2020 in paleoichthyology

New taxa

Jawless vertebrates

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Clavulaspis[1]	Nom. nov	Valid	Elliott, Lassiter & Geyer	Devonian (Givetian)	Yahatinda Formation	Canada ( Alberta)	A member of the family Pteraspididae; a replacement name for Helaspis Elliott et al. (2000).
Ecphymaspis[1]	Gen. et comb. nov	Valid	Elliott, Lassiter & Geyer	Devonian (Eifelian)	Spring Mountain beds	United States ( Idaho)	A member of the family Pteraspididae; a new genus for "Psephaspis" idahoensis Denison (1968).
Parathelodus liaokuoensis[2]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xishancun Xitun	China	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Parathelodus wangi[2]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xishancun Xitun	China	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Parathelodus xitunensis[2]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xitun	China	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Psarkosteus[3]	Gen. et sp. nov	Valid	Dec	Devonian (Givetian)	Skały	Poland	A member of the family Psammosteidae. The type species is P. mediocris.
Rumporostralis[4]	Gen. et comb. et sp. nov	Valid	Shan et al.	Silurian (Telychian)	Xikeng Formation	China	A member of Galeaspida belonging to the group Eugaleaspiformes and the family Sinogaleaspidae. The type species is "Sinogaleaspis" xikengensis Pan & Wang (1980); genus also includes new species R. shipanensis.
Scutellaspis[1]	Gen. et sp. nov	Valid	Elliott, Lassiter & Geyer	Devonian (Eifelian)	Spring Mountain beds	United States ( Idaho)	A member of the family Pteraspididae. Genus includes new species S. wilsoni.

Placoderms

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Actinolepis zaikai[5]	Sp. nov	Valid	Plax & Newman	Devonian (Emsian)	Lepel Beds	Belarus
Bulongosteus[6]	Gen. et sp. nov	Valid	Liu, Zong & Gong	Late Devonian	Zhulumute Formation	China	A member of the family Selenosteidae. Genus includes new species B. liui.
Johannaspis[7]	Gen. et comb. nov	Valid	Vaškaninová	Early Devonian	Prague Basin	Czech Republic	A homostiid arthrodire. Genus includes "Asterolepis" bohemicus Barrande (1872).
Minjinia[8]	Gen. et sp. nov	Valid	Brazeau et al.	Devonian (Pragian)		Mongolia	A placoderm closely related to the gnathostome crown group. Genus includes new species M. turgenensis.
Stipatosteus[5]	Gen. et sp. nov	Valid	Plax & Newman	Devonian (Emsian)	Lepel Beds	Belarus	A phlyctaeniid arthrodire. Genus includes new species S. svidunovitchi.

Acanthodians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Fallodentus[9]	Gen. et sp. nov	In press	Newman, Den Blaauwen & Burrow	Devonian (Givetian)	Mey Flagstone Formation	United Kingdom	A cheiracanthid acanthodian. Genus includes new species F. davidsoni.

Cartilaginous fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Amamriabatis[10]	Gen. et sp. nov	Valid	Adnet et al.	Eocene		Tunisia  Egypt?	A ray belonging to the family Mobulidae. The type species is A. heni.
Asteracanthus dunaii[11]	Sp. nov	Valid	Szabó & Főzy	Jurassic		Hungary
Atlantitrygon[12]	Gen. et comb. et sp. nov	Valid	Sambou et al.	Paleocene (Thanetian) and Eocene (Lutetian)	Matam	Niger  Senegal  Togo	A putative marine potamotrygonid. The type species is "Dasyatis" sudrei Cappetta (1972); genus also includes new species A. senegalensis.
Canadodus[13]	Gen. et sp. nov	Valid	Popov, Johns & Suntok	Late Oligocene	Sooke	Canada ( British Columbia)	A member of the family Chimaeridae. Genus includes new species C. suntoki.
Carcharhinus kasserinensis[10]	Sp. nov	Valid	Adnet et al.	Eocene (Bartonian)		Tunisia  Jamaica?	A species of Carcharhinus.
Coupatezia cristata[10]	Sp. nov	Valid	Adnet et al.	Eocene		Egypt  Tunisia	A ray belonging to the superfamily Dasyatoidea.
Cretasquatina[14]	Gen. et sp. nov	Valid	Maisey, Ehret & Denton	Late Cretaceous (Campanian)	Mooreville Chalk Navesink Formation?	United States ( Alabama  New Jersey?)	A member of the family Squatinidae . The type species is C. americana.
Eoplatyrhina[15]	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca	Italy	A member of the family Platyrhinidae. Genus includes "Platyrhina" bolcensis Heckel (1851).
Eorhinobatos[16]	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca	Italy	A guitarfish; a new genus for "Rhinobatos" primaevus.
Ferromirum[17]	Gen. et sp. nov	Valid	Frey et al.	Devonian (Famennian)	Ibâouane	Morocco	A member of Symmoriiformes . The type species is F. oukherbouchi.
Himantura souarfortuna[10]	Sp. nov	Valid	Adnet et al.	Eocene (Bartonian)		Tunisia	A whiptail stingray, a species of Himantura.
Lamarodus[18]	Gen. et sp. nov	Valid	Ivanov in Ivanov et al.	Permian (Guadalupian)	Bell Canyon Formation	United States ( Texas)	A member of Hybodontoidea . Genus includes new species L. triangulus. Announced in 2018; the final version of the article naming it was published in 2020.
Leptocharias tunisiensis[10]	Sp. nov	Valid	Adnet et al.	Eocene		Tunisia  Egypt?  Jordan?	A relative of the barbeled houndshark.
Mecotrygon[10]	Gen. et sp. nov	Valid	Adnet et al.	Eocene		Egypt  Tunisia	A whiptail stingray belonging to the subfamily Neotrygoninae. The type species is M. asperodentulus.
Mennerotodus mackayi[19]	Sp. nov	Valid	Cicimurri, Ebersole & Martin	Paleocene (Danian)	Clayton	United States ( Alabama)	A sand shark
Mennerotodus parmleyi[19]	Sp. nov	Valid	Cicimurri, Ebersole & Martin	Eocene (Bartonian)	Clinchfield	United States ( Georgia (U.S. state))	A sand shark
Microtriftis[12]	Gen. et sp. nov	Valid	Sambou et al.	Eocene (Ypresian to Lutetian)	Matam	Senegal	A dasyatoid of uncertain phylogenetic placement. Genus includes new species M. matami.
Nanocetorhinus zeitlingeri[20]	Sp. nov	Valid	Feichtinger, Pollerspöck & Harzhauser	Oligocene (Chattian)	Eferding	Austria	A member of Neoselachii of uncertain phylogenetic placement.
Odontorhytis priemi[12]	Sp. nov	Valid	Sambou et al.	Paleocene (Thanetian) and Eocene (Ypresian)	Matam	Mali  Morocco  Senegal  Tunisia	A member of Neoselachii of uncertain phylogenetic placement
Ouledia lacuna[10]	Sp. nov	Valid	Adnet et al.	Eocene (Bartonian)		Tunisia	A butterfly ray.
Pachygymnura[10]	Gen. et comb. nov	Valid	Adnet et al.	Eocene		Egypt  Tunisia	A butterfly ray. The type species is "Coupatezia" attiai Cook in Murray et al. (2010).
Plesiozanobatus[15]	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca	Italy	A relative of panray. Genus includes "Torpedo" egertoni De Zigno (1876).
Pristiophorus austriacus[21]	Sp. nov	Valid	Reinecke et al.	Miocene (Aquitanian)	Ebelsberg	Austria	A species of Pristiophorus.
Pristiophorus borealis[21]	Sp. nov	Valid	Reinecke et al.	Oligocene (Chattian)	Sülstorf	Germany	A species of Pristiophorus.
Pristiophorus tortonicus[21]	Sp. nov	Valid	Reinecke et al.	Miocene (Tortonian)	Mica Clay	Germany	A species of Pristiophorus.
Pristiophorus ungeri[21]	Sp. nov	Valid	Reinecke et al.	Miocene (Burdigalian)	Neuhofen	Germany	A species of Pristiophorus.
Pseudorhinobatos[16]	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca	Italy	A guitarfish; a new genus for "Rhinobatos" dezignii.
Sculptospina[22]	Gen. et sp. nov	Valid	Lebedev in Lebedev, Ivanov & Linkevich	Devonian (Famennian)		Russia ( Lipetsk Oblast)	A member of Ctenacanthiformes of uncertain phylogenetic placement. The type species is S. makhlaevi.
Sphyrna guinoti[10]	Sp. nov	Valid	Adnet et al.	Eocene		Tunisia  Egypt?	A species of Sphyrna.
Stegostoma tethysiensis[10]	Sp. nov	Valid	Adnet et al.	Eocene		Tunisia  Egypt?	A relative of the zebra shark.
Taklamakanolepis[23]	Gen. et sp. nov		Andreev et al.	Early Silurian	Ymogantau	China	A member of Mongolepidida. The type species is T. asiaticus.
Tielikewatielepis[23]	Gen. et sp. nov		Andreev et al.	Early Silurian	Tataertag Ymogantau	China	A member of Mongolepidida. The type species is T. sinensis.
Xiaohaizilepis[23]	Gen. et sp. nov		Andreev et al.	Early Silurian	Tataertag Ymogantau	China	A member of Mongolepidida. The type species is X. liui.
Yuanolepis[23]	Gen. et sp. nov		Andreev et al.	Early Silurian	Ymogantau	China	A cartilaginous fish of uncertain phylogenetic placement. The type species is Y. bachunensis.

Ray-finned fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Agassizilia[24]	Gen. et sp. nov	Valid	Cooper & Martill	Cenomanian	Kem Kem	Morocco	Possibly a member of the family Pycnodontidae. The type species is A. erfoudina.
Altmuehlfuro[25]	Gen. et sp. nov	Valid	Ebert	Late Jurassic	Solnhofen	Germany	A member of Halecomorphi belonging to the group Ophiopsiformes. Genus includes new species A. boomerang.
Amakusaichthys[26]	Gen. et sp. nov	Valid	Yabumoto, Hirose & Brito	Late Cretaceous (Santonian)	Hinoshima	Japan	A member of Ichthyodectiformes. Genus includes new species A. goshouraensis. Announced in 2018; the final version of the article naming it was published in 2020.
Ampheristus americanus[27]	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian) and Paleocene (Danian)	Clayton Kemp Clay	United States ( Arkansas  Texas)	A cusk-eel .
Anguilla? chickasawae[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian) and Paleocene (Danian)[27]	Clayton[27] Ripley	United States ( Arkansas[27]  Mississippi)	A member or a relative of the family Anguillidae
Anomoeodus wolfi[29]	Sp. nov	Valid	Capasso	Late Cretaceous (Cenomanian)	Del Rio	United States ( Texas)	A member of the family Pycnodontidae.
Anorevus[30]	Gen. et sp. nov.	Valid	Bannikov & Zorzin	Eocene (Ypresian)	Monte Bolca	Italy	A member of Percomorpha of uncertain phylogenetic placement. The type species is A. lorenzonii.
Apateodus crenellatus[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley	United States ( Mississippi)
Ariopsis castilloensi[31]	Sp. nov	In press	Aguilera & Rodriguez in Aguilera et al.	Late Oligocene to early Miocene	Castillo	Venezuela	A species of Ariopsis.
Armigatus carrenoae[32]	Sp. nov	Valid	Alvarado-Ortega, Than-Marchese & Melgarejo-Damián	Early Cretaceous (Albian)	Tlayúa	Mexico	A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Armigatidae .
Bagre ornatus[31]	Sp. nov	In press	Aguilera & Rodriguez in Aguilera et al.	Late Oligocene to early Miocene	Castillo	Venezuela	A species of Bagre.
Baringochromis[33]	Gen. et 3 sp. nov	Valid	Altner & Reichenbacher	Late Miocene		Kenya	A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is B. senutae; genus also includes B. sonyii and B. tallamae. Announced in 2020; the final version of the article naming it was published in 2021.
Benthophilus aprutinus[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy	A tadpole goby.
Benthophilus labronicus[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy	A tadpole goby.
Boreolates[35]	Gen. et sp. nov	Valid	Weems	Eocene	Nanjemoy	United States ( Virginia)	A member of the family Latidae. The type species is B. debernardi.
Brachyplatystoma elbakyani[36]	Sp. nov	In press	Agnolin & Bogan	Late Miocene		Argentina	A species of Brachyplatystoma
Buenia pisiformis[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian and Messinian)		Greece	A species of Buenia.
Burguklia minichorum[37]	Sp. nov	Valid	Bakaev & Kogan	Permian (Wordian–Capitanian)		Russia	An early ray-finned fish. Announced in 2019; the final version of the article naming it was published in 2020.
Caspiosoma lini[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy  Romania	A species of Caspiosoma .
Caspiosoma paulisulcata[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy	A species of Caspiosoma.
Caucombrus[38]	Gen. et sp. nov	Valid	Bannikov	Early Oligocene		Abkhazia  Russia	A member of the family Scombridae. Genus includes new species C. histiopterygius.
Chaychanus[39]	Gen. et sp. nov	Valid	Cantalice Severiano, Alvarado Ortega & Bellwood	Paleocene		Mexico	A member of the family Pomacentridae. Genus includes new species C. gonzalezorum. Announced in 2019; the final version of the article naming it was published in 2020.
Cheirolepis aleshkai[40]	Sp. nov	Valid	Plax	Devonian (Eifelian)		Belarus	An early ray-finned fish.
Choctawichthys[28]	Gen. et comb. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)	A member of Paracanthopterygii of uncertain phylogenetic placement. The type species is "genus Perciformorum" cepoloides Nolf & Dockery (1990).
Chromogobius? primigenius[41]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene			Possibly a species of Chromogobius.
Clupea hanishinaensis[42]	Nom. nov	Valid	Yabumoto & Nazarkin	Miocene	Bessho	Japan	A species of Clupea; a replacement name for Clupea macrocephala Yabumoto & Nazarkin (2018).
Congrophichthus[27]	Gen. et sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (late Campanian and Maastrichtian) and Paleocene (Danian)	Clayton Coffee Sand Kemp Clay	United States ( Alabama  Arkansas  Mississippi  Texas)	A member of the family Congridae. The type species is C. transterminus.
Cowetaichthys carnevalei[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)	A beardfish
Cyclothone duhoensis[43]	Sp. nov	Valid	Nam & Nazarkin	Miocene	Duho	South Korea	A species of Cyclothone. Announced in 2020; the final version of the article naming it was published in 2021.
Dakotaichthys[27]	Gen. et sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian)	Fox Hills Kemp Clay	United States ( North Dakota  Texas)	Possibly a member of the family Gadidae. The type species is D. hogansoni.
Eleotris omuamuaensis[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian)		Italy	A species of Eleotris.
Eleotris tyrrhenicus[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian)		Italy	A species of Eleotris.
Ellimma longipectoralis[44]	Sp. nov	Valid	Polck et al.	Early Cretaceous (Aptian)	Barra Velha Formation	Brazil	A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae . Announced in 2019; the final version of the article naming it was published in 2020.
Ellimmichthys spinosus[45]	Sp. nov	In press	De Figueiredo & Gallo	Early Cretaceous	Recôncavo Basin	Brazil	A member of Clupeomorpha belonging to the group Ellimmichthyiformes.
Elongofuro[25]	Gen. et sp. nov	Valid	Ebert	Late Jurassic		Germany	A member of Halecomorphi belonging to the group Ophiopsiformes. Genus includes new species E. woelflei.
Elopothrissus carsonsloani[27]	Sp. nov	Valid	Schwarzhans & Stringer	Paleocene (Danian)	Clayton	United States ( Arkansas)	A relative of the Japanese gissu.
Engdahlichthys[46]	Gen. et sp. nov	Valid	Murray et al.	Paleocene	Fort Union	United States ( Montana)	A sturgeon. Genus includes new species E. milviaegis.
Enigmacottus[34]	Gen. et sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy	Possibly a member of the family Psychrolutidae. The type species is E. socialis.
Epaelops[47]	Gen. et sp. nov	In press	Alves, Alvarado Ortega & Brito	Early Cretaceous (Albian)	Tlayúa	Mexico	A member of Elopiformes. Genus includes new species E. martinezi. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Eutawichthys choctawae[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Kemp Clay[27] Ripley	United States ( Mississippi  Texas[27])	Probably a member of Beryciformes
Feroxichthys[48]	Gen. et sp. nov	Valid	Xu	Middle Triassic (Anisian)	Guanling	China	A member of the family Colobodontidae. The type species is F. yunnanensis.
Globanomoeodus[29]	Gen. et sp. nov	Valid	Capasso	Late Cretaceous (Cenomanian)	Del Rio	United States ( Texas)	A member of the family Pycnodontidae. The type species is G. dentespassim.
Gobius peloponnesus[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Pliocene (Zanclean)		Greece	A species of Gobius.
Gobius supraspectabilis[41]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene		Slovakia	A species of Gobius.
Gregoriopycnodus[49]	Gen. et comb. nov	Valid	Taverne, Capasso & Del Re	Early Cretaceous (Albian)		Italy	A member of the family Pycnodontidae. The type species is "Palaeobalistum" bassanii d'Erasmo (1914).
Hadromos[50]	Gen. et sp. nov	Valid	Murray	Early Cenozoic (possibly Eocene)	Sangkarewang	Indonesia	A member of Cyprinoidea . Genus includes new species H. sandersae.
Hastichthys totonacus[51]	Sp. nov	In press	Alvarado-Ortega & Díaz-Cruz	Late Cretaceous (Turonian)	Agua Nueva	Mexico	A member of Aulopiformes belonging to the family Dercetidae.
Heckelichthys preopercularis[52]	Sp. nov	In press	Baños Rodríguez et al.	Cretaceous (Albian–Cenomanian)	El Doctor	Mexico	A member of Ichthyodectiformes
Hesperichthys gironeae[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian and Messinian)		Italy	A goby.
Hesperichthys iugosus[41]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene		Moldova	A goby. Originally described as a species of Hesperichthys, but subsequently transferred to the genus Sarmatigobius.[53]
Hiascoactinus[54]	Gen. et sp. nov	In press	Kim et al.	Late Triassic	Amisan	South Korea	A member of Redfieldiiformes. Genus includes new species H. boryeongensis. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Hoeseichthys[41]	Gen. et comb. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene and Pliocene[55]		Czech Republic  Germany  India[56]  Italy[34]  Morocco[55]	A goby. Genus includes "Otolithus (Gobius)" praeclarus Procházka (1893) and H. laevis (Weiler, 1942), as well as "Gobiida" bicornuta Lin, Girone & Wolf (2015) and "Gobiida" brioche Lin, Girone & Wolf (2015).[34]
Holzmadenfuro[57]	Gen. et sp. nov	Valid	Ebert, Thies & Hauff	Early Jurassic (Toarcian)	Posidonia	Germany	A member of Halecomorphi. Genus includes new species H. rebmanni.
Hoplopteryx langfordi[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)
Hypsocormus posterodorsalis[58]	Sp. nov	Valid	Maxwell et al.	Late Jurassic (Kimmeridgian)	Nusplingen	Germany
Knipowitschia etrusca[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy	A species of Knipowitschia.
Knipowitschia polonica[41]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene		Poland  Ukraine[59]	A species of Knipowitschia.
Kokenichthys navis[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley	United States ( Mississippi)	A member of Osteoglossiformes of uncertain phylogenetic placement
Kradimus[60]	Gen. et sp. nov	Valid	Veysey, Brito & Martill	Late Cretaceous (Turonian)	Akrabou Formation	Morocco	A member of Crossognathiformes. Genus includes new species K. asflaensis. Announced in 2019; the final version of the article naming it was published in 2020.
Lavinia stuwilliamsi[61]	Sp. nov	Valid	McClellan & Smith	Late Miocene	Salt Lake Formation	United States ( Utah)	A species of Lavinia .
Lebrunichthys[62]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)		Lebanon	A member of Pachyrhizodontidae . The type species is L. nammourensis.
Lesueurigobius stazzanensis[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian)		Italy	A species of Lesueurigobius.
Lindoeichthys[63]	Gen. et sp. nov.	Valid	Murray et al.	Late Cretaceous (Maastrichtian)	Scollard Formation	Canada ( Alberta)	A member of Percopsiformes. Genus includes new species L. albertensis. Announced in 2019; the final version of the article naming it was published in 2020.
Louwoichthys[64]	Gen. et sp. nov	Valid	Xu	Middle Triassic (Anisian)		China	A member of the stem group of Neopterygii. Genus includes new species L. pusillus.
Luganoia fortuna[65]	Sp. nov	Valid	Xu	Middle Triassic (Ladinian)	Falang	China
Megalops? nolfi[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)	Probably member of the family Megalopidae
Monosmilus[66]	Gen. et sp. nov		Capobianco et al.	Eocene (Lutetian)	Domanda	Pakistan	A stem-engraulid. The type species is M. chureloides.
Muraenanguilla unionensis[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley	United States ( Mississippi)	An eel of uncertain phylogenetic placement
Neilpeartia[67]	Gen. et sp. nov	Valid	Carnevale et al.	Eocene (Ypresian)	Monte Bolca	Italy	A frogfish. Genus includes new species N. ceratoi.
Neomesturus[68]	Gen. et sp. nov	In press	Cooper & Martill	Turonian	Akrabou	Morocco	A member of Pycnodontiformes. Genus includes new species N. asflaensis.
Neoproscinetes africanus[24]	Sp. nov	Valid	Cooper & Martill	Cenomanian	Kem Kem	Morocco	A member of the family Pycnodontidae
Njoerdichthys[69]	Gen. et sp. nov	In press	Cawley et al.	Late Cretaceous (Turonian)	Hesseltal	Germany	A member of the family Pycnodontidae. The type species is N. dyckerhoffi.
Nunaneichthys[70]	Gen. et sp. nov	In press	Hernández-Guerrero et al.	Cretaceous (Albian-Cenomanian)		Mexico	A Pterothrissinae . Genus includes new species N. mexicanus.
Odondebuenia agiadiae[41]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene		Poland  Slovakia  Ukraine[59]	A relative of the coralline goby.
Ohmdenfuro[57]	Gen. et sp. nov	Valid	Ebert, Thies & Hauff	Early Jurassic (Toarcian)	Posidonia	Germany	A member of Halecomorphi. Genus includes new species O. bodmani.
Osmeroides mississippiensis[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley	United States ( Mississippi)
Ossulcus[28]	Gen. et sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)	Probably a member of Beryciformes. The type species is O. labiatus.
Otolithopsis cumatilis[28]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)	An otolith of a fish of uncertain phylogenetic placement.
Palaeogadus weltoni[27]	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian)	Kemp Clay	United States ( Texas)
Palealbula korchinskyi[71]	Sp. nov	Valid	Schwarzhans & Mironenko	Late Jurassic		Russia	Possibly a albuliform
Palealbula moscoviensis[71]	Sp. nov	Valid	Schwarzhans & Mironenko	Late Jurassic		Russia	Possibly a albuliform .
Paranursallia cavini[68]	Sp. nov	In press	Cooper & Martill	Turonian	Akrabou	Morocco	A member of Pycnodontiformes.
Pauciuncus[50]	Gen. et comb. nov	Valid	Murray	Early Cenozoic (possibly Eocene)	Sangkarewang	Indonesia	A member of Cyprinoidea. Genus includes "Puntius" bussyi.
Polcynichthys[72]	Gen. et sp. nov	In press	London & Shimada	Late Cretaceous (Cenomanian)	Tarrant	United States ( Texas)	A member of the family Pachyrhizodontidae . Genus includes new species P. lloydhilli.
Proterorhinus cretensis[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Pliocene (Zanclean)		Greece	A species of Proterorhinus.
Pythonichthys arkansasensis[27]	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian) and Paleocene (Danian)	Clayton Kemp Clay	United States ( Arkansas  Texas)	A species of Pythonichthys.
Rachycentron stremphaencus[73]	Sp. nov	Valid	Godfrey & Carnevale	Miocene (Tortonian)	St. Marys	United States ( Maryland)	A relative of the cobia.
Rebekkachromis valyricus[74]	Sp. nov	Valid	Kevrekidis & Reichenbacher in Kevrekidis et al.	Miocene	Ngorora	Kenya	A cichlid belonging to the tribe Oreochromini.
Rebekkachromis vancouveringae[74]	Sp. nov	Valid	Kevrekidis & Reichenbacher in Kevrekidis et al.	Miocene	Ngorora	Kenya	A cichlid belonging to the tribe Oreochromini.
Rhynchoconger brettwoodwardi[27]	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian)	Kemp Clay	United States ( Texas)	A species of Rhynchoconger.
Rupelia[75]	Gen. et comb. nov	Valid	Baykina & Kovalchuk in Kovalchuk et al.	Oligocene (Rupelian)		Russia	A member of family Clupeidae. Genus includes R. rata (Daniltshenko, 1959).
Salwaichthys[76]	Gen. et sp. nov	Valid	Bannikov	Early Oligocene		Abkhazia  Poland  Russia ( Adygea)	A member of Perciformes belonging to the new family Salwaichthyidae. Genus includes new species S. paratethyensis.
Sanctusichthys[77]	Gen. et sp. nov	Valid	López-Arbarello, Maxwell & Schweigert	Late Jurassic (Kimmeridgian)	Nusplingen Limestone	Germany	A member of Halecomorphi. Genus includes new species S. rieteri.
Sangkarewangia[50]	Gen. et sp. nov	Valid	Murray	Early Cenozoic (possibly Eocene)	Sangkarewang	Indonesia	A member of Cyprinoidea. Genus includes new species S. sumatranus.
Scleropages sanshuiensis[78]	Sp. nov	Valid	Zhang	Early Eocene	Huachong	China	A species of Scleropages. Announced in 2019; the final version of the article naming it was published in 2020.
Scombroclupea javieri[79]	Sp. nov	In press	Than Marchese et al.	Late Cretaceous (Cenomanian)	Cintalapa	Mexico	A member of Clupeomorpha of uncertain phylogenetic placement
Siniperca ikikoku[80]	Sp. nov	Valid	Yabumoto	Miocene		Japan	A species of Siniperca.
Simocormus[58]	Gen. et sp. nov	Valid	Maxwell et al.	Late Jurassic (Kimmeridgian–Tithonian)		Germany	A member of the family Pachycormidae . The type species is S. macrolepidotus.
Stanhopeichthys[81]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)		Lebanon	A member of Pachyrhizodontidae . The type species is S. libanicus.
Stefanichthys[82]	Gen. et sp. nov.	Valid	Bannikov & Zorzin	Eocene]] (Ypresian)	Monte Bolca	Italy	A member of Percoidei of uncertain phylogenetic placement. The type species is S. mariannae.
Thorogobius petilus[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy	A species of Thorogobius.
Thrax[28]	Gen. et sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley	United States ( Mississippi)	A member of Aulopiformes belonging to the family Ichthyotringidae. The type species is T. acutus.
Tippaha[28]	Gen. et sp. et comb. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley	United States ( Mississippi)	A member of Holocentriformes of uncertain phylogenetic placement. The type species is T. mythica; genus also includes "genus Ophidiidarum" cavatus Nolf & Stringer (1996).
Vachalia[83]	Gen. et sp. nov	Valid	Přikryl & Carnevale	Early Oligocene		Czech Republic	A member of the family Platytroctidae. Genus includes new species V. moraviensis.
Vanderhorstia prochazkai[41]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene		Ukraine[59]	A species of Vanderhorstia.
Vegrandichthys[84]	Gen. et sp. nov	Valid	Díaz-Cruz, Alvarado-Ortega & Giles	Late Cretaceous (Cenomanian)	Cintalapa	Mexico	A member of Aulopiformes belonging to the family Enchodontidae. The type species is V. coitecus.
Warilochromis[85]	Gen. et sp. nov	Valid	Altner, Ruthensteiner & Reichenbacher	Late Miocene	Ngorora	Kenya	A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is W. unicuspidatus.
Zosterisessor exsul[34]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)		Italy  Romania	A relative of the grass goby.

Lobe-finned fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Ferganoceratodus annekempae[86]	Sp. nov	Valid	Cavin, Deesri & Chanthasit	Late Jurassic	Phu Kradung	Thailand	A lungfish.
Megalichthys mullisoni[87]	Sp. nov	Valid	Downs & Daeschler	Devonian (Famennian)	Catskill	United States ( Pennsylvania)
Metaceratodus baibianorum[88]	Sp. nov	Valid	Panzeri et al.	Late Cretaceous	La Colonia	Argentina	A lungfish.
Neoceratodus potkooroki[89]	Sp. nov	Valid	Kemp & Berrell	Cretaceous		Australia	A lungfish.
Ptychoceratodus oldhami[90]	Sp. nov	Valid	Bhat & Ray	Late Triassic (Carnian)	Tiki	India	A lungfish. Announced in 2018; the final version of the article naming it was published in 2020.
Rossichthys[91]	Gen. et sp. nov	Valid	Johanson et al.	Carboniferous (Tournaisian)	Ballagan Formation	United Kingdom	A member of Rhizodontida. The type species is R. clackae.

Research

• A study on the morphology of the osteostracans, evaluating different methods used to determine the morphological variation within this group and its evolution, is published by Ferrón et al. (2020).^[92]
• A study on the morphological diversity of osteostracan headshields, aiming to determine the relationship between their morphological diversity and hydrodynamic performance and its implications for the knowledge of the ecological diversity of the osteostracans, is published by Ferrón et al. (2020).^[93]
• A study aiming to test the alternative hypotheses of
  placoderm jaw bone homologies, and evaluating their implications for the knowledge of evolution of jaw bones in early jawed vertebrates, is published by King & Rücklin (2020).^[94]
• A study on the fossil dentitions of acanthothoracids is published by Vaškaninová et al. (2020), who report that the teeth of acanthothoracids differed fundamentally from those of arthrodires, and argue that the characteristic traits of acanthothoracid dentition might be ancestral for all jawed vertebrates.^[95]
• Redescription of the anatomy of Walterilepis speciosa, based on data from new fossil material, and a study on the phylogenetic relationships of this species is published online by Lukševičs (2020).^[96]
• Description of the neurocranial anatomy of Ellopetalichthys scheii is published by Castiello et al. (2020).^[97]
• A study aiming to determine whether
  suspension feeder, focusing on mechanical properties of its jaw, is published by Coatham et al. (2020).^[98]
• The earliest fossilized vertebrate embryos reported so far, preserved with an adult specimen of Watsonosteus fletti from the Givetian Eday Flagstone Formation (Orcadian Basin; Scotland, United Kingdom), are described by Newman et al. (2020).^[99]
• Burrow, Newman & den Blaauwen (2020) describe external spiracular elements in Middle Devonian acanthodians from northern Scotland, differing from spiracles of all known extant and extinct fishes, and report the oldest record of elastic cartilage in the fossil record.^[100]
• A study examining the factors influencing the long-term variations of genus-level diversity of elasmobranchs and ray-finned fishes throughout their evolutionary history is published by Guinot & Cavin (2020).^[101]
• Carrillo-Briceño et al. (2020) describe a new elasmobranch assemblage from the Oligocene–Miocene boundary in the Dos Bocas Formation (Ecuador), and evaluate the implications of this assemblage for chronostratigraphic inferences and the knowledge of local paleoenvironment.^[102]
• Two large vertebrae of sharks belonging to the genus
  Ptychodontidae, are described from the Santonian of Spain by Jambura & Kriwet (2020).^[103]
• A study on the evolution of body size in lamniform sharks, including the evolution of gigantism in the lineage of Otodus megalodon, is published by Shimada, Becker & Griffiths (2020).^[104]
• Taxonomic revision of the Oligocene and Miocene sand sharks is published by Hovestadt (2020).^[105]
• A study aiming to determine the linear body dimensions of Otodus megalodon at different life stages is published by Cooper et al. (2020).^[106]
• A study on the class structure of assemblages of specimens of Otodus megalodon in eight previously known formations and in a newly described Miocene locality from northeastern Spain is published by Herraiz et al. (2020), who interpret their findings as indicative of existence of five potential nurseries of these sharks ranging from the Langhian to the Zanclean.^[107]
• An isolated tooth of Cosmopolitodus hastalis from the Miocene of South Korea is described by Yun (2020).^[108]
• A study on the fossil record of the great white shark from the Pliocene of Peru and Chile is published by Villafaña et al. (2020), who interpret their findings as indicating that great white sharks used the Coquimbo locality in Chile as a nursery and Pisco (Peru) and Caldera (Chile) localities as feeding grounds during the Pliocene.^[109]
• A study on the anatomy and phylogenetic relationships of "Urolophus" crassicaudatus is published by Marramà et al. (2020), who transfer this species to the genus Arechia.^[110]
• Collareta et al. (2020) describe a fossil stinger a stingray from the Pliocene (Piacenzian) locality La Serra (Italy), twice as long as the longest caudal spines reported from any living stingray species of the Mediterranean Sea, and possibly representing the longest stingray stinger ever reported from both the fossil and the recent records.^[111]
• A study on the morphology of the marginal dentition of Lophosteus superbus is published by Chen et al. (2020), who reconstruct the dental ontogeny in this taxon, and evaluate its implications for the knowledge of the evolution of teeth of bony fishes.^[112]
• Redescription of the anatomy of Tanyrhinichthys mcallisteri is published by Stack et al. (2020).^[113]
• Fragmentary fossil material of Gyrosteus mirabilis is reported from the Toarcian of the Ahrensburg erratics assemblage (Schleswig-Holstein, Germany) by Hornung & Sachs (2020), expanding known geographic range of this species, and representing the first record of a chondrosteid species beyond its type area.^[114]
• Redescription of the skeletal anatomy of Yanosteus longidorsalis is published by Hilton, Grande & Jin (2020).^[115]
• Revision and a study on the phylogenetic relationships of members of the subfamily Pycnodontinae is published by Poyato-Ariza (2020).^[116]
• A study on the skeletal anatomy and phylogenetic relationships of Lombardina decorata is published by Taverne (2020).^[117]
• A study on the degree of preservation of the skin of an
  aspidorhynchid specimen from the Barremian Paja Formation (Colombia), representing the first instance of soft tissue preservation in vertebrates from the Early Cretaceous of northern South America, is published by Alfonso-Rojas & Cadena (2020).^[118]
• A study on the diversity and distribution of non-marine teleost fishes in the Western Interior of North America during the late Maastrichtian, based on fossils from the Hell Creek Formation, the Lance Formation and the Scollard Formation, is published online by Brinkman et al. (2020).^[119]
• Description of new fossil material of Abisaadichthys libanicus and Eusebichthys byblosi, providing new information on the skeletal anatomy of these taxa, is published by Taverne & Capasso (2020).^[120]
• Fossil material of Xiphactinus is described from the latest Maastrichtian Salamanca Formation (Chubut Province, Argentina) by De Pasqua, Agnolin & Bogan (2020), representing the first record of this genus from southern part of South America.^[121]
• A methodology for assessing locomotion energetics in extinct bony fishes is presented by Ferrón (2020), who interprets his findings as providing evidence of endothermy in Xiphactinus audax.^[122]
• Redescription and a study on the phylogenetic relationships of Laeliichthys ancestralis is published by Brito, Figueiredo & Leal (2020).^[123]
• A study on the skeletal anatomy of Pirskenius, aiming to resolve whether Pirskeniidae can be sustained as a separate family, is published by Reichenbacher et al. (2020).^[124]
• A study aiming to infer the genetic basis of the reduction of pelvic skeleton in a Miocene stickleback fish Gasterosteus doryssus is published by Stuart, Travis & Bell (2020).^[125]
• A fish larva sharing anatomical similarities with the so‐called tholichthys larval stage of butterflyfishes is described from the Eocene (Bartonian) locality of Gornyi Luch (Krasnodar Krai, Russia) by Carnevale & Bannikov (2020).^[126]
• New fossil material of Mawsonia gigas, including one of the anatomically most informative specimens referable to the genus Mawsonia, is described from the Mesozoic Tacuarembó Formation (Uruguay) by Toriño et al. (2020).^[127]
• New fossil material of Axelrodichthys megadromos is described from several Campanian and Maastrichtian sites in southern France by Cavin et al. (2020), who present a reconstruction of the skull of this species, and study its phylogenetic relationships and ecology.^[128]
• Redescription of the anatomy of the skull of Durialepis edentatus is published by Mondéjar‐Fernández, Friedman & Giles (2020).^[129]
• Description of new material of tristichopterids from the Devonian (Famennian) locality of Strud (Belgium), and a study on the phylogenetic relationships of tristichopterids, is published by Olive et al. (2020).^[130]
• Description of a new, 1.57-metre-long articulated specimen of Elpistostege watsoni from the Upper Devonian of Canada, and a study on the implications of this specimen for the knowledge of the early evolution of the vertebrate hand, is published by Cloutier et al. (2020).^[131]
• A study aiming to determine the potential significance of tides for the evolution of bony fish and early tetrapods from the Late Silurian to early Late Devonian is published by Byrne et al. (2020).^[132]
• Evidence of enhanced fish production during the extreme global warmth of the early Paleogene is presented by Britten & Sibert (2020).^[133]
• A study aiming to determine the impact of changes in the Earth system during the Eocene–Oligocene transition on pelagic fish production and biodiversity is published by Sibert et al. (2020).^[134]