ^ a b c Aldrich, 1980:59
- ^ de Montfort, 1801
- ^ Freeman, 2017:55
- ^ a b [Anon.], 1933
- ^ Glaubrecht & Salcedo-Vargas, 2004:64
- ^ e.g. Labails, 2009:24
- ^ Bonnell, 1963:6
- ^ Gould, 1886; Gibson, 1887; Aldrich, 1983; Ellis, 1994a, 1998a:10–30
- ^ Lee, 1875, 1883; Buel, 1887:81; Bullen, 1898:139; Allan, 1955a, b; Heuvelmans, 1958; Aldrich & Sullivan, 1978; Garcin & Raynal, 2011; Salvador & Tomotani, 2014; Walker, 2016; Naish, 2017
- ^ Buckland, 1886:399; Oudemans, 1892; Bartsch, 1917:364, 1931:347, 1937:403; Aldrich & Sullivan, 1978; Aldrich, 1979:67; Ellis, 1994b
- ^ Lee, 1883:64; Ellis, 1998a:15; but see Paxton et al., 2005
- ^ [Anon.], 1849:264; Myklebust, 1946; Ellis, 1998a:20
- ^ Lee, 1883:79; Ellis, 1998a:20; Switek, 2011; but see Galbreath, 2015:42
- ^ Carrington, 1957:35
- ^ Carrington, 1957:36; Paxton, 2021:314–315
- ^ Stead, 1933; Ellis, 1998a:28; Dalton, 2021
- ^ Ellis, 1998a:13–14
- ^ Ellis, 1998a:12, 14–15
- ^ Ley, 1941; Aldrich & Sullivan, 1978; Aldrich, 1990b; Ellis, 1998a:10–11; Lotzof, 2021; Guerra, 2022
- ^ a b Lotzof, 2021
- ^ Greener, 2010
- ^ Kirk, 1880:310; Lotzof, 2021
- ^ e.g. [Anon.], 1934
- ^ Paxton & Shine, 2016:27, 32
- ^ Paxton, 2003; France, 2016a, b, 2017
- ^ Lee, 2014; Mikkelson, 2014; Evon, 2016b; see also Ellis, 1994a:163, and April Fools' Day joke of Carstens, 2016, taken seriously by [Anon.], 2020
- ^ e.g. Evon, 2016a, 2017; Howard, 2017
- ^ Gerhardt, 1966:171; Muntz, 1995; Ellis, 1998a:11–12; Guerra, 2022; see Aristotle, c. 350 BC; Pliny, AD 77–79
- ^ [Anon.], 2000
- ^ Roper et al., 2015:78
- ^ Ellis, 1994a:379
- ^ Ellis, 1998a:5, 257; Sweeney & Roper, 2001:[27]; Guerra et al., 2004a:428; but see Paxton & Holland, 2005
- ^ Gudger, 1953:199; Lane, 1957:129; Ellis, 1994a:144
- ^ Vedel, 1575
- ^ Hamer, 1546; Rondelet, 1554; Gessner, 1558
- ^ Holberg, 1733
- ^ Huitfeldt, [1595]; Stephanius, [c. 1650]
- ^ Hamer, 1546
- ^ Muus, 1959:170
- ^ Carrington, 1957:63
- ^ Paxton & Holland, 2005:39
- ^ Jónsson, [c. 1645]; see Paxton & Holland, 2005:40
- ^ Roeleveld, N.d.
- ^ Barber & Riches, 1971:26; Aldrich, 1980:57
- ^ Carrington, 1957:63; Russell & Russell, 1975:97
- ^ Steenstrup, 1849:950–951/[9]
- ^ Packard, 1873:87
- ^ Steenstrup, 1849:950–952/[9–10]; for the original Icelandic text, see there and Jónsson, [c. 1645]:238
- ^ Steenstrup, 1849:952/[10–11]
- ^ Verrill, 1875c:214
- ^ a b c Tryon, 1879b:185
- ^ Hooke et al., c. 1674
- ^ see More, 1875a
- ^ [Anon.], c. 1673
- ^ see Sueur-Hermel, 2017
- ^ a b c Kristensen & Knudsen, 1983:222
- ^ Robson, 1933:690
- ^ from unpublished manuscript by Steenstrup cited in Verrill, 1875b:84
- ^ a b c [NHMD], 2019
- ^ Steenstrup
- ^ Verrill, 1875b:84 citing Harting, 1860
- ^ Tambs-Lyche, 1946:288
- ^ Harting, 1860
- ^ a b c d e Tryon, 1879b:184
- ^ a b c d Verrill, 1880a
- ^ a b Dell, 1970:27
- ^ a b Stephen, 1962:154
- ^ Crosse & Fischer, 1862:138
- ^ Dunning, 1998:428
- ^ MacAlaster, 1977:14
- ^ a b c Earle, 1977:53
- ^ Verrill, 1874a:158
- ^ a b Sweeney & Roper, 2001:[9]
- ^ a b Verrill, 1874a
- ^ Verrill, 1875b
- ^ J.E. Gray cited in Lee, 1875:114; see also Kent, 1874a:179
- ^ Kent, 1874a:178
- ^ Verrill, 1881b:401
- ^ Pfeffer, 1912:31
- ^ Owen, 1881:163
- ^ Glaubrecht & Salcedo-Vargas, 2000:276
- ^ a b Verrill, 1875a:34
- ^ Verrill, 1880a:181
- ^ Verrill, 1875b:79
- ^ Buckland, 1875:212–214
- ^ a b Dery, 2013
- ^ Buckland, 1875:213
- ^ Maunder, 1991
- ^ Hatton & Harvey, 1883:239
- ^ Agassiz, 1874
- ^ Packham, 1998
- ^ a b Duncan, 1906:34; Ellis, 1998a:82
- ^ Reed, 1995; Ellis, 1998a:82, 187
- ^ Ellis, 1998a:187
- ^ Ellis, 1998a:186
- ^ [Anon.], 1966; Ellis, 1997a:35
- ^ [Anon.], 2004a
- ^ Frank, 2014
- ^ Ellis, 1998a:201
- ^ Aldrich, 1978; Aldrich, 1990a:5
- ^ Reed, 1995; Ellis, 1998a:187
- ^ Lu et al., 1995
- ^ Sweeney & Roper, 2001:[12]
- ^ O'Connor, 1875
- ^ a b Nunn & Holmes, 2008
- ^ a b Pfeffer, 1912:32
- ^ a b Lau, 2021
- ^ a b [OM], 2021
- ^ Copedo, 2022
- ^ a b Verrill, 1880a:191, b:285
- ^ a b Ganeri, 1990:24; [Anon.], c. 2001
- ^ a b McWhirter & McWhirter, 1971:88
- ^ a b [Anon.], 1968
- ^ McWhirter & McWhirter, 1968:59, 1971:88
- ^ see Wood, 1982:190; Ellis, 1998a:107
- ^ MacGinitie & MacGinitie, 1949
- ^ Heuvelmans, 1958
- ^ MacGinitie & MacGinitie, 1968
- ^ Akimushkin, 1963; Wood, 1982:190
- ^ [Anon.], c. 2001; Harvey, 2004; Martin & Benoit, 2013; Marsh, 2016
- ^ Marsh, 2016
- ^ [Anon.], c. 2001
- ^ Harvey, 2004
- ^ McWhirter & McWhirter, 1968:59
- ^ e.g. Wood, 1982:189; Carwardine, 1995:240; Glenday, 2014:62; but see #121 for an exception
- ^ McWhirter & Page, 1956:101
- ^ Hickey, 2009
- ^ Hickey, 2010; [Anon.], N.d.
- ^ McConvey, 2015a
- ^ Verrill, 1882d:477
- ^ a b c d Marshall, 1996:45
- ^ Stevens, 1980:213, 1988:149–150
- ^ Kirk, 1880:312
- ^ a b c Marshall, 1996:46
- ^ see Dell, 1970:28
- ^ a b Ellis, 1998a:7
- ^ Sweeney & Roper, 2001:[87]
- ^ Joubin, 1895:34
- ^ [Anon.], 1890:190
- ^ a b Pfeffer, 1912:27
- ^ Martens, 1894
- ^ Kilias, 1967:491
- ^ Glaubrecht & Salcedo-Vargas, 2004
- ^ Kilias, 1993
- ^ a b Voss, 1956:136
- ^ a b Belloc, 1950:6; listed incorrectly therein as station 558, per Sweeney & Roper, 2001:[33]; Sweeney, 2017:[105]
- ^ a b Eivindsen, 2011
- ^ Roeleveld, 2002:727
- ^ after Voss, 1998:104; Sweeney & Roper, 2001:[5]; Sweeney & Young, 2003; Roper et al., 2015:82; Sweeney, 2017
- ^ Gervais, 1875:93
- ^ Lipiński et al., 2000:106
- ^ Nesis, 1987:218
- ^ a b Pfeffer, 1912:2
- ^ Owen, 1881:156
- ^ Flynn & Weigall, 1980; Ellis, 1998a:6–7
- ^ Clarke, 1992:72
- ^ Verrill, 1880a:240
- ^ per unpublished notes by S.S. Berry at NMNH, cited by Sweeney, 2017:[194]
- ^ a b c d Förch, 1998:89
- ^ a b Kristensen & Knudsen, 1983:223
- ^ Lu et al., 1995:324
- ^ Paxton & Holland, 2005:41
- ^ see Barber & Riches, 1971:26; Aldrich, 1980:57
- ^ O'Shea, 2003d
- ^ Heuvelmans, 2003:fig. 100
- ^ Packard, 1873:94
- ^ Johnson & Scott, 1940; Ellis, 1998a:82
- ^ Ellis, 1998a:89, fig.
- ^ Förch, 1998:22–23
- ^ a b Ellis, 1997a:36
- ^ Snell & Tucker, 2003:43
- ^ a b Ellis, 1997a:44
- ^ Ellis, 1997a:44; see also Snell & Tucker, 2003:97
- ^ Ellis, 1998c
- ^ Ellis, 1997a:41
- ^ Ellis, 1997a:42
- ^ Harvey, 2004; [Anon.], N.d.
- ^ Hickey, 2009, 2010
- ^ see e.g. Wood, 1982:189; Carwardine, 1995:240; Glenday, 2014:62
- ^ Eivindsen, 2011; Lervik, 2011
- ^ Bullen, 1898:ill. facing p. 143
- ^ Bartsch, 1917:364–366, 1931:347–349, 1937:403
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