2020 in paleobotany

Source: Wikipedia, the free encyclopedia.
List of years in paleobotany
In paleontology
2017
2018
2019
2020
2021
2022
2023
In arthropod paleontology
2017
2018
2019
2020
2021
2022
2023
In paleoentomology
2017
2018
2019
2020
2021
2022
2023
In paleomalacology
2017
2018
2019
2020
2021
2022
2023
In reptile paleontology
2017
2018
2019
2020
2021
2022
2023
In archosaur paleontology
2017
2018
2019
2020
2021
2022
2023
In mammal paleontology
2017
2018
2019
2020
2021
2022
2023
In paleoichthyology
2017
2018
2019
2020
2021
2022
2023

This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

Flowering plants

Alismatales

Name Novelty Status Authors Age Type locality Location Notes Images

Limnobiophyllum pedunculatum[1]

Sp. nov

Valid

Low, Su & Xing in Low et al.

Late Oligocene

 China

A member of the family Araceae.

Apiales

Name Novelty Status Authors Age Type locality Location Notes Images

Paleopanax puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

 United States
( Tennessee)

A member of the family Araliaceae.

Arecales

Name Novelty Status Authors Age Type locality Location Notes Images

Echimonocolpites chicxulubensis[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Arecaceae.

Palmoxylon ceroxyloides[4]

Sp. nov

In press

Khan, Hazra & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

 India

A petrified palm stem of a member of the subfamily Ceroxyloideae.

Palmoxylon dindoriensis[5]

Sp. nov

Valid

Khan, Roy & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

 India

A petrified palm stem.

Sabalites dawsonii[6]

Sp. nov

Valid

Greenwood & Conran

Eocene

 Canada
( British Columbia)

Sabalites karondiensis[7]

Sp. nov

In press

Roy, Hazra & Khan in Roy et al.

Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian)

Deccan Intertrappean Beds

 India

A palm frond .

Spinizonocolpites riochiquensis[8]

Sp. nov

Valid

Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal

Late Cretaceous (Maastrichtian)

Lago Colhué Huapí Formation

 Argentina

A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Buxales

Name Novelty Status Authors Age Type locality Location Notes Images

Pachysandra europaea[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Pachysandra. Announced in 2019; the final version of the article naming it was published in 2020.

Caryophyllales

Name Novelty Status Authors Age Type locality Location Notes Images

Gomphrenipollis garciae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant, possibly produced by members of the family Amaranthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Chloranthales

Name Novelty Status Authors Age Type locality Location Notes Images

Sarcandraxylon[11]

Gen. et sp. nov

Valid

Pipo, Iglesias & Bodnar

Late Cretaceous (early–middle Campanian)

Santa Marta Formation

Antarctica

A member of the family Chloranthaceae. Genus includes new species S. sanjosense.

Cornales

Name Novelty Status Authors Age Type locality Location Notes Images

Amersinia littletonensis[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

Langtonia parva[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Mastixicarpum hoodii[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Nyssa gergoei[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A tupelo.

Nyssa gyoergyi[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A tupelo.

Platycrater iljinskajae[14]

Sp. nov

In press

Denk et al.

Late Oligocene

 Russia

Portnallia alexanderi[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Crossosomatales

Name Novelty Status Authors Age Type locality Location Notes Images
Staphylea woodworthensis[15] Sp. nov In press Zhu & Manchester Oligocene Renova Formation  United States
( Montana)
A species of Staphylea.

Cucurbitales

Name Novelty Status Authors Age Type locality Location Notes Images

Coriaripites goodii[16]

Sp. nov

Valid

Barreda, Palazzesi & Tellería in Renner et al.

Late Cretaceous (Campanian–early Maastrichtian)

Lopez de Bertodano Formation
Santa Marta Formation
Snow Hill Island Formation

Antarctica

Pollen grains similar to those of extant members of the genus Coriaria.

Echitriporites jolyi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021.

Ericales

Name Novelty Status Authors Age Type locality Location Notes Images

Andrewsiocarpon puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

 United States
( Tennessee)

A member of the family Theaceae.

Anubiscarpon[17]

Gen. et sp. nov

Valid

Smith & Manchester

Middle Eocene

Clarno Formation

 United States
( Oregon)

A member of the family Theaceae. Genus includes new species A. andersonae.

Fabales

Name Novelty Status Authors Age Type locality Location Notes Images

Cercis zekuensis[18]

Sp. nov

Valid

Li et al.

Early Miocene

 China

A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021.

Gleditsia pliocaenica[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020.

Menendoxylon lutzi[19] Sp. nov In press Baez & Crisafulli Miocene Chiquimil  Argentina Fossil wood of a member of the family Fabaceae.

Parkiidites marileae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021.

Prioria martineziorum[20]

Sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

Santiago Formation

 Panama

A species of Prioria.

Psilastephanocolporites deoliverae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Psilastephanocolporites endoporatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Striatopollis grahamii[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of an eudicot, probably a member of the family Fabaceae.

Fagales

Name Novelty Status Authors Age Type locality Location Notes Images

Alnus chaybulakensis[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

An alder.

Berryophyllum hainanensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

Carpinus asymmetrica[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

Maguan

 China

A species of

Carpinus
.

Carpinus symmetrica[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

Maguan Basin

 China

A species of

Carpinus

Carya pipecreekensis[24]

Sp. nov

In press

Swinehart & Farlow

Late Neogene

Pipe Creek Sinkhole

 United States
( Indiana)

A hickory.

Castaneophyllum hainanensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

Castaneophyllum lanceolata[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

Castanopsis bulgarica[25] Sp. nov Valid Mantzouka, Ivanov & Bozukov Late Miocene–early Pliocene (late Messinian–early Zanclean) Pokrovnik  Bulgaria A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021.

Lithocarpus changchangensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A species of Lithocarpus.

Quercus borissovii[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

An oak.

Quercus changchangensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Quercus paleoargyrotricha[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Quercus paleohypargyrea[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Quercus paleolamellosa[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Garryales

Name Novelty Status Authors Age Type locality Location Notes Images

Eucommia szaferi[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Eucommia. Announced in 2019; the final version of the article naming it was published in 2020.

Gentianales

Name Novelty Status Authors Age Type locality Location Notes Images

Asclepiadospermum[26]

Gen. et 2 sp. nov

Valid

Del Rio et al.

Early Eocene

Niubao Formation

 China

An asclepiadoid Apocynaceae genus.
Included species A. marginatum and A. ellipticum.

Margocolporites carinae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021.

Icacinales

Name Novelty Status Authors Age Type locality Location Notes Images

Iodes elliptica[27]

Sp. nov

In press

Del Rio et al.

Early Oligocene

Wenshan Basin

 China

A member of the family Icacinaceae.

Iodes passiciensis[28]

Sp. nov

Valid

Del Rio & De Franceschi

Early Eocene

 France

A member of the family Icacinaceae.

Manchesteria[29]

Gen. et sp. nov

In press

Stull & Rozefelds in Rozefelds et al.

Cenozoic (mid-Miocene or, more likely, middle Eocene)

 Australia

A member of the family Icacinaceae. Genus includes new species M. australis.

Pyrenacantha simonsii[30]

Sp. nov

Valid

Stull et al.

Early Oligocene

 Egypt

A species of Pyrenacantha.

Lamiales

Name Novelty Status Authors Age Type locality Location Notes Images

Echitricolpites cruziae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021.

Multiareolites? reticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites curvimuratus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites pardoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021.

Laurales

Name Novelty Status Authors Age Type locality Location Notes Images

Actinodaphnoxylon[31]

Gen. et sp. nov

In press

Akkemik et al.

Eocene (Lutetian)

 Turkey

A member of the family Lauraceae. Genus includes new species A. zileensis.

Mezilaurinoxylon oleiferum[32]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Patagonoxylon[32]

Gen. et sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Thymolepis[33]

Gen. et sp. nov

Valid

Chambers & Poinar

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.

Valviloculus[34]

Gen. et sp. nov

Valid

Poinar et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis.

Liliales

Name Novelty Status Authors Age Type locality Location Notes Images

Smilax fujianensis[35]

Sp. nov

Valid

Dong et al.

Middle Miocene

 China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Smilax zhangpuensis[35]

Sp. nov

Valid

Dong et al.

Middle Miocene

 China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Magnoliales

Name Novelty Status Authors Age Type locality Location Notes Images

Magnolia nanningensis[36]

Sp. nov

Valid

Huang et al.

Late Oligocene

Nanning Basin

 China

A species of Magnolia.

Magnolia waltheri[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020.

Melloniflora[37]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Potomac Group

 United States
( Virginia)

A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis.

Malpighiales

Name Novelty Status Authors Age Type locality Location Notes Images

Salix palaeofutura[38]

Sp. nov

Valid

Narita et al.

Miocene

Bifuka Formation

 Japan

A willow.

Malvales

Name Novelty Status Authors Age Type locality Location Notes Images
Bastardioxylon[19] Gen. et sp. nov In press Baez & Crisafulli Miocene Chiquimil  Argentina Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua.
Dipterocarpus dindoriensis[39] Sp. nov Valid Khan, Spicer & Bera in Khan et al. Late Cretaceous (Maastrichtian) Deccan Intertrappean Beds  India A species of Dipterocarpus.

Echiperiporites germeraadii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites jaramilloi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites titanicus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolporites elizabeteae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021.

Veraguasoxylon[20]

Gen. et sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

Santiago Formation

 Panama

A member of the family Malvaceae. Genus includes new species V. panamense.

Myrtales

Name Novelty Status Authors Age Type locality Location Notes Images

Eucalyptus xoshemium[40]

Sp. nov

Valid

Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon

Eocene (Ypresian)

Huitrera Formation

 Argentina

A species of Eucalyptus.

Mangroveoxylon[41]

Gen. et comb. nov

In press

Moya & Brea

Late Miocene?

Ituzaingó Formation

 Argentina

A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979).

Myrceugenellites grandiporosum[32]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Primotrapa[42]

Gen. et sp. nov

Valid

Li & Li in Li et al.

Late Eocene to early Miocene

Hannuoba

 China
 Czech Republic
 France
 Germany

A member of

Trapoideae
. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018).

Nymphaeales

Name Novelty Status Authors Age Type locality Location Notes Images

Praenymphaeapollenites[43]

Gen. et sp. nov

Valid

Barrón, Peris & Labandeira in Peris et al.

Cenomanian

Burmese amber

 Myanmar

Pollen of a member of Nymphaeaceae.
Genus includes new species P. cenomaniensis.

Oxalidales

Name Novelty Status Authors Age Type locality Location Notes Images

Cunoniantha[44]

Gen. et sp. nov

Valid

Jud & Gandolfo

Paleocene (early Danian)

Salamanca Formation

 Argentina

A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021.

Elaeocarpus nanningensis[45]

Sp. nov

In press

Liu et al.

Late Oligocene

Yongning Formation

 China

A species of Elaeocarpus.

Elaeocarpus prelacunosus[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus preprunifolioides[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus prerugosus[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus preserratus[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

Elaeocarpus presikkimensis[45]

Sp. nov

In press

Liu et al.

Miocene

Erzitang Formation

 China

A species of Elaeocarpus.

Poales

Name Novelty Status Authors Age Type locality Location Notes Images

Monoporopollenites scabratus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Rhizomatites[46]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

 Argentina

A member of Cyperaceae. Genus includes new species R. cyperoides.

Proteales

Name Novelty Status Authors Age Type locality Location Notes Images

Banksia microphylla[47]

Sp. nov

Valid

Carpenter in Carpenter & Milne

Late Eocene

 Australia

A species of Banksia.

Banksieaeidites zanthus[47]

Sp. nov

Valid

Milne in Carpenter & Milne

Late Eocene

 Australia

A Banksia-like pollen.

Platanus emryi[48]

Sp. nov

Valid

Huegele, Spielbauer & Manchester

Miocene

 United States

A species of Platanus.

Ranunculales

Name Novelty Status Authors Age Type locality Location Notes Images

Cissampelos defranceschii[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021.

Clematis csabae[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A species of Clematis.

Diploclisia praeaffinis[50]

Sp. nov

Valid

Jia et al.

Late Miocene

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites bangorensis[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites haominae[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites tibetica[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Paleoorbicarpum[51]

Gen. et sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

 China

A member of the family Menispermaceae. Genus includes new species P. parvum.

Stephania bangorensis[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Stephania geniculata[51]

Sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

 China

A species of Stephania

Stephania ornamenta[51]

Sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

 China

A species of Stephania

Stephania shuangxingii[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Rosales

Name Novelty Status Authors Age Type locality Location Notes Images

Berhamniphyllum junrongii[52]

Sp. nov

Valid

Zhou, Wang & Huang in Zhou et al.

Late Eocene

Markam Basin

 China

A member of the family Rhamnaceae

Crataegus pentagynoides[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020.

Hemiptelea kryshtofovichii[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

A member of the family Ulmaceae.

Scabrastephanoporites[3]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

Sapindales

Name Novelty Status Authors Age Type locality Location Notes Images

Acer dombeyopsis[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Acer viburnoides[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Acer vitiforme[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Brosipollis reticulatus[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Burseraceae.

Choerospondias fujianensis[53]

Sp. nov

In press

Wang et al.

Miocene

 China

A species of Choerospondias.

Llanodelacruzoxylon[54] Gen. et sp. nov Rodríguez-Reyes, Estrada-Ruiz & Gasson Oligocene–Miocene Santiago Formation  Panama A member of the family Anacardiaceae. Genus includes new species L. sandovalii.

Manchestercarpa[55]

Gen. et sp. nov

Valid

Atkinson

Late Cretaceous (Campanian)

 Canada
( British Columbia)

A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.

Parametopioxylon[56]

Gen. et sp. nov

Valid

Franco et al.

Miocene

Ituzaingó Formation

 Argentina

A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.

Psilastephanocolporites hammenii[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Meliaceae

Quinquala[57]

Gen. et sp. nov

Valid

Manchester & Disney in Manchester, Disney & Pham

Eocene

Clarno Formation
Tepee Trail Formation

 United States
( Oregon
 Wyoming)

A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata.

Rousea cavitata[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Saxifragales

Name Novelty Status Authors Age Type locality Location Notes Images

Corylopsis grisea[58]

Sp. nov

Valid

Quirk & Hermsen

Early Pliocene

Gray Fossil Site

 United States
( Tennessee)

A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021.

Itea polyneura[59]

Sp. nov

In press

Huang & Tian in Tian et al.

Oligocene

Huazhige Formation

 China

A species of Itea.

Protoaltingia[60]

Gen. et sp. nov

Valid

Scharfstein, Stockey & Rothwell

Late Cretaceous (Coniacian)

 Canada
( British Columbia)

A member of the family Altingiaceae. Genus includes new species P. comoxense.

Solanales

Name Novelty Status Authors Age Type locality Location Notes Images

Physalis hunickenii[61]

Sp. nov

Valid

Deanna, Wilf & Gandolfo

Early Eocene

Laguna del Hunco Formation

 Argentina

A species of Physalis.

Trochodendrales

Name Novelty Status Authors Age Type locality Location Notes Images

Eotrochion[62]

Gen. et sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

 United States
( Wyoming)

A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021.

Paraconcavistylon[62]

Gen. et comb. nov

Valid

Manchester, Kvaček & Judd

Ypresian

Klondike Mountain Formation

 United States
( Washington)

A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021.

Trochodendron infernense[62]

Sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

 United States
( Wyoming)

A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021.

Vitales

Name Novelty Status Authors Age Type locality Location Notes Images

Yua jiangxiensis[63]

Sp. nov

Valid

He & Wang

Miocene

Toupi Formation

 China

A species of Yua. Announced in 2020; the final version of the article naming it was published in 2021.

Other angiosperms

Name Novelty Status Authors Age Type locality Location Notes Images

Aextoxicoxylon kawasianus[64]

Sp. nov

Valid

Vera et al.

Late Cretaceous

Puntudo Chico Formation

 Argentina

A fossil dicot wood

Atlantocarpus[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 Portugal
 United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis.

Carpolithes gergoei[13]

Sp. nov

Valid

Hably & Erdei in Hably

Miocene

 Hungary

A fossil fruit of a flowering plant of uncertain phylogenetic placement.

Catanthus[66]

Gen. et sp. nov

In press

Friis, Crane & Pedersen

Early Cretaceous

 Portugal

An early flowering plant. Genus includes new species C. dolichostemon.

Cavilignum[67]

Gen. et sp. nov

Valid

Siegert & Hermsen

Early Pliocene

Gray Fossil Site

 United States
( Tennessee)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil

endocarps
. Genus includes new species C. pratchettii.

Chainandra[68]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species C. zeugostylus.

Cichoreacidites? flammulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Crotonoidaepollenites echinatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Cyathitepala[69]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species C. papillosa.

Dasykothon[70]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.

Dinganthus[71]

Gen. et sp. nov

Valid

Liu et al.

Miocene

Dominican amber

 Dominican Republic

A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera.

Echistephanoporites annulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Eofructus[72]

Gen. et sp. nov

Valid

Han & Wang

Early Cretaceous

Yixian Formation

 China

An infructescence including a central axis and five fruits resembling Liaoningfructus. Genus includes new species E. liutiaogouensis.

Foveotricolporites crassus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Inaperturopollenites microechinatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Ladakhipollenites carmoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Lambertiflora[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans.

Malvacipolloides diversus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides echibaculatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides romeroae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Menatanthus[73]

Gen. et sp. nov

Valid

Uhl, Paudayal & El Atfy in Uhl et al.

Paleocene (Thanetian)

 France

A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021.

Mugideiriflora[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Aptian-early Albian)

Almargem Formation

 Portugal

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica.

Phantophlebia[74]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.

Platanites willigeri[75]

Sp. nov

Valid

Halamski & Kvaček in Halamski et al.

Late Cretaceous (Santonian)

Czerna Formation

 Poland

Trifoliolate platanoid leaves.

Psilaperiporites delicatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Psilaperiporites lunaris[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Ranunculacidites reticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Rasenganus[76]

Gen. et sp. nov

Valid

Xing & Gu

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A possible epizoochorous fruit. Genus includes new species R. auricularus.

Retibrevitricolpites microreticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites costaporus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites? toigoae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retipollenites solimoensis[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retitriporites crassoreticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Rhoipites alfredii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Singpuria[77]

Gen. et sp. nov

Valid

Ramteke, Manchester & Nagrale in Ramteke et al.

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

 India

A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei.

Sinoherba[78]

Gen. et sp. nov

Valid

Liu & Wang in Liu, Chen & Wang

Early Cretaceous (BarremianAptian)

Yixian Formation

 China

An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021.

Varifructus[79]

Gen. et sp. nov

In press

Liu et al.

Early Cretaceous

Yixian Formation

 China

An early flowering plant. Genus includes new species V. lingyuanensis.

Wireroadia[80]

Gen. et sp. et comb. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian to Santonian)

 United States
( Alabama
 Massachusetts
 New Jersey
 New York)

A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick).

Pinales

Name Novelty Status Authors Age Type locality Location Notes Images

Agathis ledongensis[81]

Sp. nov

Valid

Oskolski et al.

Late Oligocene–early Miocene

Qiutangling Formation

 China

A species of Agathis

Agathoxylon cozzoi[82]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

Chihuido Formation

 Argentina

Araucaria cuneoi[83]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

 Argentina

A species of Araucaria.

Araucaria famii[84]

Sp. nov

Valid

Stockey & Rothwell

Late Cretaceous (Campanian)

 Canada
( British Columbia)

A species of Araucaria.

Araucaria fildesensis[85]

Sp. nov

Valid

Shi et al.

Eocene

Fossil Hill Formation

Antarctica
(King George Island)

A species of Araucaria

Araucaria huncoensis[86]

Sp. nov

Valid

Rossetto‐Harris in Rossetto‐Harris et al.

Early Eocene

Laguna del Hunco Formation

 Argentina

A species of Araucaria.

Araucaria stockeyana[83]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

 Argentina

A species of Araucaria.

Araucarites pachacuteci[87] Sp. nov

In press

Martínez in Martínez et al.

Early Cretaceous (BerriasianValanginian)

Huancané Formation

 Peru

A member of the family Araucariaceae

Brachyoxylon zhouii[88]

Sp. nov

Valid

Jiang et al.

Early Cretaceous

Guantou Formation

 China

A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021.

Brachyphyllum sattlerae[89] Sp. nov Valid Batista et al. Early Cretaceous (Aptian) Crato Formation  Brazil A member of the family Pinidae.

Callialastrobus[90]

Gen. et sp. nov

In press

Kvaček & Mendes

Early Cretaceous (late Aptian–early Albian)

Lusitanian Basin

 Portugal

A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai.

Cedrus anatolica[91]

Sp. nov

Valid

Akkemik

Early Miocene

Hançili Formation

 Turkey

A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021.

Circoporoxylon tibetense[92]

Sp. nov

In press

Xia et al.

Middle Jurassic (Callovian)

Xiali Formation

 China

Possibly a member of the family Podocarpaceae.

Comoxostrobus[93]

Gen. et sp. nov

Valid

Stockey, Rothwell & Atkinson

Late Cretaceous (early Coniacian)

 Canada
( British Columbia)

A member of the family

Taiwanioideae
. Genus includes new species C. rossii.

Cupressinoxylon klimovii[94]

Nom. nov

Valid

Blokhina

Miocene

 Russia

A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989).

Cupressinoxylon llantenesense[82]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

Llantenes Formation

 Argentina

Cupressinoxylon manuelii[95] Sp. nov In press Ríos-Santos, Cevallos-Ferriz & Pujana Late Cretaceous (Campanian-Maastrichtian) Cabullona Group  Mexico

Ductoagathoxylon wangii[96]

Sp. nov

In press

Gou & Feng in Gou et al.

Middle Jurassic

Xishanyao Formation

 China

A conifer stem.

Friisia[97]

Gen. et sp. nov

In press

Mendes & Kvaček

Early Cretaceous (late Aptian – early Albian)

Lusitanian Basin

 Portugal

An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.

Juniperoxylon acarcaea[98]

Sp. nov

In press

Akkemik

Early Miocene

Hançili Formation

 Turkey

A member of the family Cupressaceae.

Lesbosoxylon kemaliyensis[99]

Sp. nov

Valid

Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım

Miocene

Divriği Formation

 Turkey

A member of the family Pinaceae.

Marskea cuspidata[100]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Prisayan Formation

 Russia

A member of the family Taxaceae.

Mukawastrobus[101]

Gen. et sp. nov

Valid

Stockey, Nishida & Rothwell

Late Cretaceous (late Campanian—early Maastrichtian)

 Japan

A member of the family

Taiwanioideae
. Genus includes new species M. satoi.

Piceoxylon yumeniense[102]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

 China

Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Pinuxylon selmeierianum[103]

Sp. nov

Valid

Dolezych & Reinhardt

Paleogene

Eureka Sound Group

 Canada
( Nunavut)

A member of the family Pinaceae described on the basis of fossil wood

Protophyllocladoxylon chijinense[102]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

 China

Announced in 2020; the final version of the article naming it was published in 2021.

Tsuga asiatica[104]

Sp. nov

Valid

Wu & Zhou in Wu et al.

Late Paleogene

 China

A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020.

Other seed plants

Name Novelty Status Authors Age Type locality Location Notes Images

Amyelon turpanense[105]

Sp. nov

In press

Shi, Yu & Sun

Permian (Lopingian)

Wutonggou

 China

A root of a member of Cordaitales

Androstrobus obovatus[106]

Sp. nov

In press

Bodnar et al.

Late Triassic

Potrerillos Formation

 Argentina

A member of

Cycadales
.

Araripestrobus[107]

Gen. et sp. nov

Valid

Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al.

Early Cretaceous (Aptian-Albian)

Crato

 Brazil

A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus.

Archaeopetalanthus[108]

Gen. et sp. nov

Valid

Naugolnykh

Carboniferous

Listvjanskaya

 Russia

A member of

Pinophyta belonging to the group Vojnovskyales
. Genus includes new species A. progressus.

Battenispermum[109]

Gen. et sp. nov

In press

Mendes, Pedersen & Friis

Early Cretaceous

 Portugal

A seed plant belonging to the informal grouping

Gnetales
. Genus includes new species B. hirsutum.

Carpolithus volantus[110]

Sp. nov

Valid

Gómez et al.

Early Cretaceous (Aptian)

La Cantera Formation

 Argentina

A fossil seed, possibly produced by a member of

Gnetales
.

Ductolobatopitys[111]

Gen. et sp. nov

Valid

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.

Filigigantopteris[112]

Gen. et 2 sp. nov

In press

Zhou et al.

Late Permian

Nayixiong Formation

 China

A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia.

Ginkgo pediculata[113]

Sp. nov

In press

Deng, Yang & Zhou

Early Cretaceous

 China

A species of Ginkgo.

Jianchangia[114]

Gen. et sp. nov

Valid

Yang, Wang & Ferguson

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Ephedraceae. Genus includes new species J. verticillata.

Johniphyllum[115]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales. Genus includes new species J. multinerve.

Jordaniopteris[116]

Gen. et comb. nov

Valid

Anderson in Anderson et al.

Permian (possibly Lopingian)

Um Irna Formation

 Jordan

A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).

Kaokoxylon brasiliensis[111]

Sp. nov

Valid

Conceição, Neregato & Iannuzzi in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A conifer described on the basis of fossil wood.

Nilssonia mirovanae[117]

Sp. nov

Valid

Čepičková & Kvaček

Late Cretaceous (Cenomanian)

Peruc-Korycany Formation

 Czech Republic

A cycad.

Novaiorquepitys[118]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.

Ovalocarpus butmanii[119]

Sp. nov

Valid

Naugolnykh & Linkevich

Permian (Artinskian)

 Russia
( Sverdlovsk Oblast)

A member of Ginkgoales belonging to the family Cheirocladaceae.

Palaeocupressinoxylon[120]

Gen. et sp. nov

In press

Wan, Yang & Wang

Late Permian

Turpan–Hami Basin

 China

A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.

Phoenicopsis anadyrensis[121]

Sp. nov

Valid

Nosova in Zolina et al.

Late Cretaceous–Paleocene (MaastrichtianDanian)

Rarytkin

 Russia
( Chukotka Autonomous Okrug)

A member of Czekanowskiales

Pseudovoltzia sapflorensis[115]

Sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales

Pterostoma neehoffii[122]

Sp. nov

Valid

Conran et al.

Middle Miocene

 New Zealand

A cycad

Pteruchus frenguellii[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A pollen organ of a seed fern

Pteruchus lepidus[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A pollen organ of a seed fern

Samaropsis jinchangensis[124]

Sp. nov

Valid

Hua & Sun in Hua et al.

Early Permian

 China

A seed fossil. Announced in 2019; the final version of the article naming was published in 2020.

Umaltolepis involuta[125]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

Umaltolepis sogdianica[125]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

Umkomasia aequatorialis[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A cupulate structure of a seed fern

Wantus[115]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales. Genus includes new species W. acaulis.

Wudaeophyton[126]

Gen. et sp. nov

Valid

Pšenička et al.

Early Permian

Taiyuan Formation

 China

A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.

Yangopteris[127]

Gen. et comb. nov

In press

Zhou et al.

Permian (Asselian)

 China

A seed fern; a new genus for "Alethopteris" ascendens Halle.

Yvyrapitys[118]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

Other plants

Name Novelty Status Authors Age Type locality Location Notes Images

Annularia paisii[128]

Sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

 Portugal

Blasiites huolinhensis[129]

Sp. nov

In press

Li et al.

Early Cretaceous

Huolinhe Formation

 China

A liverwort belonging to the family Blasiaceae.

Botryopteris multifolia[130]

Sp. nov

In press

He et al.

Permian (Lopingian)

Junlian Formation

 China

A fern

Birisia mandshurica[131]

Sp. nov

Valid

Golovneva, Grabovskiy & Zolina

Early Cretaceous (Albian)

Frentsevka Formation

 Russia
( Primorsky Krai)

A fern belonging to the family Dicksoniaceae.

Calamites cambrensis[132]

Sp. nov

Valid

Thomas

Carboniferous (Pennsylvanian)

 United Kingdom

Catenuporella[133]

Gen. et sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

 China

A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia.

Charaxis spicatus[134]

emend. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

 Spain

Thallus of Echinochara lazarii. Both taxa has been firstly found anatomically attached

Chlamydomonas hanublikanus[135]

Sp. nov

Vršanská & Hinkelman

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Chlamydomonas

Circinites[46]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

 Argentina

A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.

Clavatisporites cenomaniana[136]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of

Filicopsida
of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Clavator calcitrapus var. jiangluoensis[134]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

 Spain

Clavatoracean species.

Collarecodium? nezpercae[137] Sp. nov Valid Bucur & Rigaud in Bucur et al. Late Triassic (Norian)  United States
( Idaho)
A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.

Collarisporites minor[136]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of

Filicopsida
of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Coniopteris sandaolingensis[138]

Sp. nov

Valid

Yuan & Sun in Yuan et al.

Middle Jurassic

Xishanyao Formation

 China

Dimicheleodendron[139]

Gen. et comb. nov

Valid

Thomas & Cleal

Carboniferous

 United Kingdom

A lycophyte; a new genus for "Lepidodendron" hickii.

Drynaria diplosticha[140]

Sp. nov

Valid

Yu & Xie in Yu et al.

Late Miocene

Bangmai Formation

 China

A fern belonging to the family Polypodiaceae.

Echinochara lazarii[134]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et MArtín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

 Spain

A member of Clavatoraceans.

Equicalastrobus pusillus[141]

Sp. nov

Valid

Zhang & Yan in Zhang et al.

Late Triassic

Baojishan Basin

 China

A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021.

Equisetum yenbaiense[142]

Sp. nov

Valid

Aung et al.

Late Miocene

 Vietnam

A species of Equisetum

Equisetum yongpingense[142]

Sp. nov

Valid

Aung et al.

Late Pliocene

Sanying Formation

 Vietnam

A species of Equisetum

Filippoporella[143] Gen. et sp. nov In press Sokač & Grgasović Early Paleocene  Croatia A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi.

Frullania partita[144]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A liverwort, a species of Frullania

Frullania vanae[145]

Sp. nov

Valid

Mamontov et al.

Eocene

Rovno amber

 Ukraine

A liverwort, a species of Frullania

Gippslandites[146]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

 Australia

A member of Zosterophyllaceae. Genus includes new species G. minutus.

Gmujij[147] Gen. et sp. nov Valid Pfeiler & Tomescu Devonian (Emsian) Battery Point  Canada
( Quebec)
An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021.
Griphoporella minuta[137] Sp. nov Valid Bucur & Peybernes in Bucur et al. Late Triassic  Japan A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Hansopteris[148] Gen. et sp. nov In press Zhou et al. Permian (Asselian)  China An anachoropterid fern. Genus includes new species H. uncinatus.
Holosporella magna[137] Sp. nov Valid Bucur & Fucelli in Bucur et al. Late Triassic (Norian)  United States
( Nevada)
A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella? rossanae[137] Sp. nov Valid Bucur & Del Piero in Bucur et al. Late Triassic (Norian)  Canada
( Yukon)
A green alga belonging to the group Dasycladales and the family Triploporellaceae.

Inocladus[149]

Gen. et comb. nov

Valid

LoDuca et al.

Silurian

 United States

An alga related to the group Bryopsidales. Genus includes "Buthotrephis" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021.

Intermurella ordosensis[133]

Sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

 China

A green alga belonging to the group Dasycladales.

Jurafructus[150]

Gen. et sp. nov

Valid

Chen et al.

MiddleLate Jurassic

Jiulongshan Formation

 China

A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.

Keraphyton[151]

Gen. et sp. nov

Valid

Champreux, Meyer-Berthaud & Decombeix

Devonian (Famennian)

Mandowa Mudstone Formation

 Australia

A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.

Khasurtya[152]

Gen. et sp. nov

Valid

Mamontov in Kopylov et al.

Early Cretaceous

 Russia

A liverwort belonging to the group

Marchantiidae
. Genus includes new species K. ginkgoides.

Lepidodendron demkinae[153]

Sp. nov

Valid

Mosseichik

Carboniferous (Viséan)

 Russia

Lobatannularia linjiaensis[154]

Sp. nov

Valid

Xu et al.

Middle Triassic

Linjia Formation

 China

A member of Equisetales.

Lygodium sanshuiense[155]

Sp. nov

Valid

Naugolnykh et al.

Paleocene

Buxin Formation

 China

A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020.

Metzgeriites kujiensis[156]

Sp. nov

Valid

Katagiri in Katagiri & Shinden

Late Cretaceous (Santonian)

Tamagawa Formation

 Japan

A liverwort.

Munieria martinclosasi[134]

comb. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Lebanon

 Lebanon

Clavatoracean thallus. Formerly known as Charaxis martinclosasi

Neoarthropitys[157] Gen. et sp. nov In press Gnaedinger et al. Middle Triassic Quebrada de los Fósiles  Argentina A member of Equisetales. Genus includes new species N. gondwanaensis.

Osmundacaulis asiatica[158]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of the family Osmundaceae

Osmundacaulis sinica[158]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of the family Osmundaceae

Ovoidites circumplicatus[159]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

 Argentina

A green alga belonging to the group Zygnematales.

Ovoidites tripartitus[159]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

 Argentina

A green alga belonging to the group Zygnematales.

Palaeostachya guanglongii[160]

Sp. nov

In press

Liu et al.

Permian (Asselian)

Taiyuan Formation

 China

A member of the family Calamitaceae.

Parazosterophyllum[146]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

 Australia

A member of Zosterophyllaceae. Genus includes new species P. timsiae.

Patruliuspora oregonica[137] Sp. nov Valid Bucur & Rigaud in Bucur et al. Late Triassic (Norian)  United States
( Oregon)
A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Patruliuspora pacifica[137] Sp. nov Valid Bucur, Del Piero & Peyrotty in Bucur et al. Late Triassic (Norian)  Canada
( Yukon)
A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Pellites hamiensis[161] Sp. nov Valid Li et al. Middle Jurassic Xishanyao  China A liverwort belonging to the family Pelliaceae.

Plenasium (Aurealcaulis) elegans[162]

Sp. nov

In press

Hiller et al.

Eocene

Na Duong Formation

 Vietnam

A member of Osmundaceae

Pleuromeia shaolinii[163]

Sp. nov

Valid

Zhang & Wang in Zhang et al.

Middle Triassic

Linjia

 China

Polycingulatisporites multiverrucata[136]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla

 Argentina

Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020.

Polypodiisporites minutiverrucatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

Polysporia baetica[164]

Sp. nov

In press

Álvarez-Vázquez, Bek & Drábková

Carboniferous (Pennsylvanian)

Peñarroya-Belmez-Espiel Coalfield

 Spain

A member of Isoetales

Polystichum pacltovae[165]

Sp. nov

Valid

Kvaček in Kvaček & Teodoridis

Oligocene

 Czech Republic

A fern, a species of Polystichum

Proodontosoria[166]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.

Proterocladus antiquus[167]

Sp. nov

Valid

Tang et al.

Mesoproterozoic
circa 1 Ga

Nanfen

 China

An early siphonocladalean chlorophyte

Psilatriletes cozzuolii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

Psilochara monevaensis[168]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Middle Miocene

 Spain

A charophyte.

Qianshouia[169]

Gen. et sp. nov

Valid

Huang et al.

Late Devonian

Wutong Formation

 China

A plant of uncertain phylogenetic placement, possibly a

sphenopsid
. Genus includes new species Q. mira.

Scolecopteris minuta[170]

Sp. nov

In press

Wan et al.

Early Permian

Taiyuan Formation

 China

A fern belonging to the group

Marattiales
.

Sigillaria pfefferkornii[171]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

 China

Sigillaria wudensis[171]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

 China

Sphaerochara miocenica[168]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Miocene

 Lebanon
 Spain

A charophyte.

Thyrsopteris cretacea[172]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Thyrsopteris

Tomiostrobus sinensis[173]

Sp. nov

Valid

Feng in Feng et al.

Early Triassic (Induan)

Kayitou Formation

 China

A member of the family Isoetaceae.

Tumidopteris astra[174]

Sp. nov

Valid

Naugolnykh

Permian (Roadian)

Pechora coal basin

 Russia

A fern belonging to the family Gleicheniaceae.

Ufadendron elongatum[175]

Sp. nov

Valid

Tang et al.

Late Permian

Linxi

 China

A

lycopsid belonging to the family Tomiodendraceae

Uzhurodendron[176]

Gen. et sp. nov

Valid

Mosseichik & Filimonov

Carboniferous (Tournaisian)

Bystrianskaya

 Russia
( Krasnoyarsk Krai)

A member of Lycopodiopsida. Genus includes new species U. asiaticum.

Zeilleria fosteri[177]

Sp. nov

Valid

Thomas et al.

Carboniferous (Bashkirian)

 United Kingdom

A fern

General research

References

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