2021 in arthropod paleontology
| |||
---|---|---|---|
| |||
2021 in arthropod paleontology is a list of new arthropod fossil taxa, including arachnids, crustaceans, insects, trilobites, and other arthropods that were announced or described, as well as other significant arthropod paleontological discoveries and events which occurred in 2021.
Arachnids
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Schwarze et al. |
Europe (Baltic Sea region) |
A pseudoscorpion belonging to the family Pseudotyrannochthoniidae. |
||||
Gen. et sp. nov |
Valid |
Wunderlich & Müller |
Cretaceous |
Possibly a member of the family Dipluridae. The type species is A. spicula. |
||||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of the family Tetrablemmidae. The type species is A. plenfemur. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the family Pholcochyroceridae. |
|||
Sp. nov |
Valid |
Khaustov et al. |
Late Eocene |
A mite belonging to the group Raphignathoidea and the family Barbutiidae. |
||||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. The type species is B. crassifemora. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the family Salticoididae. |
|||
Burmadictyna fissura[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the family Salticoididae. |
||
Burmadictyna similis[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the family Salticoididae. |
||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Araneoidea and the family Zarqaraneidae. The type species is B. trispinae. |
|||
Nom. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Nemesiidae; a replacement name for Burmesia Wunderlich (2020). |
|||
Sp. nov |
Valid |
Jiang & Li in Xin et al. |
Cretaceous |
Burmese amber |
A spider belonging to the family Oonopidae. |
|||
Burmorchestina convexa[4] |
Sp. nov |
Valid |
Jiang & Li in Xin et al. |
Cretaceous |
Burmese amber |
A spider belonging to the family Oonopidae. |
||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the family Burmorsolidae. |
|||
Sp. nov |
In press |
Schwarze et al. |
Eocene |
Bitterfeld amber
|
A pseudoscorpion belonging to the family Pseudotyrannochthoniidae. |
|||
Sp. nov |
Valid |
Lourenço in Lourenço & Velten |
Cretaceous |
Burmese amber |
A scorpion belonging to family Chaerilobuthidae. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Theridiidae. The type species is C. concavum. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Crassicephalidae. The type species is C. parvibulbus. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Araneoidea and the family Zarqaraneidae. |
|||
Gen. et sp. nov |
Valid |
Lourenço in Lourenço & Velten |
Cretaceous |
Burmese amber |
A scorpion belonging to the family Protoischnuridae. The type species is C. smeelei. |
|||
Gen. et sp. nov |
Valid |
Downen & Selden |
A spider belonging to the family Palpimanidae. Genus includes new species C. vittari. |
|||||
Sp. nov |
Valid |
Khaustov et al. |
Late Eocene |
A mite belonging to the group Heterostigmata and the family Dolichocybidae .
|
||||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Dubiodeinopsidae. The type species is D. spinifemora. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Dubiouloboridae. The type species is D. incompletus. |
|||
Gen. et 2 sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Dubiouloboridae. The type species is D. praeta; genus also includes D. procerembolus. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of the family Tetrablemmidae. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider of uncertain phylogenetic placement, possibly a member of an early branch of the RTA clade. The type species is E. cretaceus. |
|||
Sp. nov |
Valid |
Jiang & Li in Xin et al. |
Late Cretaceous |
Burmese amber |
A spider belonging to the family Pacullidae. |
|||
Furcembolus fengzheni[9] |
Sp. nov |
Valid |
Jiang & Li in Xin et al. |
Late Cretaceous |
Burmese amber |
A spider belonging to the family Pacullidae. |
||
Furcembolus martensi[9] |
Sp. nov |
Valid |
Jiang & Li in Xin et al. |
Late Cretaceous |
Burmese amber |
A spider belonging to the family Pacullidae. |
||
Gen. et sp. nov |
Valid |
Guo, Selden & Ren |
Cretaceous |
Burmese amber |
A spider belonging to the family Lagonomegopidae. Genus includes new species H. spinalis. |
|||
Sp. nov |
Valid |
Khaustov et al. |
Late Eocene |
Rovno amber |
A mite belonging to the group Heterostigmata and the family Tarsocheylidae. |
|||
Gen. et sp. nov |
In press |
Szabó et al. |
Late Cretaceous (Santonian) |
Ajka Coal Formation |
A spider belonging to the family Hersiliidae . Genus includes new species H. verdesi.
|
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the family Pholcochyroceridae. The type species is K. oblonga. |
|||
Gen. et sp. nov |
Valid |
Klimov et al. |
A mite belonging to the group Astigmata . The type species is L. sidorchukae.
|
|||||
Sp. nov |
In press |
Arillo & Subías in Arillo, Subías & Álvarez-Parra |
Early Cretaceous (Albian) |
A mite belonging to the family Liacaridae. |
||||
Sp. nov |
Junior homonym |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the family Pholcochyroceridae. The specific name is preoccupied Longissipalpus cochlea Wunderlich (2017). Wunderlich (2022) coined a replacement name Longissipalpus aliter.[14] |
|||
Longissipalpus impudicus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the family Pholcochyroceridae. |
||
Gen. et sp. nov |
Valid |
Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the new family Megasetidae. The type species is M. colphepeiroides. |
|||
Sp. nov |
Valid |
Mitov, Perkovsky & Dunlop |
Eocene |
A harvestman, a species of Metaphalangium. |
||||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Palpimanoidea and the family Micropalpimanidae. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Theridiidae. The type species is M. longissispinae. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
|||
Nom. nov |
Ceccolini & Cianferoni |
A spider belonging to the family Trochanteriidae; a replacement name for Veterator Petrunkevitch (1963). |
||||||
Sp. nov |
Valid |
Jiang & Li in Xin et al. |
Cretaceous |
Burmese amber |
A spider belonging to the family Oonopidae, a species of Orchestina. |
|||
Gen. et sp. nov |
In press |
Rivas & Vega |
A mite belonging to the family Johnstonianidae. The type species is P. microscutum. |
|||||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Leptonetidae. |
|||
Sp. nov |
Valid |
Selden |
A spider belonging to the group Mesothelae and to the new family Palaeothelidae. |
|||||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Mecysmaucheniidae. The type species is P. depressa. |
|||
Sp. nov |
Valid |
Khaustov et al. |
Late Eocene |
Rovno amber |
A mite belonging to the group Heterostigmata and the family Acarophenacidae. |
|||
Gen. et comb. nov |
Valid |
Mitov, Perkovsky & Dunlop |
Eocene |
Bitterfeld amber Rovno amber |
A harvestman belonging to the family Nemastomatidae; a new genus for "Nemastoma" tuberculatum Koch & Berendt (1854). |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
|||
Paramiagrammopes curvatus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes furca[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes granulatus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes inaequalis[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes inclinatus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes multifemurspinae[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes paracurvatus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes pilosus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes pollex[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes semiapertus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes simplex[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes sulcus[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes texter[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Paramiagrammopes unibrevispina[2] |
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Palpimanoidea and the family Planarchaeidae. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Palpimanoidea and the family Planarchaeidae. The type species is P. longicorpus. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Palpimanoidea and the family Vetiatoridae. The type species is P. circulus. |
|||
Gen. et sp. nov |
Valid |
Wriedt et al. |
Cretaceous |
A pseudoscorpion belonging to the family Chthoniidae. Genus includes new species P. burmiticus. |
||||
Sp. nov |
Valid |
Magalhaes et al. |
Burmese amber |
A spider belonging to the family Psilodercidae. |
||||
Sp. nov |
Valid |
Khaustov et al. |
Late Eocene |
Rovno amber |
A mite belonging to the group Heterostigmata and the family Acarophenacidae. |
|||
Gen. et 2 sp. nov |
In press |
Geißler et al. |
Late Cretaceous (Cenomanian) |
Burmese amber |
A pseudoscorpion belonging to the family Ideoroncidae. Genus includes new species P. gracilis and P. compactus. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Leptonetoidea and the family Protoaraneoididae. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of the family Tetrablemmidae. The type species is P. deformans. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Palpimanoidea and the family Vetiatoridae. The type species is P. fruticosus. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of Araneomorphae belonging to the family Eopsilodercidae. |
|||
Sp. nov |
Valid |
Selden |
Carboniferous (Kasimovian) |
A spider belonging to the group Mesothelae and the family Arthrolycosidae. |
||||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. The type species is P. tuberculatus. |
|||
Gen. et sp. nov |
Valid |
Guo, Selden & Ren |
Cretaceous |
Burmese amber |
A spider belonging to the family Lagonomegopidae. Genus includes new species S. fax. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Scutuloboridae. The type species is S. admirabilis. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Scutuloboridae. The type species is S. pumilio. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Deinopoidea and the new family Scutuloboridae. The type species is S. spiralembolus. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Archaeidae. The type species is S. aberrans. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the group Araneoidea and the family Zarqaraneidae. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Uloboridae. The type species is S. crux. |
|||
Sp. nov |
Valid |
Lourenço |
Cretaceous |
Burmese amber |
A scorpion belonging to the group Buthoidea and the family Palaeoburmesebuthidae. |
|||
Sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A spider belonging to the family Telemidae, possibly a species of Telemofila. |
|||
Gen. et sp. nov |
Valid |
Wunderlich in Wunderlich & Müller |
Cretaceous |
Burmese amber |
A member of the family Tetrablemmidae. The type species is T. penicillus. |
Research
- Revision of the fossil record of whip spiders is published by Haug & Haug (2021).[23]
- Guo, Selden & Ren (2021) describe an adult lagonomegopid female, part of an egg sac and lagonomegopid spiderlings preserved in four pieces of Burmese amber, and interpret this finding as evidence of maternal care in fossil spiders.[24]
- An exuvium of a member of the genus Myrmecarchaea is described from the Eocene Cambay amber (India) by Wood, Singh & Grimaldi (2021), representing the first member of the family Archaeidae from Cambay amber reported to date.[25]
- An indeterminate deutonymph belonging to the family Sejidae is described by Joharchi, Vorontsov and Walter (2021) from Burmese amber, which represents the oldest record of the parasitiform mite clade Mesostigmata.[26]
Crustaceans
New taxa
Malacostracans
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Schweitzer, Hyžný & Feldmann |
A crab belonging to the family Parthenopidae. |
|||||
Gen. et sp. nov |
In press |
Ferratges, Hyžný & Zamora |
A member of Axiidea belonging to the family Anacalliacidae. Genus includes new species A. enigma. |
|||||
Gen. et sp. nov |
In press |
Ferratges, Hyžný & Zamora |
Early Cretaceous (Aptian) |
Forcall Formation |
A member of Axiidea. Genus includes new species A. longimanus. |
|||
Gen. et sp. nov |
Valid |
Sánchez-García et al. |
Spanish amber |
A member of Isopoda belonging to the family Trichoniscidae. The type species is A. resinicola. |
||||
Gen. et sp. nov |
Valid |
Van Bakel et al. |
A crab belonging to the superfamily Homolodromioidea and the family Prosopidae . The type species is B. piardi.
|
|||||
Gen. et sp. nov |
Valid |
Winkler |
Altmühltal Formation
|
A member of Caridea, possibly belonging to the family Palaemonidae. The type species is B. haereri. |
||||
Gen. et sp. nov |
In press |
Nyborg et al. |
Early to middle Miocene |
A crab belonging to the group Cyclodorippoida and the family Cyclodorippidae. The type species is B. bretoni. |
||||
Gen. et sp. nov |
In press |
De Angeli & Garassino |
A crab belonging to the family Epialtidae. The type species is B. bretoni. |
|||||
Sp. nov |
Valid |
Winkler |
Late Jurassic (Tithonian) |
Altmühltal Formation |
A member of Caridea. |
|||
Sp. nov |
Valid |
Wallaard et al. |
Late Jurassic (Tithonian) |
A crab belonging to the group Homolodromioidea and the family Bucculentidae .
|
||||
Bucculentum horstkuscheli[35] |
Sp. nov |
Valid |
Krzemińska et al. |
Late Jurassic (Kimmeridgian) |
A crab belonging to the group Homolodromioidea and the family Bucculentidae. |
|||
Bucculentum plettenbergense[35] |
Sp. nov |
Valid |
Krzemińska et al. |
Late Jurassic (Oxfordian) |
A crab belonging to the group Homolodromioidea and the family Bucculentidae. |
|||
Sp. nov |
Valid |
De Angeli & Alberti |
Eocene (Bartonian-Priabonian) |
A crab, a species of Carpilius. |
||||
Sp. nov |
In press |
Bruce et al. |
A member of Isopoda, a species of Cirolana. |
|||||
Sp. nov |
In press |
Bruce et al. |
Early Cretaceous (Aptian) |
Sierra Madre Formation |
A member of Isopoda, a species of Cirolana. |
|||
Sp. nov |
In press |
Bruce et al. |
Early Cretaceous (Aptian) |
Sierra Madre Formation |
A member of Isopoda, a species of Cirolana. |
|||
Gen. et sp. nov |
Valid |
Beschin, Busulini & Tessier |
Eocene |
A crab belonging to the family Mathildellidae. The type species is C. spinosa. |
||||
Sp. nov |
Valid |
Schweitzer, Hyžný & Feldmann |
A crab belonging to the family Inachoididae. |
|||||
Gen. et sp. nov |
In press |
Ferratges, Hyžný & Zamora |
Early Cretaceous (Aptian) |
Forcall Formation |
A hermit crab. Genus includes new species C. josaensis. |
|||
Gen. et sp. nov |
Valid |
Luque in Luque et al. |
Late Cretaceous (Cenomanian) |
Burmese amber |
A crab belonging to the group Eubrachyura and to the new family Cretapsaridae. The type species is C. athanata. |
|||
Gen. et sp. nov |
Valid |
Beschin, Busulini & Tessier |
Eocene |
A member of the family Munidopsidae. The type species is C. striata. |
||||
Sp. nov |
In press |
Ferratges, Hyžný & Zamora |
Early Cretaceous (Aptian) |
Forcall Formation |
A member of Axiidea belonging to the family Callianideidae. |
|||
Gen. et sp. nov |
Valid |
Schädel et al. |
Late Cretaceous (Cenomanian) |
Burmese amber |
A member of Isopoda belonging to the group Epicaridea. The type species is C. nidis. |
|||
Sp. nov |
Valid |
Fraaije, Van Bakel & Jagt |
A hermit crab belonging to the family Schobertellidae. |
|||||
Gen. et sp. nov |
Disputed |
Schädel, Hyžný & Haug |
A member of Isopoda belonging to the group Cymothoida. The type species is E. madelineae. Bruce & Rodcharoen (2023) considered Electrolana to be a junior synonym of the genus Cirolana, though the authors maintained E. madelineae as a distinct species within the latter genus.[43] |
|||||
Gen. et comb. nov |
Valid |
O'Flynn, Audo & Kawai |
A member of the family Astacidae; a new genus for "Astacus" edwardsi Van Straelen (1928). |
|||||
Sp. nov |
Valid |
Ferratges, Artal & Zamora |
A hermit crab belonging to the family Calcinidae. |
|||||
Sp. nov |
In press |
Garassino, Feldmann & Schweitzer |
A crab belonging to the family Epialtidae. |
|||||
Gen. et sp. nov |
Valid |
Sánchez-García et al. |
Early Cretaceous (Albian) |
Spanish amber |
A member of Isopoda belonging to the family Ligiidae. The type species is E. tarraconensis. |
|||
Sp. nov |
Valid |
De Angeli & Garassino |
Late Eocene |
A crab, a species of Ethusa. |
||||
Sp. nov |
Valid |
Wei et al. |
Late Neogene |
A member of Amphipoda. |
||||
Sp. nov |
Valid |
Becker, Fraaije & Mulder |
A member of the family Glypheidae .
|
|||||
Sp. nov |
Valid |
Winkler |
Late Jurassic (Tithonian) |
Altmühltal Formation |
A member of Caridea. |
|||
Gen. et sp. nov |
Valid |
Sánchez-García et al. |
Early Cretaceous (Albian) |
Spanish amber |
A member of Isopoda belonging to the group Crinocheta, possibly a member of the family Detonidae. The type species is H. helenae. |
|||
Sp. nov |
Valid |
Garassino, Pasini & Nyborg in Garassino et al. |
A species of Homarus. |
|||||
Sp. nov |
In press |
Feldmann & Schweitzer |
Late Cretaceous (Campanian) |
A lobster. |
||||
Gen. et sp. nov |
Valid |
Winkler et al. |
Late Jurassic (Tithonian) |
A member of Stenopodidea. The type species is J. frattigianii. |
||||
Sp. nov |
Valid |
Feldmann et al. |
A species of Linuparus. |
|||||
Sp. nov |
Valid |
Garassino, Pasini & Nyborg in Garassino et al. |
Early Cretaceous (Albian) |
A member of the family Mecochiridae .
|
||||
Sp. nov |
Valid |
Gašparič et al. |
Early Miocene |
A species of Metanephrops. Announced in 2021; validated in 2023.[55] |
||||
Sp. nov |
In press |
Ferratges, Hyžný & Zamora |
Early Cretaceous (Aptian) |
Forcall Formation |
A member of Axiidea. |
|||
Sp. nov |
Valid |
Charbonnier et al. |
A member of the family Mecochiridae .
|
|||||
Gen. et sp. nov |
Valid |
Garassino, Pasini & Nazarkin |
A member of the family Crangonidae. Genus includes new species M. tartaricus. |
|||||
Sp. nov |
In press |
Stinnesbeck et al. |
Late Cretaceous (Turonian) |
A member of Isopoda belonging to the family Cymothoidae. |
||||
Gen. et comb. nov |
Valid |
Beschin, Busulini & Tessier |
Eocene |
Szépvölgy Formation |
A member of the family Munididae. The type species is "Protomunida" pentacantha Müller & Collins (1991); genus also includes "Eumunida" veronensis Beschin, Busulini & Tessier in Beschin et al. (2019). |
|||
Sp. nov |
Valid |
Poschmann |
A member of Phyllocarida. |
|||||
Gen. et sp. nov |
In press |
De Angeli & Garassino |
Eocene (Priabonian) |
A crab belonging to the family Dairidae . The type species is O. bericus.
|
||||
Sp. nov |
Valid |
Garassino, Pasini & Nyborg in Garassino et al. |
Early Cretaceous (Albian) |
A member of the family Erymidae. |
||||
Sp. nov |
Valid |
Lima, Aguilera & Tavares |
Late Miocene |
A spider crab belonging to the family Inachoididae. |
||||
Gen. et sp. nov |
Valid |
Ferratges, Artal & Zamora |
Eocene (Ypresian) |
A hermit crab belonging to the family Diogenidae. The type species is P. robustus. |
||||
Sp. nov |
Valid |
Ferratges, Domínguez & Ossó |
Eocene (Ypresian) |
Roda Formation |
A crab belonging to the family Homolidae. |
|||
Gen. et sp. nov |
Valid |
Beschin et al. |
Late Eocene |
A hermit crab. Genus includes new species P. scaligera. |
||||
Gen. et sp. nov |
Valid |
Van der Wal et al. |
Late Eocene |
An isopod belonging to the family Cymothoidae. The type species is P. dvorakorum. |
||||
Gen. et sp. nov |
Valid |
Schweigert |
A crab belonging to the group Homolodromioidea and the family Prosopidae. The type species is P. thauckei. Announced in 2021 in an online-only journal; validated in 2023.[65] |
|||||
Sp. nov |
In press |
Bruce et al. |
Early Cretaceous (Aptian) |
Sierra Madre Formation |
A member of Isopoda belonging to the family Cirolanidae. |
|||
Gen. et sp. nov |
Valid |
Pazinato et al. |
A member of Isopoda. The type species is P. odysseus. |
|||||
Sp. nov |
In press |
Andrada et al. |
A member of the family Axiidae. |
|||||
Sp. nov |
In press |
Nyborg et al. |
Late Cretaceous (Coniacian) |
A crab belonging to the group Raninoida and the family Paranecrocarcinidae. |
||||
Sp. nov |
Valid |
Devillez & Charbonnier |
A member of Erymoidea .
|
|||||
Gen. et sp. nov |
Valid |
Haug & Haug |
Altmühltal Group (Eichstätt Subformation) |
A mantis shrimp. The type species is T. laurae. |
||||
Gen. et sp. nov |
Valid |
Beschin, Busulini & Tessier |
Eocene |
A crab belonging to the superfamily Trapezioidea; the type genus of the new family Vicetitrapeziidae. The type species is V. exaltissimo. |
||||
Gen. et sp. nov |
Valid |
Lima, Aguilera & Tavares |
Early-middle Miocene |
A spider crab belonging to the family Inachoididae. Genus includes new species W. santanai. |
||||
Sp. nov |
Valid |
Schweitzer, Hyžný & Feldmann |
A species of Xanthodius. |
|||||
Gen. et sp. nov |
Valid |
Xing et al. |
Late Cretaceous (Cenomanian) |
Burmese amber |
A member of the family Penaeidae. Genus includes new species X. electrinus. |
Ostracods
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Vázquez García et al. |
Cretaceous (Albian–Cenomanian) |
|||||
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
A member of Palaeocopida belonging to the family Kirkbyidae. |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Roadian) |
Cutoff Formation |
A member of the family Bairdiidae. |
|||
Sp. nov |
Valid |
Melnikova et al. |
Syadai Formation |
|||||
Nom. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
In press |
Wang et al. |
Middle Eocene to Oligocene |
A species of Candona. |
||||
Sp. nov |
In press |
Kshetrimayum et al. |
A species of Candona. |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
||||||
Sp. nov |
Valid |
Forel in Tarnac et al. |
Permian (Capitanian) |
A member of the family Bairdiidae. |
||||
Ceratobairdia pratti[73] |
Sp. nov |
Valid |
Forel in Tarnac et al. |
Permian (Roadian) |
Cutoff Formation |
A member of the family Bairdiidae. |
||
Ceratobairdia sexagintaduella[73] |
Sp. nov |
Valid |
Forel in Tarnac et al. |
Permian (Roadian) |
Cutoff Formation |
A member of the family Bairdiidae. |
||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
In press |
Choi et al. |
Early Cretaceous (Albian) |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Wannenmacher et al. |
A member of the family Cytheruridae. |
|||||
Sp. nov |
In press |
Vázquez García et al. |
Cretaceous (Albian–Cenomanian) |
Riachuelo Formation |
||||
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Capitanian) |
Bell Canyon Formation |
A member of Podocopida belonging to the family Microcheilinellidae. |
|||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Wannenmacher et al. |
Middle Jurassic (Aalenian) |
Achdorf Formation |
A member of the family Cytheruridae. |
|||
Sp. nov |
In press |
Maia et al. |
Late Pleistocene |
|||||
Sp. nov |
Valid |
Wannenmacher et al. |
Middle Jurassic (Aalenian) |
A member of the family Cytheruridae. |
||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Melnikova et al. |
Silurian |
Syadai Formation |
||||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
Bonaparte Basin |
||||
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Roadian) |
Cutoff Formation |
A member of Palaeocopida belonging to the family Geisinidae. |
|||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
|||||
Sp. nov |
In press |
Kshetrimayum et al. |
Late Cretaceous (Maastrichtian) |
|||||
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Roadian) |
Cutoff Formation |
A member of Podocopa belonging to the family Healdiidae. |
|||
Healdia mckittrickensis[73] |
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Roadian to Capitanian) |
Bell Canyon Formation |
A member of Podocopa belonging to the family Healdiidae. |
||
Healdia veeversi[80] |
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
Bonaparte Basin |
|||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
Bonaparte Basin |
||||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
Bonaparte Basin |
||||
Sp. nov |
In press |
Forel in Forel, Poulet-Crovisier & Korat |
Late Permian and Early Triassic |
Daye Formation |
A member of Palaeocopida. |
|||
Hollinella williamsranchensis[73] |
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Roadian to Capitanian) |
Bell Canyon Formation |
A member of Palaeocopida belonging to the family Hollinellidae. |
||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
In press |
Song et al. |
Late Devonian |
|||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
In press |
Vázquez García et al. |
Cretaceous (Albian–Cenomanian) |
Riachuelo Formation |
||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
Bonaparte Basin |
||||
Sp. nov |
In press |
Vázquez García et al. |
Cretaceous (Albian–Cenomanian) |
Riachuelo Formation |
||||
Sp. nov |
Valid |
Crasquin in Tarnac et al. |
Permian (Roadian to Capitanian) |
Cutoff Formation |
A member of Podocopa belonging to the family Paraparchitidae. |
|||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
|||||
Sp. nov |
Valid |
Glinskikh & Tesakova |
||||||
Sp. nov |
Valid |
Forel in Forel & Grădinaru |
A member of the family Polycopidae. |
|||||
Sp. nov |
Valid |
Forel in Forel & Grădinaru |
Triassic |
A member of the family Polycopidae. |
||||
Sp. nov |
Valid |
Forel in Forel & Grădinaru |
Triassic |
A member of the family Polycopidae. |
||||
Sp. nov |
Valid |
Forel in Forel & Grădinaru |
Triassic |
A member of the family Polycopidae. |
||||
Sp. nov |
Valid |
Forel in Forel & Grădinaru |
Triassic |
A member of the family Polycopidae. |
||||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Gen. et sp. nov |
Valid |
Sciuto in Sciuto & Benkhedda |
A member of the family Trachyleberididae. The type species is P. benkheddai. |
|||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Gen. et sp. nov |
Valid |
Sciuto in Sciuto & Benkhedda |
Miocene (Tortonian) |
A member of the family Trachyleberididae. The type species is Q. tevestaensis. |
||||
Sp. nov |
In press |
Vázquez García et al. |
Cretaceous (Albian–Cenomanian) |
Riachuelo Formation |
||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
Bonaparte Basin |
||||
Sp. nov |
Valid |
Slipper |
Late Cretaceous (Turonian) |
|||||
Gen. et sp. nov |
Valid |
Melnikova et al. |
Silurian |
Syadai Formation |
The type species is S. tumidosa. |
|||
Sp. nov |
Valid |
Sciuto & Reitano |
Early Pliocene |
|||||
Sp. nov |
Valid |
Jones, Kelman & Laurie |
Carboniferous (Mississippian) |
|||||
Sp. nov |
Valid |
Sciuto, Temani & Ammar |
Miocene (Messinian) |
Other crustaceans
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Li, Teng & Wimbledon |
Late Jurassic/Early Cretaceous |
A clam shrimp. |
||||
Sp. nov |
In press |
Collareta & Newman in Collareta et al. |
Chake-Chake beds |
A barnacle, a species of Chelonibia. |
||||
Sp. nov |
In press |
Gale |
A barnacle. |
|||||
Sp. nov |
In press |
Gale |
A barnacle. |
|||||
Sp. nov |
Valid |
Liao & Shen |
Early Devonian |
A clam shrimp. |
||||
Sp. nov |
Valid |
Gale |
Plio-Pleistocene |
A barnacle. |
||||
Gen. et sp. nov |
Valid |
Tang, Mychko, Feldmann & Schweitzer in Tang et al. |
A member of Cyclida. Genus includes new species M. terengganuensis. Announced in 2021; validated in 2023.[95] |
|||||
Sp. nov |
Valid |
Liao & Shen |
Early Devonian |
A clam shrimp. |
||||
Sp. nov |
Valid |
Gale |
Plio-Pleistocene |
A barnacle. |
Research
- A study on the anatomy and phylogenetic relationships of the species "Penaeus" natator from the Santonian of Lebanon is published by Audo, Winkler & Charbonnier (2021), who interpret this species as a relative of Pseudodrobna kenngotti from the Late Jurassic of Germany, and transfer it to the genus Pseudodrobna[96]
- A study on the anatomy and morphological variation in Beurlenia araripensis, based on data from fossil samples from the Crato Formation (Brazil), is published by Barros et al. (2021).[97]
- A study on the anatomy of the eyes of Callichimaera perplexa, and on their growth during the ontogeny of this crab, is published by Jenkins, Briggs & Luque (2021).[98]
- A study on the anatomy and phylogenetic relationships of Oxyuropoda is published by Robin et al. (2021), who interpret this arthropod as the oldest known member of the crown group of Peracarida.[99]
- Fossil evidence of early colonization of estuarine settings by ostracods is reported from the Silurian Si Ka Formation (Vietnam) by McGairy et al. (2021)[100]
- A study on evolutionary trends in
Insects
Radiodonts
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Pates et al. |
||||||
Gen. et comb. nov |
Valid |
Wu et al. |
A tamisiocaridid radiodont. |
|||||
Gen. et sp. et comb. nov |
In press |
Wu et al. |
Cambrian |
An anomalocaridid radiodont. |
||||
Sp. nov |
Valid |
Jiao et al. |
Wulongqing Formation |
|||||
Gen. et sp. nov |
Valid |
Caron & Moysiuk |
Cambrian (Wuliuan) |
A hurdiid radiodont. |
Research
- A study on the morphologies of frontal appendages and probable modes of feeding of radiodonts from the Burgess Shale is published by De Vivo, Lautenschlager & Vinther (2021).[107]
Trilobites
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Feist & Klapper |
A member of the family Phacopidae. |
|||||
Gen. et 2 sp. et comb. nov |
In press |
Feist & Klapper |
Late Devonian |
A member of the family Phacopidae. The type species is A. medius; genus also includes new species A. tchrafinensis, as well as "Phacops" erfoudensis Richter & Richter (1943). |
||||
Sp. nov |
Valid |
Flick & Shiino |
Permian (probably Wordian) |
Hoso-o Formation |
||||
Gen. et comb. nov |
Valid |
Holloway |
A member of Lichida belonging to the family Lichidae; a new genus for "Amphilichas" shallopensis Twenhofel (1928). |
|||||
Sp. nov |
Valid |
Holloway |
Silurian |
A member of Lichida belonging to the family Lichidae. |
||||
Sp. nov |
In press |
Wernette et al. |
Cambrian (Furongian) |
Myet-Ye Formation |
||||
Sp. nov |
Valid |
Flick |
Devonian |
Greifenstein Limestone |
||||
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Bollandiinae. |
||||
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Brachymetopidae belonging to the subfamily Brachymetopinae. |
||||
Gen. et sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. Genus includes new species C. celadensis. |
||||
Sp. nov |
Valid |
Flick & Shiino |
Permian (probably Wordian) |
Hoso-o Formation |
||||
Sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
A member of the family Phacopidae. |
||||
Chlupacops clapassousensis[108] |
Sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
A member of the family Phacopidae. |
|||
Chlupacops migrans[108] |
Sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
Serre Formation |
A member of the family Phacopidae. |
||
Chlupacops narbonnensis[108] |
Sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
A member of the family Phacopidae. |
|||
Chlupacops rectannulatus[108] |
Sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
Serre Formation |
A member of the family Phacopidae. |
||
Sp. nov |
Valid |
Flick |
Devonian |
Günterod Limestone |
||||
Gen. et sp. nov |
Valid |
Gandl |
Carboniferous |
Possibly a member of the family Phillipsiidae. Genus includes new species C. matallanensis. |
||||
Gen. et 3 sp. et comb. nov |
In press |
Feist & Klapper |
Late Devonian |
Serre Formation |
A member of the family Phacopidae. The type species is C. gallica; genus also includes new species C. postera and C. expansa, as well as "Phacops" zofiae Chlupáč (1993). |
|||
Sp. nov |
Valid |
Nielsen & Andersen |
Cambrian (Furongian) |
A member of the family Olenidae. |
||||
|
Sp. nov |
Valid |
Adrain & Pérez-Peris |
A member of Acanthoparyphinae. |
||||
Sp. nov |
Valid |
Holloway |
Silurian |
A member of the family Odontopleuridae. |
||||
Sp. nov |
Valid |
Holloway |
Silurian |
A member of Lichida belonging to the family Lichidae. |
||||
Dicranopeltis kallaion[110] |
Sp. nov |
Valid |
Holloway |
Silurian |
A member of Lichida belonging to the family Lichidae. |
|||
Sp. nov |
Valid |
Flick & Shiino |
Permian (probably Wordian) |
Hoso-o Formation |
||||
Ditomopyge (Carniphillipsia) cantamudensis[113] |
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. |
|||
Ditomopyge (Carniphillipsia) castelleriae[113] |
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. |
|||
Sp. nov |
Valid |
Van Viersen |
||||||
Dohmiella pooka[117] |
Sp. nov |
Valid |
Van Viersen |
Devonian (Eifelian) |
||||
Sp. nov |
In press |
Geyer, Landing & Żylińska |
Cambrian (Miaolingian) |
Jbel Wawrmast Formation |
A member of the family Paradoxididae. |
|||
Eccaparadoxides? hestia[118] |
Sp. nov |
In press |
Geyer, Landing & Żylińska |
Cambrian (Miaolingian) |
Jbel Wawrmast Formation |
A member of the family Paradoxididae. |
||
Eccaparadoxides zelus[118] |
Sp. nov |
In press |
Geyer, Landing & Żylińska |
Cambrian (Miaolingian) |
Delitzsch Formation |
A member of the family Paradoxididae. |
||
Sp. nov |
Valid |
Peel |
A member of the family Alokistocaridae. |
|||||
Sp. nov |
Valid |
Smith & Laurie |
Ordovician (Darriwilian) |
|||||
Sp. nov |
Valid |
Peel |
Cambrian (Guzhangian) |
Blue Cliffs Formation |
A member of the family Bolaspididae. |
|||
Sp. nov |
Valid |
Peel |
Telt Bugt Formation |
A member of the family Zacanthoididae. |
||||
Sp. nov |
Valid |
Van Viersen |
Devonian (Eifelian) |
|||||
Gen. et sp. nov |
Valid |
Smith & Laurie |
Ordovician (Darriwilian) |
Genus includes new species G. ritchiei. |
||||
Gen. et 2 sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
A member of the family Phacopidae. The type species is G. konradbartzschi; genus also includes new species G. consimilis, and possibly also "Phacops" liopyga Richter (1863). |
||||
Sp. nov |
Valid |
Adrain & Karim |
Possibly a member of the family Dimeropygidae. |
|||||
Gonioteloides pankowskii[122] |
Sp. nov |
Valid |
Adrain & Karim |
Ordovician (Tremadocian) |
Possibly a member of the family Dimeropygidae. |
|||
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Griffithidinae. |
||||
Gen. et sp. nov |
Valid |
Adrain & Pérez-Peris |
Ordovician (Darriwilian) |
Table Cove Formation |
A member of Pilekiinae. The type species is H. plurima. |
|||
Sp. nov |
Valid |
Wei et al. |
Ordovician (Darriwilian) |
|||||
Gen. et sp. nov |
Valid |
Smith & Laurie |
Ordovician (Darriwilian) |
Genus includes new species I. schoonorum. |
||||
Sp. nov |
In press |
Hughes et al. |
Cambrian (Miaolingian) |
|||||
Sp. nov |
Valid |
Adrain & Pérez-Peris |
Ordovician (Darriwilian) |
Table Cove Formation |
A member of Acanthoparyphinae. |
|||
Sp. nov |
Valid |
Alberti & Müller |
Devonian (Emsian) |
Rupbach Shales |
A member of the family Odontopleuridae. |
|||
Sp. nov |
Valid |
Flick |
Devonian |
Günterod Limestone |
||||
Koneprusites cervisiaemontis[126] |
Sp. nov |
Valid |
Flick |
Devonian |
Günterod Limestone |
|||
Sp. nov |
Valid |
Flick & Flick |
Devonian (Emsian) |
Ballersbach Limestone |
A member of the family Odontopleuridae. |
|||
Sp. nov |
Valid |
Smith & Laurie |
Ordovician (Darriwilian) |
|||||
Gen. et sp. nov |
Valid |
Adrain & Pérez-Peris |
Ordovician (Darriwilian) |
Table Cove Formation |
A member of Deiphoninae. The type species is M. becki. |
|||
Sp. nov |
Valid |
Wei et al. |
||||||
Sp. nov |
Valid |
Holloway |
Silurian |
A member of the family Odontopleuridae. |
||||
Sp. nov |
Valid |
Flick |
Devonian |
Greifenstein Limestone |
A member of the family Phacopidae. |
|||
Sp. nov |
Valid |
Müller, Hahn & Paul |
A member of the family Brachymetopidae. |
|||||
Namuropyge brooksi[129] |
Sp. nov |
Valid |
Müller, Hahn & Paul |
Carboniferous (Mississippian) |
A member of the family Brachymetopidae. |
|||
Namuropyge ulrichi[129] |
Sp. nov |
Valid |
Müller, Hahn & Paul |
Carboniferous (Viséan) |
A member of the family Brachymetopidae. |
|||
Sp. nov |
Valid |
Flick & Shiino |
Permian (probably Wordian) |
Hoso-o Formation |
||||
Gen. et sp. nov |
Valid |
Adrain & Pérez-Peris |
Ordovician (Darriwilian) |
Table Cove Formation |
Possibly a member of Sphaerexochinae. The type species is N. karimae. |
|||
Sp. nov |
Valid |
Wei et al. |
Ordovician (Darriwilian) |
|||||
Sp. nov |
Valid |
Smith & Laurie |
Ordovician (Darriwilian) |
|||||
Nyterops fugax[130] | Sp. nov | Valid | Van Viersen & Koppka | Devonian (Givetian) | Hanonet Formation | Belgium | A member of the family Phacopidae. | |
Nyterops gerolsteinensis[130] | Sp. nov | Valid | Van Viersen & Koppka | Devonian (Eifelian?) | Germany | A member of the family Phacopidae. | ||
Gen. et 2 sp. nov |
In press |
Feist & Klapper |
Devonian (Frasnian) |
A member of the family Phacopidae. The type species is O. hexagonalis; genus also includes new species O. postinflexa, and possibly also "Trimerocephaloides" linguiformis Feist et al. (2009). |
||||
Sp. nov |
Valid |
Holloway |
Silurian |
A member of the family Scutelluidae .
|
||||
Octobronteus? ormistoni[131] |
Sp. nov |
Valid |
Holloway |
Silurian |
A member of the family Scutelluidae. |
|||
Octobronteus? petasus[131] |
Sp. nov |
Valid |
Holloway |
Silurian |
A member of the family Scutelluidae. |
|||
Sp. nov |
Valid |
Wei et al. |
Ordovician (Darriwilian) |
|||||
Sp. nov |
Valid |
Wei et al. |
Ordovician (Darriwilian) |
|||||
Pedinopariops apsconditus[130] | Sp. nov | Valid | Van Viersen & Koppka | Devonian (Eifelian) | Jemelle Formation | Belgium | A member of the family Phacopidae. | |
Sp. nov |
Valid |
Holloway |
Silurian (Aeronian) |
A member of the family Scutelluidae. |
||||
Gen. et sp. nov |
Valid |
Basse & Müller |
Early Devonian |
A member of the subfamily Proetinae. Genus includes new species P. feisti. |
||||
Gen. et sp. nov |
Disputed |
Alberti |
Devonian (Emsian) |
Rupbach Shale |
A member of the family Odontopleuridae. The type species is R. paedomorpha. Van Viersen (2024) considered R. paedomorpha to be a junior synonym of Leonaspis kleini.[134] |
|||
Gen. et sp. nov |
Valid |
Wei et al. |
Ordovician (Darriwilian) |
Genus includes new species S. mirabilis. |
||||
Sp. nov |
Valid |
Sun, Bentley & Laurie |
Cambrian (Guzhangian) |
Warburton Basin
|
||||
Sp. nov |
Valid |
Van Viersen & Lerouge |
Devonian (Emsian) |
A member of the family Proetidae. |
||||
Sp. nov |
Valid |
Holloway |
Silurian |
A member of Lichida belonging to the family Lichidae. |
||||
Sp. nov |
Valid |
Flick |
Devonian |
Ballersbach Limestone |
||||
Sp. nov |
Valid |
Basse & Müller |
Early Devonian |
A member of the subfamily Proetinae. |
||||
Gen. et sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Bollandiinae. Genus includes new species V. rara. |
||||
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. |
||||
Waideggula (Waideggula) cantabrica tabardina[113] |
Ssp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. |
|||
Waideggula (Waideggula) pedregalensis[113] |
Sp. nov |
Valid |
Gandl |
Carboniferous |
A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. |
|||
Sp. nov |
Valid |
Flick & Shiino |
Permian (probably Wordian) |
Hoso-o Formation |
||||
Gen. et sp. nov |
Valid |
Wei et al. |
Ordovician (Darriwilian) |
Genus includes new species Y. punctata. |
Research
- A study on middle–late Cambrian trilobite diversity patterns in South China is published by Zhang et al. (2021).[137]
- Sun, Zeng & Zhao (2021) describe digestive structures of representatives of five trilobite genera from the Cambrian Mantou Formation and Zhangxia Formation (Liaoning, China).[138]
- Hou, Hughes & Hopkins (2021) report structural details of the upper limb branch of Triarthrus eatoni and Olenoides serratus, and interpret their findings as indicating that the upper limb branch of trilobites served a respiratory function.[139]
- A study on the morphology of Redlichia rex and Olenoides serratus, aiming to determine whether these trilobites were adapted for durophagy, is published by Bicknell et al. (2021).[140]
- A study exploring the existence and the nature of growth gradients along the main body axis of Oryctocarella duyunensis is published by Dai et al (2021), who interpret O. duyunensis as the first trilobite with documented determinate growth.[141]
- Description of all meraspid stages of Oryctocarella duyunensis, based on data from specimens from the Cambrian Balang Formation (Hunan, South China), is published by Dai et al. (2021).[142]
- A study on the ontogenic moulting sequence of Arthricocephalites xinzhaiheensis, based on data from specimens from the Balang Formation, is published by Wang et al. (2021).[143]
- A study on post-embryonic axial growth in Estaingia bilobata is published by Holmes, Paterson & García-Bellido (2021).[144]
- A study on the functional performance of Placoparia cambriensis in water is published by Esteve et al. (2021), who interpret their findings as indicating that this trilobite was not able to swim, but it was capable of hopping locomotion, and it might represent a transitional form between benthic animals without swimming skills and animals able to swim near the benthos.[145]
- A study on the phylogenetic relationships within Ordovician–Silurian extinction events on this group, is published by Beech & Lamsdell (2021).[146]
- The first known Silurian trilobite specimen preserved with soft parts reported to date (a specimen of Herefordshire Lagerstätte (United Kingdom) by Siveter et al. (2021).[147]
- A study on the long-term evolutionary history of Devonian trilobites in North Africa is published by Bault et al. (2021).[148]
- A study on the hydrodynamics of queuing behaviour in Trimerocephalus chopini is published by Song et al. (2021).[149]
- Evidence from trace and body fossils indicative of the presence of trilobites in brackish-water settings is presented by Mángano et al. (2021).[150]
- A study on the chemical changes in the exoskeleton of trilobites induced by diagenesis, based on data from pygidia of Athabaskia anax from the Miaolingian of San Isidro (Argentina), is published by D'Angelo et al. (2021), is published by D'Angelo et al. (2021), who argue that some morphological characteristics of the trilobite pygidia are in fact results of chemical and structural changes taking place during fossilization, and evaluate possible systematic implications of the chemical data, advising caution when using morphological characteristics of the exoskeletons to establish new taxa.[151]
- The study on the internal structures of eyes of trilobites belonging to the genera Asaphus and Archegonus published by Scholtz, Staude & Dunlop (2019)[152] is criticized by Schoenemann & Clarkson (2021).[153][154]
- A study on the biomechanics of the trilobite cephalon is published by Esteve et al. (2021), who interpret their findings as indicating that in the sutured trilobites the cephalon was able to withstand greater stresses than in their non‐sutured counterparts, and argue that the ability to withstand greater burrowing loads enabled trilobites to successfully invade bioturbated and more consolidated sediments during the Cambrian substrate revolution.[155]
- A study on the compound eyes of trilobites belonging to the group Phacopina is published by Schoenemann et al. (2021), who interpret their findings as indicating that these trilobites had hyper-compound eyes hiding an individual compound eye below each of the big lenses, resulting in each of the compound eyes comprising tens or hundreds of small compound eye systems.[156]
- A study on the systematics of Devonian trochurine trilobites is published by Van Viersen (2021).[157]
Other arthropods
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Bicknell & Shcherbakov |
A member of Xiphosura belonging to the family Austrolimulidae. The type species is A. superspinosus. |
|||||
Gen. et sp. nov |
In press |
Jiao et al. |
A non-trilobite artiopod. The type species is B. longicaudata. |
|||||
Gen. et sp. nov |
Valid |
Van Roy et al. |
A member of Thylacocephala. Genus includes new species B. libori. |
|||||
Gen. et sp. nov |
In press |
Ma et al. |
A bivalved arthropod related to Tuzoia. Genus includes new species D. anima. |
|||||
Gen. et 2 sp. nov |
Valid |
Moritz & Wesener |
Cretaceous |
Burmese amber |
A Cambalidea . The type species is E. ornata; genus also includes E. cretacea.
|
|||
Gen. et sp. nov |
Valid |
Laville, Haug & Haug |
Carboniferous (middle Pennsylvanian) |
A member of Thylacocephala. The type species is E. keithflinti. |
||||
Gen. et sp. nov |
Valid |
Bicknell, Hecker & Heyng |
A member of Xiphosura belonging to the family Austrolimulidae. Genus includes new species F. pochankei. |
|||||
Sp. nov |
In press |
Wang et al. |
Early Devonian |
|||||
Gen. et 2 sp. nov |
Valid |
Moritz & Wesener |
Cretaceous |
Burmese amber |
A millipede belonging to the order Spirostreptida and the suborder Cambalidea. The type species is K. muelleri; genus also includes K. distorta. |
|||
Gen. et sp. nov |
Jiao et al. |
A stem-group euarthropod of uncertain phylogenetic placement, with possible close affinities with either radiodonts or fuxianhuiids. The type species is L. ferox. |
||||||
Sp. nov |
Valid |
Riquelme & Hernández-Patricio in Riquelme, Hernández-Patricio & Álvarez-Rodríguez |
A millipede belonging to the family Pyrgodesmidae. |
|||||
Gen. et sp. nov |
Valid |
Lamsdell et al. |
A horseshoe crab. Genus includes new species O. latus. |
|||||
Gen. et sp. nov |
Valid |
Izquierdo-López & Caron |
A stem mandibulate bivalved arthropod. Genus includes new species P. apatis. |
|||||
Sp. nov |
Valid |
Jin et al. |
Cambrian |
A stem mandibulate bivalved arthropod. |
||||
Gen. et sp. nov |
Valid |
Van Roy et al. |
Ordovician (Sandbian) |
Letná Formation |
A member of Thylacocephala. Genus includes new species P. irenae. |
|||
Gen. et sp. nov |
Valid |
Wang et al. |
Silurian (Llandovery) |
Xiushan Formation and Fentou Formation |
A member of Eurypterida . Genus includes new species T. xiushanensis.
|
|||
Sp. nov |
In press |
Poschmann & Rozefelds |
Bowen Basin (Baralaba Coal Measures) |
A member of Eurypterida. |
Research
- Lan et al. (2021) report exquisite preservation of bilaterally symmetric brain in Kaili biota (China), and evaluate the implications of these fossils for the knowledge of the evolution of the central nervous system in arthropods.[173]
- A study on carapace shape variation and hydrodynamic performance of members of the genus Isoxys is published by Pates et al. (2021), who argue that members of this genus occupied a variety of distinct niches in Cambrian oceans, and some were adapted for vertical movement in the water column.[174]
- A study on the composition and microstructure of the carapace of Chuandianella ovata is published by Liu, Fu & Zhang (2021), who interpret their findings as indicating that this arthropod reinforced its carapace with phosphatic mineralization.[175]
- A study on the post-embryonic development of Chuandianella ovata is published by Liu, Fu & Zhang (2021).[176]
- Braddy & Dunlop (2021) argue that Parioscorpio venator was a cheloniellid-like arthropod with large raptorial appendages.[177]
- New information on the anatomy of the head of Fuxianhuia is presented by Aria, Zhao & Zhu (2021), who interpret fuxianhuiids as mandibulates.[178]
- Partial remains of a member of the genus Arthropleura, representing one of the largest arthropod fossils reported to date and providing new information on the exoskeleton of Arthropleura, are described from the Carboniferous (Serpukhovian) Stainmore Formation (Northumberland, England, United Kingdom) by Davies et al. (2021), who also evaluate the implications of this finding for the knowledge of arthropleurid habitat and factors that enabled the evolution of large body size in arthropleurids.[179]
- Revision of the morphological diversity, relationships and taxonomy of Early Triassic thylacocephalans is published by Laville et al. (2021).[180]
- Description of new fossil material of Mayrocaris bucculata from the Solnhofen Limestone, providing new information on the anatomy of this thylacocephalan, is published by Laville et al. (2021), who evaluate the implications of these fossils for the knowledge of the body organization and phylogenetic affinities of thylacocephalans.[181]
- A study on the morphology and possible intraspecific variability of Sinoburius lunaris is published by Schmidt et al. (2021).[182]
- A fossil larva lacking segmentation of the carapace, closely resembling the trilobite protaspis, is described from the Ordovician (Darriwilian) of central Siberia by Dzik (2021), found associated with other skeletal elements of the angarocaridid Girardevia;[183] however, Lerosey-Aubril & Laibl (2021) subsequently interpret this specimen as actually belonging to the trilobite genus Isotelus or a related taxon, and conclude that protaspid larvae represent a developmental trait unique to trilobites.[184][185]
- A study on the possible relationships between eurypterid morphology, the ease with which members of this group experienced ecdysis, and longevity of eurypterid species is published by Brandt (2021).[186]
- A geometric morphometric analysis of data from eurypterine eurypterid specimens is presented by Bicknell & Amati (2021).[187]
- Redescription of Leiopterella tetliei is published by Braddy, Dunlop & Bonsor (2021).[188]
- Bicknell, Melzer & Schmidt (2021) reconstruct prosomal appendages of Eurypterus tetragonophthalmus and Pentecopterus decorahensis, model the flexure and extension of these appendages.[189]
- A study on the range of motion of prosomal appendages in Megalograptus ohioensis and Mixopterus kiaeri, and on its implications for the knowledge of the likely foraging strategies of these eurypterids, is published by Schmidt et al. (2021).[190]
- Bicknell et al. (2021) report the discovery of exceptionally preserved central nervous system in a specimen of Euproops danae from the Carboniferous Mazon Creek fossil beds (Illinois, United States).[191]
- A specimen of Euproops danae preserving anatomical details of the prosomal musculature is described from the Carboniferous Lawrence Formation (Kansas, United States) by Bicknell et al. (2021).[192]
- Redescription and a study on the phylogenetic relationships of Prolimulus woodwardi is published by Lustri, Laibl & Bicknell (2021).[193]
- Revision of Sloveniolimulus rudkini, based on data from new fossil material from the Anisian Strelovec Formation (Slovenia), is published by Bicknell et al. (2021).[194]
- A study on the anatomy and phylogenetic relationships of Parioscorpio venator is published by Anderson et al. (2021).[195]
General research
- A study on the evolution of the arthropod labrum is published by Budd (2021), who reevaluates the morphology of the Cambrian stem-euarthropod Parapeytoia and evaluates its implications for the knowledge of the origin of the labrum.[196]
- Liu et al. (2021) report the discovery a previously undetected exite at the base of most appendages of Leanchoilia illecebrosa, as well as morphologically similar (and likely homologous) exite in the same position in Naraoia spinosa and Retifacies abnormalis, and evaluate the implications of this discovery for the knowledge of the origin of exites in arthropod phylogeny.[197]
References
- ^ S2CID 235600702.
- ^ OCLC 1249433671.
- ^ S2CID 234771916.
- ^ .
- ^ Lourenço, W. R.; Velten, J. (2021). "Early Cretaceous Burmite fossils of the genus Chaerilobuthus Lourenço & Beigel, 2011 (Scorpiones: Chaerilobuthidae) and description of a particular new species". Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire. 9 (30): 1–5.
- ^ Lourenço, W. R.; Velten, J. (2021). "One more new genus and species of scorpion from Early Cretaceous Burmese amber (Scorpiones: Protoischnuridae)". Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire. 9 (14): 1–5.
- S2CID 234364801.
- ^ S2CID 231729394.
- ^ S2CID 235687701.
- ^ .
- S2CID 244688216.
- PMID 34301989.
- S2CID 244063809.
- ISBN 978-3-931473-22-8.
- ^ S2CID 235688196.
- S2CID 244439704.
- S2CID 245030280.
- ^ a b Selden, P. A. (2021). "New spiders (Araneae: Mesothelae), from the Carboniferous of New Mexico and England, and a review of Paleozoic Araneae". New Mexico Museum of Natural History and Science Bulletin. 84: 317–358.
- S2CID 232021004.
- S2CID 233562507.
- S2CID 239104785.
- ^ Lourenço, W. R. (2021). "Further comments on the elements of the family Palaeoburmesebuthidae Lourenço, 2015 with description of a new species of Spinoburmesebuthus Lourenço, 2017 from Early Cretaceous Burmite amber (Scorpiones)". Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire. 9 (17): 1–6.
- S2CID 232434849.
- PMID 34521253.
- PMID 34887693.
- S2CID 239420481.
- ^ S2CID 236264488.
- ^ S2CID 234089134.
- ^ S2CID 236936902.
- doi:10.26879/1153.
- ^ S2CID 234095653.
- S2CID 245098970.
- ^ S2CID 245113149.
- S2CID 243872256.
- ^ S2CID 239137045.
- ^ De Angeli, A.; Alberti, R. (2021). "Carpilius cantellii n. sp. (Decapoda, Brachyura, Carpiloidea) nuovo crostaceo eocenico del territorio vicentino (Italia nordorientale)". Studi Trentini di Scienze Naturali. 101: 53–59.
- ^ S2CID 233598895.
- ^ a b c d Beschin, C.; Busulini, A.; Tessier, G. (2021). "La fauna di crostacei associati a coralli dell'Eocene inferiore dell'Alta Valle del Chiampo (Altissimo - Vicenza - Italia nordorientale)". Lavori – Società Veneziana di Scienze Naturali. 46: 67–128.
- PMID 34669480.
- S2CID 235272269.
- S2CID 233592807.
- S2CID 234105973.
- .
- S2CID 238861251.
- ^ S2CID 245116760.
- S2CID 245116153.
- S2CID 237715372.
- S2CID 233907086.
- S2CID 233956333.
- ^ S2CID 238730142.
- S2CID 245110985.
- S2CID 241658829.
- S2CID 233965804.
- S2CID 245028447.
- S2CID 260528714.
- S2CID 234137706.
- S2CID 238685807.
- S2CID 239627722.
- S2CID 231793893.
- ^ S2CID 243111372.
- S2CID 245099698.
- S2CID 233519062.
- PMID 34966568.
- S2CID 245103649.
- S2CID 259588399.
- .
- S2CID 243402521.
- S2CID 245092898.
- from the original on 2021-01-29. Retrieved 2021-01-28.
- PMID 33959413.
- PMID 36654378.
- ^ S2CID 234215087.
- ^ S2CID 240548510.
- ^ S2CID 239241428.
- ^ S2CID 245546819.
- ^ S2CID 238640282.
- ^ S2CID 231741696.
- S2CID 234073147.
- ^ S2CID 233959714.
- ^ a b c d e f g h i Jones, P. J.; Kelman, A.; Laurie, J. R. (2021). "Mississippian Ostracoda from the Bonaparte and Canning basins, NW Australia: Platycopina and Podocopida, their biostratigraphy, palaeoecology and palaeozoogeographic links". Australasian Palaeontological Memoirs. 54: 1–162.
- .
- ^ S2CID 236284253.
- S2CID 233547784.
- .
- S2CID 236572952.
- S2CID 232070786. Retrieved 2021-01-12.
- ^ S2CID 239643702.
- ^ S2CID 242275243.
- .
- S2CID 234770890.
- ^ S2CID 235526409.
- ^ S2CID 238736745.
- ^ a b Gale, A. S. (2021). "Cirripedes (Crustacea, Thoracica) from the Plio-Pleistocene of Sicily (Italy) and a review of the history of Mediterranean Cainozoic cirripedes". Cainozoic Research. 21 (1): 3–73.
- S2CID 233522378.
- S2CID 260026897.
- S2CID 233328177.
- PMID 33730028.
- S2CID 245113954.
- PMID 34129798.
- PMID 34847752.
- hdl:10400.1/18610.
- PMID 33552709.
- S2CID 235221043.
- S2CID 233565727.
- S2CID 236683029.
- PMID 34527273.
- PMID 34284622.
- ^ S2CID 246358282.
- ^ S2CID 240556158.
- ^ S2CID 237658636.
- S2CID 234832348.
- ^ Flick, U. (2021). "Belenopyge bellerophon n. sp. und Perunaspis? sp. (Trilobita) aus dem Greifenstein-Kalk der Typlokalität (Grenzbereich Emsium/Eifelium, Unter-/Mitteldevon)". Geologisches Jahrbuch Hessen. 140: 35–46.
- ^ ISBN 978-3-510-61421-9.
- ^ a b Flick, U. (2021). "Taxa der Phacopinae HAWLE & CORDA, 1847 (Trilobita) aus dem Grenzbereich Emsium/Eifelium sowie Unter-Eifelium (Unter-/Mitteldevon) der Bicken-Ense-Decke (Rheinisches Schiefergebirge)". Geologisches Jahrbuch Hessen. 140: 73–100.
- S2CID 244144045.
- ^ S2CID 239237412.
- ^ S2CID 233910967.
- ^ S2CID 245014554.
- ^ S2CID 234108940.
- ^ S2CID 237518004.
- S2CID 233352957.
- ^ S2CID 236581079.
- ^ S2CID 245106828.
- S2CID 244254193.
- ^ Alberti, M.; Müller, P. (2021). "Koneprusia dongesi n. sp. (Trilobita; Odontopleuridae) aus dem tiefen Rupbach-Schiefer (mittleres Ober-Emsium; Rhenoherzynikum)". Mainzer geowissenschaftliche Mitteilungen. 49: 67–80.
- ^ a b c Flick, U. (2021). "Die Trilobiten-Fauna im Ballersbach-Kalk (Bicken-Formation) an der Typlokalität (Trilobita; Unter-/Mitteldevon)". Geologisches Jahrbuch Hessen. 140: 5–34.
- ^ Flick, U.; Flick, H. (2021). "Eine neue Art der Gattung Leonaspis R. & E. RICHTER, 1917 (Trilobita) aus dem Ballersbach-Kalk (hohes Unter-/tiefes Mitteldevon) von Ballersbach/Lahn-Dill-Gebiet, Rheinisches Schiefergebirge". Philippia. 18: 109–122.
- S2CID 238686171.
- ^ a b c Müller, P.; Hahn, G.; Paul, T. (2021). "Neue Funde der Gattung Namuropyge aus dem nördlichen Rheinischen Schiefergebirge (Trilobita, Viseum, Deutschland)". Dortmunder Beiträge zur Landeskunde - naturwissenschaftliche Mitteilungen. 50: 131–167.
- ^ a b c van Viersen, A. P.; Koppka, J. (2021). "Type and other species of Phacopidae (Trilobita) from the Devonian of the Ardenno-Rhenish Mountains". Mainzer Geowiss. Mitt. 49: 25–66.
- ^ S2CID 238697672.
- ^ a b Basse, M.; Müller, P. (2021). "The pre-Middle Devonian Proetinae of the German Variscides, with new taxa from the Early Devonian in the Lahn Syncline (Trilobita, Rhenish Massif)" (PDF). Paläontologie, Stratigraphie, Fazies. (26), Freiberger Forschungshefte, C 559: 17–62.
- ^ Alberti, M. (2021). "Der Odontopleurid Rupbachella paedomorpha n. gen., n. sp. aus dem tiefen Rupbach-Schiefer (Trilobita; Ober-Emsium; Rheinisches Schiefergebirge)". Mainzer Naturwissenschaftliches Archiv. 58: 53–67.
- .
- S2CID 239534285.
- S2CID 232133406.
- S2CID 233670096.
- .
- PMID 33789898.
- PMID 33499790.
- hdl:11577/3391127.
- S2CID 233703345.
- PMID 34631322.
- S2CID 245132667.
- PMID 34562458.
- S2CID 238725336.
- .
- S2CID 233852361.
- S2CID 237875240.
- PMID 33529560.
- S2CID 233935342.
- PMID 31175282.
- PMID 33828091.
- PMID 33828090.
- S2CID 235554619.
- PMID 34593889.
- S2CID 236409612.
- PMID 34249518.
- S2CID 245238634.
- ^ S2CID 235557325.
- S2CID 236289449.
- ^ S2CID 237859264.
- S2CID 234474446.
- S2CID 233937535.
- S2CID 237922832.
- PMID 34804574.
- PMID 33552714.
- S2CID 234814276.
- S2CID 236284813.
- S2CID 235537677.
- PMID 36654455.
- S2CID 252467508.
- S2CID 237235251.
- PMID 34157876.
- .
- S2CID 245090299.
- S2CID 245285654.
- S2CID 233952670.
- S2CID 245401499.
- .
- S2CID 235172524.
- PMID 34154529.
- S2CID 231757917.
- S2CID 235202291.
- S2CID 234497506.
- S2CID 236276543.
- S2CID 240537443.
- S2CID 240257178.
- .
- S2CID 245414037.
- doi:10.1130/G49193.1.
- S2CID 244588555.
- PMID 33732551.
- doi:10.26879/1168.
- S2CID 234812878.
- S2CID 233277410.
- PMID 34330912.