2021 in paleoichthyology

Source: Wikipedia, the free encyclopedia.
List of years in paleoichthyology
In paleontology
2018
2019
2020
2021
2022
2023
2024
In paleobotany
2018
2019
2020
2021
2022
2023
2024
In arthropod paleontology
2018
2019
2020
2021
2022
2023
2024
In paleoentomology
2018
2019
2020
2021
2022
2023
2024
In paleomalacology
2018
2019
2020
2021
2022
2023
2024
In reptile paleontology
2018
2019
2020
2021
2022
2023
2024
In archosaur paleontology
2018
2019
2020
2021
2022
2023
2024
In mammal paleontology
2018
2019
2020
2021
2022
2023
2024

This list of

paleoichthyology
that occurred in 2021.

Jawless vertebrates

Name Novelty Status Authors Age Type locality Location Notes Images

Falxcornus[1]

Gen. et sp. nov

In press

Meng & Gai

Devonian (Lochkovian)

Xishancun Formation

 China

A member of Galeaspida belonging to the family Tridensaspidae. Genus includes new species F. liui.

Hongshanaspis[2]

Gen. et sp. nov

Valid

Liu et al.

Silurian (Telychian)

Qingshui Formation

 China

A member of Galeaspida belonging to the family Hanyangaspidae. Genus includes new species H. inexpectatus.

Jiangxialepis[3]

Gen. et sp. nov

Valid

Liu et al.

Silurian (Telychian)

Fentou Formation

 China

A member of Galeaspida belonging to the group Eugaleaspidiformes. The type species is J. retrospina.

Qushiaspis[4]

Gen. et sp. nov

Valid

Jiang et al.

Early Devonian

Xujiachong Formation

 China

A member of Galeaspida. Genus includes new species Q. elaia.

Jawless fishes research

  • A study on the phylogenetic relationships of cyathaspidids is published by Elliott, Lassiter & Blieck (2021).[5]
  • Miyashita et al. (2021) report larval and juvenile forms of four stem lampreys from the Paleozoic era (Hardistiella, Mayomyzon, Pipiscius and Priscomyzon), including a hatchling-to-adult growth series of Priscomyzon, and report that the studied larvae display features that are otherwise unique to adult modern lampreys, and lack the defining traits of ammocoetes.[6]
  • A study on the anatomy and likely feeding ecology of Mesomyzon mengae, based on data from new, well-preserved specimens, is published by Wu, Chang & Janvier (2021).[7]
  • A study on the histology of the dermal skeleton in Procephalaspis oeselensis, Aestiaspis viitaensis, Dartmuthia gemmifera and four species of Tremataspis is published by Bremer et al. (2021), who interpret their findings as indicative of the emergence of the complex pore-canal system in Tremataspis through the modification of the structures already present in other taxa.[8]
  • A study aiming to determine whether the earliest vertebrates may have swum under various conditions without a clearly-differentiated tail fin, based on data from an abstracted model of Metaspriggina walcotti, is published by Rival, Yang & Caron (2021).[9]
  • A study on the morphological and functional diversity of osteostracan and galeaspid headshields, and on its implications for the knowledge of the ecology of the immediate jawless relatives of jawed vertebrates, is published by Ferrón et al. (2021).[10]
  • Redescription of Nochelaspis maeandrine is published by Meng, Zhu & Gai (2021).[11]
  • A study on the anatomy of a dorsal head shield of Kalanaspis delectabilis is published by Tinn et al. (2021).[12]

Placoderms

Name Novelty Status Authors Age Type locality Country Notes Images

Bianchengichthys[13]

Gen. et sp. nov

Valid

Li, Zhu & Zhu in Li et al.

Silurian (Ludfordian)

Xiaoxi Formation

 China

A placoderm closely related to the last common ancestor of bony and cartilaginous fishes. The type species is B. micros.

Leptodontichthys[14]

Gen. et sp. nov

Jobbins et al.

Devonian (Givetian)

Taboumakhlouf Formation

 Morocco

A member of Arthrodira belonging to the family Plourdosteidae. The type species is L. ziregensis.

Placoderm research

  • Zhu et al. (2021) use CT scanning to reveal the endocast of Brindabellaspis stensioi, and evaluate the implications of its anatomy for the knowledge of the phylogenetic relationships of early jawed vertebrates.[15]
  • Redescription of the anatomy of the headshield of Parayunnanolepis xitunensis is published by Wang & Zhu (2021).[16]
  • Description of new fossil material of Palaeacanthaspis vasta from the Devonian (Lochkovian) Chortkiv Formation (Ukraine), and a study on the phylogenetic relationships of this species, is published by Dupret et al. (2021).[17]

Acanthodians

Name Novelty Status Authors Age Type locality Country Notes Images

Dobunnacanthus[18]

Gen. et comb. nov

Valid

Dearden et al.

Devonian

 United Kingdom

A new genus for "Vernicomacanthus" waynensis Miles (1973)

Nostolepis digitus[19]

Sp. nov

Valid

Li et al.

Devonian (Lochkovian)

Xitun Formation

 China

Nostolepis qujingensis[19]

Sp. nov

Valid

Li et al.

Devonian (Lochkovian)

Xitun Formation

 China

Acanthodian research

  • A study on the development of teeth in acanthodians, and on its implications for the knowledge of the evolution of teeth of jawed vertebrates, is published by Rücklin et al. (2021).[20]
  • A study on the anatomy of teeth, jaws and associated oral structures of acanthodians, and on their implications for the knowledge of the evolution of dentition of modern cartilaginous fishes, is published by Dearden & Giles (2021).[21]

Cartilaginous fishes

Name Novelty Status Authors Age Type locality Location Notes Images

Aquilolamna[22]

Gen. et sp. nov

Vullo et al.

Late Cretaceous (Turonian)

Agua Nueva Formation

 Mexico

A probable planktivorous shark placed in the new family Aquilolamnidae, of uncertain placement. Possibly a member of Lamniformes. The type species is A. milarcae.

Carcharhinus dudoni[23]

Sp. nov

Valid

Canevet & Lebrun

Miocene

 France

A species of Carcharhinus.

Carcharhinus pedronii[23]

Sp. nov

Valid

Canevet & Lebrun

Miocene

 France

A species of Carcharhinus.

Carcharhinus tingae[24]

Sp. nov

Valid

Cicimurri & Ebersole

Eocene (Bartonian)

 United States
( Louisiana)

A species of Carcharhinus.

Cladodus gailensis[25]

Sp. nov

Valid

Feichtinger et al.

Carboniferous (Serpukhovian)

 Austria

Dracopristis[26]

Gen. et sp. nov

Valid

Hodnett et al.

Late Carboniferous (Kasimovian
)

Atrasado Formation

 United States
( New Mexico)

A medium-sized

ctenacanthiform
shark known from a complete skeleton with soft tissue. The type species is D. hoffmanorum.

Durnonovariaodus[27]

Gen. et sp. nov

Valid

Stumpf et al.

Late Jurassic (Tithonian)

Kimmeridge Clay

 United Kingdom

A member of the family

Hybodontidae
. The type species is D. maiseyi.

Favusodus[28]

Gen. et sp. nov

Valid

Li et al.

Triassic (LadinianCarnian)

 China

A member of

Euselachii
. Genus includes new species F. orientalis.

Junggarensis[29]

Gen. et sp. nov

Valid

Roelofs et al.

Devonian (Famennian)

 Mongolia

Genus includes new species J. ambiguus.

Keichouodus[28]

Gen. et sp. nov

Valid

Li et al.

Triassic (Ladinian–Carnian)

 China

A member of Euselachii. Genus includes new species K. nimaiguensis.

Keuperodus[30]

Gen. et comb. nov

Valid

Ivanov, Duffin & Richter

Late Triassic (Carnian)

Arden Sandstone Formation

Grabfeld Formation

 Germany
 United Kingdom

A member of the family Jalodontidae. Genus includes "Phoebodus" brodiei Woodward (1893) (interpreted by Ivanov, Duffin & Richter, 2021 as a senior synonym of "Phoebodus" keuperinus Seilacher, 1948).

Lilamna[31]

Gen. et comb. nov

Valid

Greenfield

late Eocene or Cretaceous (uncertain)

 China

A possible member of the family Pseudoscapanorhynchidae. The type species is "Archaeolamna" apophysata Li (1997).

Maiseyacanthus[32]

Gen. et sp. nov

Valid

Bronson

Carboniferous (Mississippian)

Fayetteville Shale

 United States
( Arkansas)

A symmoriiform. The type species is M. ozarkanum.

Maiseyodus[33]

Gen. et comb. nov

Valid

Long et al.

Devonian (Emsian)

Cravens Peak Beds

 Australia

A member of the family

Mcmurdodontidae; a new genus for "Mcmurdodus
" whitei Turner & Young (1987).

Moskovirhynchus[34]

Gen. et sp. nov

Valid

Popov & Shapovalov

Late Jurassic

 Russia

A

Callorhinchidae
. Genus includes new species M. robustus.

Nebriimimus[35]

Gen. et sp. nov

Valid

Collareta et al.

Pliocene (Zanclean)

 Italy

A member of Rajiformes, possibly a skate. The type species is N. wardi.

Negaprion cossardi[23]

Sp. nov

Valid

Canevet & Lebrun

Miocene

 France

A species of Negaprion.

Phoebodus curvatus[36]

Sp. nov

Valid

Ivanov

Devonian (GivetianFrasnian)

 Australia
 Poland
 Russia

Protochimaera[37]

Gen. et sp. nov

Valid

Lebedev & Popov in Lebedev et al.

Carboniferous (ViséanSerpukhovian)

Dashkovo Formation

 Russia
( Moscow Oblast)

A chimaera. Genus includes new species P. mirabilis.

Pseudocorax kindlimanni[38]

Sp. nov

In press

Jambura, Stumpf & Kriwet

Late Cretaceous (Cenomanian)

Sannine Formation

 Lebanon

Ptychodus maghrebianus[39]

Sp. nov

In press

Amadori et al.

Late Cretaceous (Turonian)

 Morocco

Reifella[40]

Gen. et sp. nov

Valid

Ivanov in Ivanov et al.

Permian (Roadian)

Cutoff Formation

 United States
( Texas)

A member of the family Anachronistidae. Genus includes new species R. lata.

Rosaodus[28]

Gen. et sp. nov

Valid

Li et al.

Triassic (Ladinian–Carnian)

 China

A member of Elasmobranchii of uncertain phylogenetic placement. Genus includes new species R. xingyiensis.

Strophodus indicus[41]

Sp. nov

Valid

Sharma & Singh

Middle Jurassic (Bathonian)

Jaisalmer Formation

 India

Marine hybodont shark

Strophodus jaisalmerensis[42]

Sp. nov

In press

Kumar et al.

Jurassic

Jaisalmer Formation

 India

A hybodont shark.

Taeniurops sergiomorai[43]

Sp. nov

Laurito & Valerio

Miocene–Pliocene (MessinianPiacenzian)

Uscari Formation

 Costa Rica

A stingray, a species of Taeniurops.

Triodus aeduorum[44]

Sp. nov

Valid

Luccisano et al.

Early Permian

Autun Basin

 France

Vallisodus[45]

Nom. nov

Valid

Duffin

Late Triassic

Penarth Group

 United Kingdom

A member of

Neoselachii
; a replacement name for Vallisia Duffin (1982).

Cartilaginous fish research

  • Description of new fossil material of Gualepis elegans from the Lower Devonian of Yunnan (China), and a study on the phylogenetic relationships of this fish, is published by Cui et al. (2021).[46]
  • Mottequin et al. (2021) reject the interpretation of Spiraxis interstrialis as chondrichthyan egg cases, and evaluate the implications of this reinterpretation for the knowledge of the evolution of oviparity in cartilaginous fishes.[47]
  • Description of the first known skull remains of
    Onchopristidae.[48]
  • New, exceptionally well-preserved skeleton of
    Altmühltal Formation (Germany) by Stumpf et al. (2021), who interpret this specimen as indicating that Asteracanthus and Strophodus represent two valid genera distinct from all other hybodontiforms.[49]
  • A study on the morphological diversity of teeth of lamniform sharks from mid-Cretaceous assemblages in Australia, and on its implications for the knowledge of the composition of mid-Cretaceous shark communities and their recovery in the aftermath of the Cenomanian-Turonian boundary event, is published by Bazzi, Kear & Siversson (2021).[50]
  • A study on the biomechanics of teeth of five species of Otodus, aiming to assess the functional significance of morphological trends in otodontid teeth and to test whether the morphology of otodontid teeth enabled the transition from piscivory to predation on marine mammals and the evolution of titanic body sizes, is published by Ballell & Ferrón (2021)[51]
  • A study on a bonebed in the
    Carcharocles angustidens dominated by small teeth is published by Miller, Gibson & Boessenecker (2021), who interpret this bonebed as a nursery area for C. angustidens.[52]
  • A study on growth patterns, reproductive biology and likely lifespan of Otodus megalodon is published by Shimada et al. (2021).[53]
  • Perez, Leder & Badaut (2021) present a novel method for estimating body size in fossil lamniform sharks, and attempt to determine the body size of Otodus megalodon.[54]
  • Revision of the fossil record of the extant tiger shark and the extinct members of the tiger shark lineage is published by Türtscher et al. (2021).[55]
  • Redescription of Striatolamia tchelkarnurensis is published by Malyshkina (2021).[56]
  • Shark teeth which might represent the first occurrence of the blacknose shark in the Pacific Ocean are described from the Pliocene Upper Onzole Formation (Ecuador) by Collareta et al. (2021), who evaluate the implications of this finding for the knowledge of the evolutionary history of the blacknose shark and the whitenose shark.[57]
  • Two fossil teeth of the blacktip shark are reported from lower Pliocene marine deposits of Tuscany (Italy) by Collareta et al. (2021), representing the first known occurrence of this species in the fossil record from both Europe and the Mediterranean Basin.[58]
  • A study on the morphological diversity of extant and fossil shark teeth, and on their implications for the knowledge of the evolution of lamniform and carcharhiniform sharks throughout the last 83 million years, is published by Bazzi et al. (2021).[59]
  • A study on the evolutionary history of sharks across the Cretaceous–Paleogene extinction event, as indicated by morphological diversity of shark teeth across the Cretaceous–Paleogene interval, is published by Bazzi et al. (2021).[60]
  • Evidence of a previously unknown major extinction of sharks in the early Miocene, ~19 million years ago, is presented by Sibert & Rubin (2021);[61] the study is subsequently criticized by Naylor et al. (2021)[62][63] and Feichtinger et al. (2021).[64][65]
  • A study on shark scales from mid-Holocene (~7,000-y-old) and modern reef sediments in Bocas del Toro (Panama), aiming to determine changes of shark abundance in this area since the mid-Holocene and their possible causes, is published by Dillon et al. (2021).[66]

Ray-finned fishes

Name Novelty Status Authors Age Type locality Country Notes Images

Achirus australis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Achirus.

Achirus chungkuz[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Achirus.

Agonopsis cume[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Agonopsis.

Agoultpycnodus[68]

Gen. et comb. nov

Valid

Taverne & Capasso

Late Cretaceous (Cenomanian-Turonian)

Akrabou Formation

 Morocco

A member of the family Pycnodontidae. The type species is A. aldrovandii.

Alilepis texasensis[40]

Sp. nov

Valid

Bakaev in Ivanov et al.

Permian (Roadian)

Cutoff Formation

 United States
( Texas)

A member of the family Elonichthyidae.

Anomoeodus aegypticus[69]

Sp. nov

Valid

Capasso et al.

Late Cretaceous (Maastrichtian)

Dakhla Formation

 Egypt

A member of Pycnodontiformes.

Anomoeodus caddoi[70]

Sp. nov

Valid

Suarez et al.

Early Cretaceous (Albian)

Holly Creek Formation

 United States
( Arkansas)

A member of Pycnodontiformes.

Aphanolebias bettinae[71]

Sp. nov

In press

Bradić-Milinović, Rundić & Schwarzhans

Miocene

 Serbia

A member of the family

Valenciidae
.

Apuliadercetis gonzalezae[72]

Sp. nov

In press

Díaz-Cruz, Alvarado-Ortega & Cantalice

Late Cretaceous (Campanian)

Angostura Formation

 Mexico

A member of Aulopiformes belonging to the family Dercetidae.

Archaemacruroides vanknippenbergi[73]

Sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Maastrichtian)

Maastricht Formation

 Belgium
 Netherlands

A member of Gadiformes of uncertain phylogenetic placement.

Ariosoma mesohellenica[74]

Sp. nov

Valid

Agiadi, Koskeridou & Thivaiou

Miocene (Aquitanian)

 Greece

A species of Ariosoma.

Austelliscus[75]

Gen. et sp. nov

Figueroa, Weinschütz & Friedman

Middle Devonian or older

Paraná Basin

 Brazil

An early ray-finned fish. Genus includes new species A. ferox.

Auxis koreanus[76]

Sp. nov

Valid

Nam, Nazarkin & Bannikov

Middle Miocene

Duho Formation

 South Korea

A species of Auxis.

Bardackichthys[77]

Gen. et sp. nov

In press

Hacker & Shimada

Late Cretaceous (Cenomanian)

Woodbine Formation

 United States
( Texas)

A member of Ichthyodectiformes. Genus includes new species B. carteri.

Bobbitichthys[78]

Gen. et comb. nov

Valid

Schwarzhans, Milàn & Carnevale

Paleocene (Selandian)

Kerteminde Marl

 Denmark

A member of the family Macrouridae. The type species is "Hymenocephalus" rosenkrantzi Schwarzhans (2003).

Brauccipycnodus[79]

Gen. et comb. nov

Valid

Taverne & Capasso

Early Cretaceous (Albian)

 Italy

A member of the family Pycnodontidae. The type species is "Proscinetes" pillae Capasso (2007).

Briveichthys[80]

Gen. et sp. nov

Valid

Štamberg & Steyer

Permian

Brive Basin

 France

A pygopterid. Genus includes new species B. chantepieorum.

Capromimus undulatus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Capromimus.

Cheirolepis jonesi[81]

Sp. nov

In press

Newman et al.

Devonian (Givetian)

Tordalen Formation

 Norway

Chiloconger chilensis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Chiloconger.

Choichix[82]

Gen. et sp. nov

Valid

Cantalice, Than-Marchese & Villalobos-Segura

Late Cretaceous (Cenomanian)

 Mexico

A member of Acanthopterygii of uncertain phylogenetic placement. Genus includes new species C. alvaradoi.

Citharichthys parvisulcus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún beds
Lacui Formation
Navidad Formation
Ranquil Formation

 Chile

A species of Citharichthys.

Citharichthys vergens[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Citharichthys.

Clarotes eocenicus[83]

Sp. nov

Valid

Murray & Holmes

Late Eocene

Jebel Qatrani Formation

 Egypt

A species of Clarotes.

Coccolepis solnhofensis[84]

Sp. nov

Valid

López-Arbarello & Ebert

Late Jurassic (Tithonian)

Altmühltal Formation

 Germany

A member of Chondrostei belonging to the family Coccolepididae.

Coelorinchus fidelis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún Beds
Lacui Formation
Navidad Formation

 Chile

A species of Coelorinchus.

Coelorinchus rapelanus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Coelorinchus.

Cottunculus primaevus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Cottunculus.

Cretaserranus[73]

Gen. et sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Maastrichtian)

Maastricht Formation

 Belgium
 Netherlands

A member of Perciformes, possibly belonging to the family Serranidae. Genus includes new species C. maastrichtensis.

Ctenopharyngodon orientalis[85]

Sp. nov

Valid

Su, Chang & Chen

Miocene

Xiacaowan Formation

 China

A species of

Ctenopharyngodon
.

Ctenopharyngodon xiejiaensis[85]

Sp. nov

Valid

Su, Chang & Chen

Miocene

Xiejia Formation

 China

A species of Ctenopharyngodon.

Dezaoia[85]

Gen. et sp. nov

Valid

Su, Chang & Chen

Oligocene

Ulanbulage Formation

 China

A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is D. saintjaquensis.

Diastemapycnodus[86]

Gen. et sp. nov

In press

Abu El-Kheir et al.

Late Cretaceous (Maastrichtian)

Dakhla Formation

 Egypt

A member of Pycnodontiformes. Genus includes new species D. tavernensis.

Elongofuro augustilgi[87]

Sp. nov

Valid

Ebert

Late Jurassic (Kimmeridgian)

Nusplingen Limestone

 Germany

A member of Ophiopsiformes.

Eoctenopharyngodon[85]

Gen. et sp. nov

Valid

Su, Chang & Chen

Oligocene

Dongying Formation

 China

A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is E. liui.

Eotexachara[88]

Gen. et sp. nov

In press

Wick

Late Cretaceous (Campanian)

Aguja Formation

 United States
( Texas)

A member of Characiformes. Genus includes new species E. malateres.

Feroxichthys panzhouensis[89]

Sp. nov

Valid

Ma, Xu & Geng

Middle Triassic (Anisian)

Guanling Formation

 China

A member of the family Colobodontidae.

Garganomyctophum[90]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Santonian)

 Italy

A lanternfish. The type species is G. sorbinii.

Gnathophis elongatus[74]

Sp. nov

Valid

Agiadi, Koskeridou & Thivaiou

Miocene (Aquitanian)

 Greece

A species of Gnathophis.

Gnathophis quinzioi[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Gnathophis.

Gobiosoma? axsmithi[91]

Sp. nov

Valid

Ebersole, Cicimurri & Stringer

Oligocene (Rupelian)

Byram Formation

 United States
( Alabama)

A member of the family Gobiidae.

Guiclupea[92]

Gen. et sp. nov

Valid

Chen et al.

Oligocene

 China

A member of

Clupeomorpha belonging to the group Ellimmichthyiformes
. The type species is G. superstes.

Hellenigobius[93]

Gen. et sp. et comb. nov

Valid

Schwarzhans, Agiadi & Thivaiou

Miocene

 Czech Republic
 Greece
 Italy
 Kazakhstan
 Serbia
 Hungary?

A member of the subfamily Gobionellinae. The type species is H. praeschismatus; genus also includes "Pomatoschistus" bunyatovi Bratishko et al. (2015).

Katyagobius[94]

Gen. et sp. nov

Valid

Reichenbacher & Bannikov

Miocene

 Moldova

A member of the family Gobiidae. The type species is K. prikryli.

Kelliaichthys[95]

Nom. nov

Valid

Schultze in Schultze et al.

 Kazakhstan

A replacement name for Kellia Kazantseva-Selezneva (1980).

Klincigobus haraldahnelti[71]

Sp. nov

In press

Bradić-Milinović, Rundić & Schwarzhans

Miocene

 Serbia

A member of the family Gobiidae.

Krumvirichthys[96]

Gen. et sp. nov

Valid

Přikryl

Oligocene–early Miocene

 Czech Republic

A deep-sea smelt. The type species is K. brzobohatyi.

Kuhlia orientalis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A flagtail.

Lampanyctus ipunensis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún beds

 Chile
 New Zealand

A species of Lampanyctus.

Larimichthys koae[97]

Sp. nov

In press

Lin & Chien

Late Miocene

Tapu Formation

 Taiwan

A species of Larimichthys.

Lepophidium chonorum[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Lepophidium.

Lepophidium mapucheorum[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Lepophidium.

Maurolicus brevirostris[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Maurolicus.

Navidadichthys[67]

Gen. et sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

Possibly a member of the family

Prototroctidae
. The type species is N. mirus.

Neuburgichthys[95]

Nom. nov

Valid

Schultze in Schultze et al.

 Kazakhstan

A replacement name for Neuburgia Kazantseva-Selezneva (1980).

Nezumia epuge[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation
Navidad Formation

 Chile

A species of Nezumia.

Nibea chaoi[97]

Sp. nov

In press

Lin & Chien

Late Miocene

Tapu Formation

 Taiwan

A species of Nibea.

Oshiaichthys[95]

Nom. nov

Valid

Schultze in Schultze et al.

Triassic

 Kyrgyzstan

A replacement name for Oshia Sytchevskaya (1999).

Palaeopantodon[98]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

 Lebanon

A member of the family Pantodontidae. The type species is P. vandersypeni.

Pankowskipiscis[99]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

 Lebanon

A member of the family Pantodontidae. The type species is P. haqelensis.

Paracarapus[67]

Gen. et sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A pearlfish. The type species is P. chilensis.

Peltoperleidus asiaticus[100]

Sp. nov

Valid

Xu

Middle Triassic (Anisian)

Guanling Formation

 China

A member of Neopterygii belonging to the group Louwoichthyiformes.

Petersichthys[101]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Cenomanian)

 Lebanon

A member of the family Pantodontidae. The type species is P. libanicus.

Physiculus pichi[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Physiculus.

Pinichthys shirvanensis[102]

Sp. nov

Valid

Bannikov

Miocene

 Russia
( Krasnodar Krai)

A member of Perciformes belonging to the group Stromateoidei.

Plesiogobius[93]

Gen. et sp. et comb. nov

Valid

Schwarzhans, Agiadi & Thivaiou

Miocene (Aquitanian)

 Greece

A member of the family Gobiidae belonging to the subfamily Gobiinae. The type species is P. felliensis.

Polyipnus bandeli[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Polyipnus.

Portentosoceros[103]

Gen. et comb. nov

Valid

Nazarkin & Bannikov

Miocene

 Russia
( Sakhalin Oblast)

A member of Percoidei of uncertain phylogenetic placement; a new genus for "Pentaceros" sakhaliniсus Gretchina (1975).

Primuluchara[88]

Gen. et sp. nov

In press

Wick

Late Cretaceous (Campanian)

Aguja Formation

 United States
( Texas)

A member of Characiformes. Genus includes new species P. laramidensis.

Pseudohilsa nikosi[104]

Sp. nov

Valid

Kevrekidis, Arratia & Reichenbacher in Kevrekidis et al.

Late Miocene

 Greece

A member of the family Clupeidae.

Pseudolesueurigobius[94]

Gen. et sp. nov

Valid

Reichenbacher & Bannikov

Miocene

 Moldova

A member of the family Gobiidae. The type species is P. manfredi.

Pseudonus humilis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Ipún beds
Navidad Formation

 Chile

A species of Pseudonus.

Pteronisculus changae[105]

Sp. nov

Valid

Ren & Xu

Middle Triassic (Anisian)

Guanling Formation

 China

Pterothrissus transpacificus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of

Pterothrissus
.

Pythonichthys panulus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation

 Chile

A species of Pythonichthys.

Raususetarches[106]

Gen. et sp. nov

Valid

Yabumoto & Nazarkin

Late Miocene

Koshikawa Formation

 Japan

A member of the family Scorpaenidae. Genus includes new species R. sakurai.

Rhinocephalus cretaceus[73]

Sp. nov

In press

Schwarzhans & Jagt

Late Cretaceous (Maastrichtian)

Maastricht Formation

 Belgium

A member of the family Merlucciidae.

Rhynchoconger chiloensis[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Rhynchoconger.

Sarmatigobius[94]

Gen. et sp. et comb. nov

Valid

Reichenbacher & Bannikov

Miocene

 Moldova

A member of the family Gobiidae. The type species is S. compactus; genus also includes "Hesperichthys" iugosus Schwarzhans, Brzobohatý & Radwańska (2020).

Saurichthys sceltrichensis[107]

Sp. nov

Valid

Renesto, Magnani & Stockar

Middle Triassic (Ladinian)

Meride Limestone

  Switzerland

Severnichthus[108][109]

Gen. et sp. nov

In press

Stringer & Schwarzhans

Late Cretaceous (Maastrichtian)

Severn Formation

 United States
( Maryland)

Possibly a member of

Polymixiiformes
. Genus includes new species S. bourdoni.

Sirembola supersa[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Lacui Formation

 Chile

A cusk-eel.

Spectrunculus sparsus[67]

Sp. nov

Schwarzhans & Nielsen

Early Miocene

Navidad Formation

 Chile

A species of Spectrunculus.

Taosciaena[97]

Gen. et 2 sp. nov

In press

Lin & Chien

Late Miocene

Tapu Formation

 Taiwan

A member of the family Sciaenidae. The type species is T. jiangi; genus also includes T. hui.

Toxopyge vracevicensis[71]

Sp. nov

In press

Bradić-Milinović, Rundić & Schwarzhans

Miocene

 Serbia

A member of the family Gobiidae.

Wilsonium[110]

Gen. et comb. nov

Valid

Liu

Early Eocene

Allenby Formation

 Canada
( British Columbia)

A Catostomidae sucker.
The type species is "Amyzon" brevipinne (1893).

Yarigobius[94]

Gen. et 2 sp. nov

Valid

Reichenbacher & Bannikov

Miocene

 Moldova

A member of the family Gobiidae. The type species is Y. decoratus; genus also includes Y. naslavcensis.

Ray-finned fish research

  • Putative paraxial ossicle of a member of the family Molidae, possibly representing the first fossil find of the genus Mola from the Mediterranean Basin reported to date, is described from the Miocene Pietra Leccese Formation (Apulia, Italy) by Collareta et al. (2021).[111]
  • A platysomid specimen, representing the earliest deep-bodied actinopterygian reported to date, is described from the Carboniferous (Tournaisian) Horton Bluff Formation (Canada) by Wilson, Mansky & Anderson (2021), who evaluate the implications of this findings for the knowledge of the evolution of early ray-finned fishes.[112]
  • A review of the fossil record of Early–Middle Triassic marine bony fishes, aiming to determine the implications of poor fossil record from the late Olenekian-early middle Anisian interval on the knowledge of the Triassic radiation of bony fishes, is published by Romano (2021).[113]
  • A diverse assemblage of late Maastrichtian and Paleocene ray-finned fishes is described from Evrytania (Greece) by Argyriou & Davesne (2021).[114]
  • A study on the diversity of fishes from upper Paleocene microfossil localities in the Ravenscrag Formation (Saskatchewan, Canada) is published by Sinha et al. (2021).[115]
  • New fish fauna dating to the Paleocene–Eocene Thermal Maximum, indicating that diverse fish communities thrived in the paleotropics during this time period, is reported from Egypt by El-Sayed et al. (2021).[116]
  • Heingård et al. (2021) report preservation of residues of both internal and integumentary tissues in the form of dark organic stains in fossil fish larvae from the Eocene (Ypresian) Stolleklint Clay (Ølst Formation, Denmark).[117]
  • Revision of the fossil material of sturgeons from the Upper Miocene deposits of southern Ukraine is published by Hilton, Kovalchuk & Podoplelova (2021).[118]
  • A study on the morphological diversity and evolution of pycnodontiforms is published by Cawley et al. (2021).[119]
  • A study on fossil crushing dentitions of Pycnodus zeaformis and P. maliensis, providing evidence of a distinct pattern of gap-filling tooth addition in pycnodonts, with individual large teeth replaced by multiple small teeth, is published by Collins & Underwood (2021).[120]
  • A study on the histology of teeth and bones of Neoproscinetes penalvai and bones of Tepexichthys aranguthyorum is published by Meunier et al. (2021).[121]
  • A redescription of
    stem-group teleost, and name the new family Atacamichthyidae.[122]
  • Revision of members of the clade Archaeomaenidae is published by Bean (2021), who considers Madariscus robustus to be a junior synonym of Archaeomaene tenuis.[123]
  • A study on genome size evolution in fossil stem-group teleosts (based on data from bone cell volumes in fossil specimens), aiming to determine the timing of whole-genome duplication in the evolutionary history of teleosts, is published by Davesne et al. (2021).[124]
  • New fossil material of elopomorphs, including the earliest records of members of the genera Albula and Paralbula from Gondwana reported to date and one of the earliest records of the genus Egertonia, is described from the Upper Cretaceous Mahajanga Basin (Madagascar) by Ostrowski (2021).[125]
  • A study on the evolutionary history of lanternfishes, primarily based on the fossil record of otoliths, is published by Schwarzhans & Carnevale (2021).[126]
  • Armbruster & Lujan (2021) identify Taubateia paraiba as a member of Rhinelepinae.[127]

Lobe-finned fishes

Name Novelty Status Authors Age Type locality Country Notes Images

Ceratodus guanganensis[128]

Sp. nov

Valid

Wang et al.

Late Jurassic

Shaximiao Formation

 China

Announced in 2021; the final version of the article naming it was published in 2022.

Eusthenodon bourdoni[129]

Sp. nov

Valid

Downs, Barbosa & Daeschler

Devonian (Famennian)

Catskill Formation

 United States
( Pennsylvania)

Lobe-finned fish research

  • A coelacanth specimen belonging or related to the species Heptanema paradoxum is described from the Ladinian Meride Limestone (Switzerland) by Renesto, Magnani & Stockar (2021), representing the first known coelacanth specimen from the Middle Triassic that undoubtedly bears elongate thin ribs.[130]
  • Fossil material of mawsoniid coelacanths is described from the marine Rhaetian Bonenburg locality (Germany) by Hartung et al. (2021), who interpret this finding as indicating that mawsoniids were already present in Europe in the Late Triassic, and that they inhabited marine environments at the end of the Triassic.[131]
  • Fossil material of a member of genus
    Woodbine Formation (Texas, United States; representing the first Cretaceous North American mawsoniid coelacanth reported to date) by Cavin et al. (2021), who evaluate the implications of this finding for the knowledge of potential factors that might have made long survival of the genera Mawsonia and Latimeria possible.[132]
  • An ossified lung of a mawsoniid coelacanth is described from the Maastrichtian of Oued Zem (Morocco) by Brito et al. (2021), representing the last known record of a Mesozoic coelacanth and the first known occurrence of coelacanths in the phosphate deposits of North Africa.[133]
  • A study on the evolution of feeding modes among tetrapodomorphs, as indicated by the anatomy of the skull of Tiktaalik roseae, is published by Lemberg, Daeschler & Shubin (2021), who report the simultaneous occurrence of anatomical modifications of the skull for prey capture through biting, as well as joint morphologies suggestive of cranial kinesis that is also present in suction-feeding fish.[134]
  • A study on the phylogenetic relationships of extant and fossil coelacanths is published by Toriño, Soto & Perea (2021).[135]
  • A study on the morphology and histology of the scales of Miguashaia bureaui, and on its implications for the knowledge of the evolution of the squamation in coelacanths, is published by Mondéjar-Fernández et al. (2021).[136]
  • New fossil remains representing one of the largest known coelacanths ever reported are described from the Middle Jurassic of Normandy (France) by Cavin et al. (2021), who also compare the relationship between taxic diversity and body size diversity in coelacanths and ray-finned fishes over the Devonian–Paleocene time interval.[137]
  • A study on tooth development in Powichthys thorsteinssoni, evaluating its implications for the knowledge of the evolution of the dentition of bony fishes, is published by King, Marone & Rücklin (2021).[138]
  • A study on the anatomy and phylogenetic relationships of Cladarosymblema narrienense is published by Clement et al. (2021).[139]

General research

  • A study on the morphology of the earliest osteocytes in Tremataspis mammillata and Bothriolepis trautscholdi is published by Haridy et al. (2021), who interpret their findings as indicating that the earliest known osteocytes in the fossil record had similar morphology and likely similar physiological capabilities to their modern counterparts, and attempt to determine initial driver favoring evolution of cellular (osteocytic) over acellular (anosteocytic) bones in vertebrates.[140]
  • Two Permian fish assemblages consisting of cartilaginous fishes and ray-finned fishes are reported from the Madumabisa Mudstone Formation (Zambia) by Peecook et al. (2021), who compare these assemblages with middle and late Permian freshwater fish faunas from Australia, Brazil, Chile and South Africa.[141]
  • A middle Miocene freshwater fish fossil fauna is described from the Castilletes Formation (Colombia) by Ballen et al. (2021), report the presence of members of fish groups known from extant Amazonian faunas east of the Andes but absent from faunas west of the Andes, and interpret their presence as evidence that the riverine systems of the Guajira Peninsula were connected to Amazonia during the middle Miocene.[142]

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