Human: Difference between revisions
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'''Humans''' (''[[Homo sapiens]]'') are the only [[Extant taxon|extant]] members of the [[subtribe]] [[Hominina]]. Together with [[Pan (genus)|chimpanzees]], [[gorilla]]s, and [[orangutan]]s, they are part of the family [[Hominidae]] (the great apes, or ''hominids''). A [[terrestrial animal]], humans are characterized by their [[Human skeletal changes due to bipedalism|erect posture]] and [[Bipedalism|bipedal locomotion]]; high [[manual dexterity]] and heavy tool use compared to other [[animal]]s; open-ended and complex [[Language|language use]] compared to other [[Animal language|animal communications]]; larger, more complex brains than other animals; and highly advanced and organized [[social animal|societies]].<ref>{{cite journal |vauthors=Goodman M, Tagle D, Fitch D, Bailey W, Czelusniak J, Koop B, Benson P, Slightom J |title=Primate evolution at the DNA level and a classification of hominoids |journal=J Mol Evol |volume=30 |issue=3 |pages=260–66 |year=1990 |pmid=2109087 |doi=10.1007/BF02099995|bibcode=1990JMolE..30..260G }}</ref><ref>{{cite web |title=Hominidae Classification |work=Animal Diversity Web @ UMich |url=http://animaldiversity.ummz.umich.edu/site/accounts/classification/Hominidae.html |accessdate=25 September 2006 |url-status=live |archiveurl=https://web.archive.org/web/20061005035254/http://animaldiversity.ummz.umich.edu/site/accounts/classification/Hominidae.html |archivedate=5 October 2006 }}</ref> |
'''Humans''' (''[[Homo sapiens]]'') are the only [[Extant taxon|extant]] members of the [[subtribe]] [[Hominina]]. Together with [[Pan (genus)|chimpanzees]], [[gorilla]]s, and [[orangutan]]s, they are part of the family [[Hominidae]] (the great apes, or ''hominids''). A [[terrestrial animal]], humans are characterized by their [[Human skeletal changes due to bipedalism|erect posture]] and [[Bipedalism|bipedal locomotion]]; high [[manual dexterity]] and heavy tool use compared to other [[animal]]s; open-ended and complex [[Language|language use]] compared to other [[Animal language|animal communications]]; larger, more complex brains than other animals; and highly advanced and organized [[social animal|societies]].<ref>{{cite journal |vauthors=Goodman M, Tagle D, Fitch D, Bailey W, Czelusniak J, Koop B, Benson P, Slightom J |title=Primate evolution at the DNA level and a classification of hominoids |journal=J Mol Evol |volume=30 |issue=3 |pages=260–66 |year=1990 |pmid=2109087 |doi=10.1007/BF02099995|bibcode=1990JMolE..30..260G }}</ref><ref>{{cite web |title=Hominidae Classification |work=Animal Diversity Web @ UMich |url=http://animaldiversity.ummz.umich.edu/site/accounts/classification/Hominidae.html |accessdate=25 September 2006 |url-status=live |archiveurl=https://web.archive.org/web/20061005035254/http://animaldiversity.ummz.umich.edu/site/accounts/classification/Hominidae.html |archivedate=5 October 2006 }}</ref> |
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Early hominins—particularly the [[australopithecine]]s, whose brains and anatomy are in many ways more similar to ancestral non-human [[ape]]s—are less often referred to as "human" than hominins of the genus ''[[Homo]]''.<ref>{{cite journal |author1=Tattersall Ian |author2=Schwartz Jeffrey |year=2009 |title=Evolution of the Genus Homo |url= |journal=Annual Review of Earth and Planetary Sciences |volume=37 |issue= 1|pages=67–92 |doi=10.1146/annurev.earth.031208.100202|bibcode=2009AREPS..37...67T }}</ref> Several of these hominins [[Control of fire by early humans|used fire]], [[Early human migrations|occupied much of Eurasia]], and the lineage that gave rise to ''Homo sapiens'' is thought to have diverged in Africa around 500,000 years ago, with the earliest fossil evidence of evidence of early ''Homo sapiens'' appearing (also in Africa) around 300,000 years ago.<ref name=":0">{{Cite journal|last=Scerri|first=Eleanor M. L.|last2=Thomas|first2=Mark G.|last3=Manica|first3=Andrea|last4=Gunz|first4=Philipp|last5=Stock|first5=Jay T.|last6=Stringer|first6=Chris|last7=Grove|first7=Matt|last8=Groucutt|first8=Huw S.|last9=Timmermann|first9=Axel|last10=Rightmire|first10=G. Philip|last11=d’Errico|first11=Francesco|date=1 August 2018|title=Did Our Species Evolve in Subdivided Populations across Africa, and Why Does It Matter?|url=https://www.cell.com/trends/ecology-evolution/abstract/S0169-5347(18)30117-4|journal=Trends in Ecology & Evolution|language=English|volume=33|issue=8|pages=582–594|doi=10.1016/j.tree.2018.05.005|issn=0169-5347|pmid=30007846|pmc=6092560}}</ref> The oldest early ''H. sapiens'' fossils were found in [[Jebel Irhoud]], [[Morocco]] dating to about 315,000 years ago.<ref name="EA-20190320">{{cite news |author=University of Huddersfield |title=Researchers shed new light on the origins of modern humans – The work, published in Nature, confirms a dispersal of Homo sapiens from southern to eastern Africa immediately preceded the out-of-Africa migration |url=https://www.eurekalert.org/pub_releases/2019-03/uoh-nrs032019.php |date=20 March 2019 |work=[[EurekAlert!]] |access-date=23 March 2019 |author-link=University of Huddersfield }}</ref><ref name="SR-20190318">{{cite journal | vauthors = Rito T, Vieira D, Silva M, Conde-Sousa E, Pereira L, Mellars P, Richards MB, Soares P | display-authors = 6 | title = A dispersal of Homo sapiens from southern to eastern Africa immediately preceded the out-of-Africa migration | journal = Scientific Reports | volume = 9 | issue = 1 | pages = 4728 | date = March 2019 | pmid = 30894612 | pmc = 6426877 | doi = 10.1038/s41598-019-41176-3 | bibcode = 2019NatSR...9.4728R }}</ref><ref>{{cite web|url=https://www.inverse.com/article/39908-ancient-human-evolution-science|title=Everything We Learned in One Year About Thousands of Years of Human Evolution|last=Sloat|first=Sarah|date=4 January 2018|website=Inverse|archiveurl=https://web.archive.org/web/20180126012411/https://www.inverse.com/article/39908-ancient-human-evolution-science|archivedate=26 January 2018|url-status=live|access-date=}}</ref><ref>{{cite journal |author1=Antón, Susan C. |author2=Swisher III, Carl C. |year=2004 |title=Early Dispersals of homo from Africa |url= |journal=Annual Review of Anthropology |volume=33 |issue= |pages=271–96 |doi=10.1146/annurev.anthro.33.070203.144024}}</ref><ref>{{cite journal |author=Trinkaus Erik |year=2005 |title=Early Modern Humans |url= |journal=Annual Review of Anthropology |volume=34 |issue= |pages=207–30 |doi=10.1146/annurev.anthro.34.030905.154913}}</ref> Omo-Kibish I (Omo I) from southern [[Ethiopia]] is the oldest [[anatomically modern Homo sapiens]] skeleton currently known (196 ± 5 ka).<ref>{{Cite journal|last=Hammond|first=Ashley S.|last2=Royer|first2=Danielle F.|last3=Fleagle|first3=John G.|date=Jul 2017|title=The Omo-Kibish I pelvis|journal=Journal of Human Evolution|volume=108|pages=199–219|doi=10.1016/j.jhevol.2017.04.004|issn=1095-8606|pmid=28552208}}</ref><ref>{{Cite journal|last=Fleagle|first=John G.|last2=Brown|first2=Francis H.|last3=McDougall|first3=Ian|date=17 February 2005|title=Stratigraphic placement and age of modern humans from Kibish, Ethiopia|journal=Nature|language=en|volume=433|issue=7027|pages=733–736|doi=10.1038/nature03258|issn=1476-4687|pmid=15716951}}</ref><ref>{{Cite journal|last=López|first=Saioa|last2=van Dorp|first2=Lucy|last3=Hellenthal|first3=Garrett|date=2016-04-21|title=Human Dispersal Out of Africa: A Lasting Debate|url=https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4844272/|journal=Evolutionary Bioinformatics Online|volume=11|issue=Suppl 2|pages=57–68|doi=10.4137/EBO.S33489|issn=1176-9343|pmc=4844272|pmid=27127403}}</ref> Humans began to exhibit evidence of [[behavioral modernity]] at least by about 100-70,000 years ago<ref>{{cite journal | last1 = Henshilwood | first1 = C. S. | last2 = d'Errico | first2 = F. | last3 = Yates | first3 = R. | last4 = Jacobs | first4 = Z. | last5 = Tribolo | first5 = C. | last6 = Duller | first6 = G. A. T. | last7 = Mercier | first7 = N. | last8 = Sealy | first8 = J. C. | last9 = Valladas | first9 = H. | last10 = Watts | first10 = I. | last11 = Wintle | first11 = A. G. | year = 2002 | title = Emergence of modern human behavior: Middle Stone Age engravings from South Africa | url = | journal = Science | volume = 295 | issue = 5558| pages = 1278–1280 | doi=10.1126/science.1067575 | pmid=11786608| bibcode = 2002Sci...295.1278H }}</ref><ref name="Backwell">{{cite journal | author = Backwell L, d'Errico F, Wadley L | year = 2008 | title = Middle Stone Age bone tools from the Howiesons Poort layers, Sibudu Cave, South Africa | url = | journal = Journal of Archaeological Science | volume = 35 | issue = | pages = 1566–1580 | doi = 10.1016/j.jas.2007.11.006 }}</ref><ref name="McBrearty Brooks 2000">{{cite journal|last1=McBrearty|first1=Sally|last2=Brooks|first2=Allison|date=2000|title=The revolution that wasn't: a new interpretation of the origin of modern human behavior|journal=Journal of Human Evolution|volume=39|issue=5|pages=453–563|doi=10.1006/jhev.2000.0435|pmid=11102266}}</ref><ref name="Henshilwood Marean 2003">{{cite journal|last1=Henshilwood|first1=Christopher|last2=Marean|first2=Curtis|date=2003|title=The Origin of Modern Human Behavior: Critique of the Models and Their Test Implications|journal=Current Anthropology|volume=44|issue=5|pages=627–651|doi=10.1086/377665}}</ref><ref>{{citation|last1=Brown|first1=Kyle S. |last2=Marean| first2=Curtis W. |last3=Herries |first3=Andy I.R. |last4=Jacobs |first4=Zenobia |last5=Tribolo |first5=Chantal |last6=Braun |first6=David |last7=Roberts |first7=David L. |last8=Meyer |first8=Michael C. |author9=Bernatchez, J. |date=14 August 2009 |title=Fire as an Engineering Tool of Early Modern Humans| journal=Science |volume=325 |issue=5942 |pages=859–862 |doi=10.1126/science.1175028 |pmid=19679810|bibcode=2009Sci...325..859B }}</ref><ref name="Henshilwood et al. 2011">{{cite journal | author = Henshilwood Christopher S | display-authors = etal | year = 2011 | title = A 100,000-Year-Old Ochre-Processing Workshop at Blombos Cave, South Africa | url = | journal = Science | volume = 334 | issue = | pages = 219–222 | doi = 10.1126/science.1211535 }}</ref> and (according to recent evidence) as far back as around 300,000 years ago, in the [[Middle Stone Age]],<ref name="Brooks">{{Cite journal|title=Long-distance stone transport and pigment use in the earliest Middle Stone Age|journal=Science|volume=360|issue=6384|pages=90–94|year=2018|doi = 10.1126/science.aao2646|pmid=29545508|vauthors=Brooks AS, Yellen JE, Potts R, Behrensmeyer AK, Deino AL, Leslie DE, Ambrose SH, Ferguson JR, d'Errico F, Zipkin AM, Whittaker S, Post J, Veatch EG, Foecke K, Clark JB|bibcode=2018Sci...360...90B}}</ref><ref name="The Atlantic-555674">{{cite news |last=Yong |first=Ed |authorlink=Ed Yong |title=A Cultural Leap at the Dawn of Humanity - New finds from Kenya suggest that humans used long-distance trade networks, sophisticated tools, and symbolic pigments right from the dawn of our species. |url=https://www.theatlantic.com/science/archive/2018/03/a-deeper-origin-of-complex-human-cultures/555674/ |date=15 March 2018 |work=[[The Atlantic]] |accessdate=15 March 2018 }}</ref><ref name="SahlePLOS1">{{Cite journal |last1=Sahle |first1=Y. |last2=Hutchings |first2=W. K. |last3=Braun |first3=D. R. |last4=Sealy |first4=J. C. |last5=Morgan |first5=L. E. |last6=Negash |first6=A. |last7=Atnafu |first7=B. |editor1-last=Petraglia |editor1-first=Michael D |title=Earliest Stone-Tipped Projectiles from the Ethiopian Rift Date to >279,000 Years Ago |doi=10.1371/journal.pone.0078092 |journal=PLoS ONE |volume=8 |issue=11 |pages=e78092 |year=2013 |pmid=24236011 |pmc=3827237 |bibcode=2013PLoSO...878092S }}</ref> (with some features of behavioral modernity possibly beginning earlier, and possibly in parallel with evolutionary brain globularization in ''H. sapiens''). In several [[Early human migrations|waves of migration]], ''H. sapiens'' ventured out of Africa and populated most of the world.<ref name="evolutionthe1st4billionyears">{{cite book |title=Evolution: The First Four Billion Years |author=McHenry, H.M |chapter=Human Evolution |editor1=Michael Ruse |editor2=Joseph Travis |year=2009 |publisher=The Belknap Press of Harvard University Press |location=Cambridge, Massachusetts |isbn=978-0-674-03175-3 |page=[https://archive.org/details/evolutionfirstfo00mich/page/265 265] |chapter-url=https://archive.org/details/evolutionfirstfo00mich/page/265 }}</ref><ref>{{Cite journal|last=Neubauer|first=Simon|last2=Hublin|first2=Jean-Jacques|last3=Gunz|first3=Philipp|date=1 January 2018|title=The evolution of modern human brain shape|url=https://advances.sciencemag.org/content/4/1/eaao5961|journal=Science Advances|language=en|volume=4|issue=1|pages=eaao5961|doi=10.1126/sciadv.aao5961|issn=2375-2548}}</ref> |
Early hominins—particularly the [[australopithecine]]s, whose brains and anatomy are in many ways more similar to ancestral non-human [[ape]]s—are less often referred to as "human" than hominins of the genus ''[[Homo]]''.<ref>{{cite journal |author1=Tattersall Ian |author2=Schwartz Jeffrey |year=2009 |title=Evolution of the Genus Homo |url= |journal=Annual Review of Earth and Planetary Sciences |volume=37 |issue= 1|pages=67–92 |doi=10.1146/annurev.earth.031208.100202|bibcode=2009AREPS..37...67T }}</ref> Several of these hominins [[Control of fire by early humans|used fire]], [[Early human migrations|occupied much of Eurasia]], and the lineage that gave rise to ''Homo sapiens'' is thought to have diverged in Africa around 500,000 years ago, with the earliest fossil evidence of evidence of early ''Homo sapiens'' appearing (also in Africa) around 300,000 years ago.<ref name=":0">{{Cite journal|last=Scerri|first=Eleanor M. L.|last2=Thomas|first2=Mark G.|last3=Manica|first3=Andrea|last4=Gunz|first4=Philipp|last5=Stock|first5=Jay T.|last6=Stringer|first6=Chris|last7=Grove|first7=Matt|last8=Groucutt|first8=Huw S.|last9=Timmermann|first9=Axel|last10=Rightmire|first10=G. Philip|last11=d’Errico|first11=Francesco|date=1 August 2018|title=Did Our Species Evolve in Subdivided Populations across Africa, and Why Does It Matter?|url=https://www.cell.com/trends/ecology-evolution/abstract/S0169-5347(18)30117-4|journal=Trends in Ecology & Evolution|language=English|volume=33|issue=8|pages=582–594|doi=10.1016/j.tree.2018.05.005|issn=0169-5347|pmid=30007846|pmc=6092560}}</ref> The oldest early ''H. sapiens'' fossils were found in [[Jebel Irhoud]], [[Morocco]] dating to about 315,000 years ago.<ref name="EA-20190320">{{cite news |author=University of Huddersfield |title=Researchers shed new light on the origins of modern humans – The work, published in Nature, confirms a dispersal of Homo sapiens from southern to eastern Africa immediately preceded the out-of-Africa migration |url=https://www.eurekalert.org/pub_releases/2019-03/uoh-nrs032019.php |date=20 March 2019 |work=[[EurekAlert!]] |access-date=23 March 2019 |author-link=University of Huddersfield }}</ref><ref name="SR-20190318">{{cite journal | vauthors = Rito T, Vieira D, Silva M, Conde-Sousa E, Pereira L, Mellars P, Richards MB, Soares P | display-authors = 6 | title = A dispersal of Homo sapiens from southern to eastern Africa immediately preceded the out-of-Africa migration | journal = Scientific Reports | volume = 9 | issue = 1 | pages = 4728 | date = March 2019 | pmid = 30894612 | pmc = 6426877 | doi = 10.1038/s41598-019-41176-3 | bibcode = 2019NatSR...9.4728R }}</ref><ref>{{cite web|url=https://www.inverse.com/article/39908-ancient-human-evolution-science|title=Everything We Learned in One Year About Thousands of Years of Human Evolution|last=Sloat|first=Sarah|date=4 January 2018|website=Inverse|archiveurl=https://web.archive.org/web/20180126012411/https://www.inverse.com/article/39908-ancient-human-evolution-science|archivedate=26 January 2018|url-status=live|access-date=}}</ref><ref>{{cite journal |author1=Antón, Susan C. |author2=Swisher III, Carl C. |year=2004 |title=Early Dispersals of homo from Africa |url= |journal=Annual Review of Anthropology |volume=33 |issue= |pages=271–96 |doi=10.1146/annurev.anthro.33.070203.144024}}</ref><ref>{{cite journal |author=Trinkaus Erik |year=2005 |title=Early Modern Humans |url= |journal=Annual Review of Anthropology |volume=34 |issue= |pages=207–30 |doi=10.1146/annurev.anthro.34.030905.154913}}</ref> Discovered in 1967, Omo-Kibish I (Omo I) from southern [[Ethiopia]] is, {{as of|2017|lc=y}}<!-- might be justified to use 2020, but I'm using date of latest cited source -->, the oldest [[anatomically modern Homo sapiens]] skeleton currently known (196 ± 5 ka).<ref>{{Cite journal|last=Hammond|first=Ashley S.|last2=Royer|first2=Danielle F.|last3=Fleagle|first3=John G.|date=Jul 2017|title=The Omo-Kibish I pelvis|journal=Journal of Human Evolution|volume=108|pages=199–219|doi=10.1016/j.jhevol.2017.04.004|issn=1095-8606|pmid=28552208}}</ref><ref>{{Cite journal|last=Fleagle|first=John G.|last2=Brown|first2=Francis H.|last3=McDougall|first3=Ian|date=17 February 2005|title=Stratigraphic placement and age of modern humans from Kibish, Ethiopia|journal=Nature|language=en|volume=433|issue=7027|pages=733–736|doi=10.1038/nature03258|issn=1476-4687|pmid=15716951}}</ref><ref>{{Cite journal|last=López|first=Saioa|last2=van Dorp|first2=Lucy|last3=Hellenthal|first3=Garrett|date=2016-04-21|title=Human Dispersal Out of Africa: A Lasting Debate|url=https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4844272/|journal=Evolutionary Bioinformatics Online|volume=11|issue=Suppl 2|pages=57–68|doi=10.4137/EBO.S33489|issn=1176-9343|pmc=4844272|pmid=27127403}}</ref> Humans began to exhibit evidence of [[behavioral modernity]] at least by about 100-70,000 years ago<ref>{{cite journal | last1 = Henshilwood | first1 = C. S. | last2 = d'Errico | first2 = F. | last3 = Yates | first3 = R. | last4 = Jacobs | first4 = Z. | last5 = Tribolo | first5 = C. | last6 = Duller | first6 = G. A. T. | last7 = Mercier | first7 = N. | last8 = Sealy | first8 = J. C. | last9 = Valladas | first9 = H. | last10 = Watts | first10 = I. | last11 = Wintle | first11 = A. G. | year = 2002 | title = Emergence of modern human behavior: Middle Stone Age engravings from South Africa | url = | journal = Science | volume = 295 | issue = 5558| pages = 1278–1280 | doi=10.1126/science.1067575 | pmid=11786608| bibcode = 2002Sci...295.1278H }}</ref><ref name="Backwell">{{cite journal | author = Backwell L, d'Errico F, Wadley L | year = 2008 | title = Middle Stone Age bone tools from the Howiesons Poort layers, Sibudu Cave, South Africa | url = | journal = Journal of Archaeological Science | volume = 35 | issue = | pages = 1566–1580 | doi = 10.1016/j.jas.2007.11.006 }}</ref><ref name="McBrearty Brooks 2000">{{cite journal|last1=McBrearty|first1=Sally|last2=Brooks|first2=Allison|date=2000|title=The revolution that wasn't: a new interpretation of the origin of modern human behavior|journal=Journal of Human Evolution|volume=39|issue=5|pages=453–563|doi=10.1006/jhev.2000.0435|pmid=11102266}}</ref><ref name="Henshilwood Marean 2003">{{cite journal|last1=Henshilwood|first1=Christopher|last2=Marean|first2=Curtis|date=2003|title=The Origin of Modern Human Behavior: Critique of the Models and Their Test Implications|journal=Current Anthropology|volume=44|issue=5|pages=627–651|doi=10.1086/377665}}</ref><ref>{{citation|last1=Brown|first1=Kyle S. |last2=Marean| first2=Curtis W. |last3=Herries |first3=Andy I.R. |last4=Jacobs |first4=Zenobia |last5=Tribolo |first5=Chantal |last6=Braun |first6=David |last7=Roberts |first7=David L. |last8=Meyer |first8=Michael C. |author9=Bernatchez, J. |date=14 August 2009 |title=Fire as an Engineering Tool of Early Modern Humans| journal=Science |volume=325 |issue=5942 |pages=859–862 |doi=10.1126/science.1175028 |pmid=19679810|bibcode=2009Sci...325..859B }}</ref><ref name="Henshilwood et al. 2011">{{cite journal | author = Henshilwood Christopher S | display-authors = etal | year = 2011 | title = A 100,000-Year-Old Ochre-Processing Workshop at Blombos Cave, South Africa | url = | journal = Science | volume = 334 | issue = | pages = 219–222 | doi = 10.1126/science.1211535 }}</ref> and (according to recent evidence) as far back as around 300,000 years ago, in the [[Middle Stone Age]],<ref name="Brooks">{{Cite journal|title=Long-distance stone transport and pigment use in the earliest Middle Stone Age|journal=Science|volume=360|issue=6384|pages=90–94|year=2018|doi = 10.1126/science.aao2646|pmid=29545508|vauthors=Brooks AS, Yellen JE, Potts R, Behrensmeyer AK, Deino AL, Leslie DE, Ambrose SH, Ferguson JR, d'Errico F, Zipkin AM, Whittaker S, Post J, Veatch EG, Foecke K, Clark JB|bibcode=2018Sci...360...90B}}</ref><ref name="The Atlantic-555674">{{cite news |last=Yong |first=Ed |authorlink=Ed Yong |title=A Cultural Leap at the Dawn of Humanity - New finds from Kenya suggest that humans used long-distance trade networks, sophisticated tools, and symbolic pigments right from the dawn of our species. |url=https://www.theatlantic.com/science/archive/2018/03/a-deeper-origin-of-complex-human-cultures/555674/ |date=15 March 2018 |work=[[The Atlantic]] |accessdate=15 March 2018 }}</ref><ref name="SahlePLOS1">{{Cite journal |last1=Sahle |first1=Y. |last2=Hutchings |first2=W. K. |last3=Braun |first3=D. R. |last4=Sealy |first4=J. C. |last5=Morgan |first5=L. E. |last6=Negash |first6=A. |last7=Atnafu |first7=B. |editor1-last=Petraglia |editor1-first=Michael D |title=Earliest Stone-Tipped Projectiles from the Ethiopian Rift Date to >279,000 Years Ago |doi=10.1371/journal.pone.0078092 |journal=PLoS ONE |volume=8 |issue=11 |pages=e78092 |year=2013 |pmid=24236011 |pmc=3827237 |bibcode=2013PLoSO...878092S }}</ref> (with some features of behavioral modernity possibly beginning earlier, and possibly in parallel with evolutionary brain globularization in ''H. sapiens''). In several [[Early human migrations|waves of migration]], ''H. sapiens'' ventured out of Africa and populated most of the world.<ref name="evolutionthe1st4billionyears">{{cite book |title=Evolution: The First Four Billion Years |author=McHenry, H.M |chapter=Human Evolution |editor1=Michael Ruse |editor2=Joseph Travis |year=2009 |publisher=The Belknap Press of Harvard University Press |location=Cambridge, Massachusetts |isbn=978-0-674-03175-3 |page=[https://archive.org/details/evolutionfirstfo00mich/page/265 265] |chapter-url=https://archive.org/details/evolutionfirstfo00mich/page/265 }}</ref><ref>{{Cite journal|last=Neubauer|first=Simon|last2=Hublin|first2=Jean-Jacques|last3=Gunz|first3=Philipp|date=1 January 2018|title=The evolution of modern human brain shape|url=https://advances.sciencemag.org/content/4/1/eaao5961|journal=Science Advances|language=en|volume=4|issue=1|pages=eaao5961|doi=10.1126/sciadv.aao5961|issn=2375-2548}}</ref> |
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The spread of the [[world population|large and increasing population]] of humans has profoundly [[Holocene extinction|affected]] much of the biosphere and millions of species worldwide. Advantages that explain this evolutionary success include a [[encephalization|larger brain]] with a well-developed [[neocortex]], [[prefrontal cortex]] and [[temporal lobe]]s, which enable advanced abstract [[reasoning]], [[language]], [[problem solving]], [[sociality]], and culture through social learning. Humans use tools more frequently and effectively than any other animal: they are the only extant species to build fires, [[cooking|cook food]], [[clothing|clothe]] themselves, and create and use numerous other [[technology|technologies]] and [[art]]s. |
The spread of the [[world population|large and increasing population]] of humans has profoundly [[Holocene extinction|affected]] much of the biosphere and millions of species worldwide. Advantages that explain this evolutionary success include a [[encephalization|larger brain]] with a well-developed [[neocortex]], [[prefrontal cortex]] and [[temporal lobe]]s, which enable advanced abstract [[reasoning]], [[language]], [[problem solving]], [[sociality]], and culture through social learning. Humans use tools more frequently and effectively than any other animal: they are the only extant species to build fires, [[cooking|cook food]], [[clothing|clothe]] themselves, and create and use numerous other [[technology|technologies]] and [[art]]s. |
Revision as of 03:07, 9 February 2020
Human Ma Middle Pleistocene – Recent
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An adult human male (left) and female (right) from the Akha tribe in Northern Thailand. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Primates |
Suborder: | Haplorhini |
Infraorder: | Simiiformes |
Family: | Hominidae |
Subfamily: | Homininae |
Tribe: | Hominini |
Genus: | Homo |
Species: | H. sapiens
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Binomial name | |
Homo sapiens Linnaeus, 1758
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Subspecies | |
Homo sapiens idaltu White et al., 2003Homo sapiens sapiens
| |
Homo sapiens population density
| |
Synonyms | |
Species synonymy[1]
|
Humans (
Early hominins—particularly the
The spread of the
Humans uniquely use such systems of symbolic communication as language and art to express themselves and exchange ideas, and also organize themselves into purposeful groups. Humans create complex
Though most of human existence has been sustained by hunting and gathering in band societies,[27] many human societies transitioned to sedentary agriculture approximately 10,000 years ago,[28] domesticating plants and animals, thus enabling the growth of civilization. These human societies subsequently expanded, establishing various forms of government, religion, and culture around the world, and unifying people within regions to form states and empires. The rapid advancement of scientific and medical understanding in the 19th and 20th centuries permitted the development of fuel-driven technologies and increased lifespans, causing the human population to rise exponentially. The global human population was estimated to be near 8.1 billion in 2019.[29]
Etymology and definition
In common usage, the word "human" generally refers to the only extant species of the genus Homo—anatomically and behaviorally modern Homo sapiens.
In scientific terms, the meanings of "
The English adjective human is a Middle English loanword from Old French humain, ultimately from Latin hūmānus, the adjective form of homō "man." The word's use as a noun (with a plural: humans) dates to the 16th century.[30] The native English term man can refer to the species generally (a synonym for humanity) as well as to human males, or individuals of either sex (though this latter form is less common in contemporary English).[31]
The species
History
million years ago ) |
Evolution and range
The genus Homo evolved and diverged from other hominins in Africa, after the human clade split from the chimpanzee lineage of the hominids (great apes) branch of the primates. Modern humans, defined as the species Homo sapiens or specifically to the single extant subspecies Homo sapiens sapiens, proceeded to colonize all the continents and larger islands, arriving in Eurasia 125,000–60,000 years ago,[35][36] Australia around 40,000 years ago, the Americas around 15,000 years ago, and remote islands such as Hawaii, Easter Island, Madagascar, and New Zealand between the years 300 and 1280.[37][38]
Evidence from molecular biology
The closest living relatives of humans are chimpanzees (genus
Evidence from the fossil record
There is little fossil evidence for the divergence of the gorilla, chimpanzee and hominin lineages.
The earliest members of the genus Homo are
Anatomical adaptations
Human evolution is characterized by a number of
The human species developed a much larger brain than that of other primates—typically 1,330 cm3 (81 cu in) in modern humans, over twice the size of that of a chimpanzee or gorilla.
The reduced degree of sexual dimorphism is primarily visible in the reduction of the male
Rise of Homo sapiens
By the beginning of the
As early Homo sapiens dispersed, it encountered varieties ofThe "out of Africa" migration of Homo sapiens took place in at least two waves, the first around 130,000 to 100,000 years ago, the second (Southern Dispersal) around 70,000 to 50,000 years ago,[80][81][82] resulting in the colonization of Australia around 65-50,000 years ago,[83][84][85] This recent out of Africa migration derived from East African populations, which had become separated from populations migrating to Southern, Central and Western Africa at least 100,000 years earlier.[86] Modern humans subsequently spread globally, replacing archaic humans (either through competition or hybridization). They inhabited Eurasia and Oceania by 40,000 years ago, and the Americas at least 14,500 years ago.[87][88]
Transition to modernity
Until about 12,000 years ago (the beginning of the
The Neolithic Revolution (the invention of agriculture) took place beginning about 10,000 years ago, first in the Fertile Crescent, spreading through large parts of the Old World over the following millennia, and independently in Mesoamerica about 6,000 years ago. Access to food surplus led to the formation of permanent human settlements, the domestication of animals and the use of metal tools for the first time in history.
Agriculture and sedentary lifestyle led to the emergence of early
Few human populations progressed to
The
With the advent of the
Habitat and population
Early human settlements were dependent on proximity to
Technology has allowed humans to colonize six of the
Human habitation within closed ecological systems in hostile environments, such as Antarctica and outer space, is expensive, typically limited in duration, and restricted to scientific, military, or industrial expeditions. Life in space has been very sporadic, with no more than thirteen humans in space at any given time.[102] Between 1969 and 1972, two humans at a time spent brief intervals on the Moon. As of June 2024, no other celestial body has been visited by humans, although there has been a continuous human presence in space since the launch of the initial crew to inhabit the International Space Station on 31 October 2000.[103] However, other celestial bodies have been visited by human-made objects.[104][105][106]
Since 1800, the human population has increased from one billion[107] to over seven billion.[108] The combined biomass of the carbon of all the humans on Earth in 2018 was estimated at ~ 60 million tons, about 10 times larger than that of all non-domesticated mammals.[109]
In 2004, some 2.5 billion out of 6.3 billion people (39.7%) lived in urban areas. In February 2008, the U.N. estimated that half the world's population would live in urban areas by the end of the year.[110] Problems for humans living in cities include various forms of pollution and crime,[111] especially in inner city and suburban slums. Both overall population numbers and the proportion residing in cities are expected to increase significantly in the coming decades.[112]
Humans have had a dramatic effect on the
Biology
Anatomy and physiology
Most aspects of human physiology are closely
Humans, like most of the other
As a consequence of bipedalism, human females have narrower
Apart from bipedalism, humans differ from chimpanzees mostly in
It is estimated that the worldwide average
Although humans appear hairless compared to other primates, with notable hair growth occurring chiefly on the top of the head,
The
Genetics
Like all mammals, humans are a diploid eukaryotic species. Each somatic cell has two sets of 23 chromosomes, each set received from one parent; gametes have only one set of chromosomes, which is a mixture of the two parental sets. Among the 23 pairs of chromosomes there are 22 pairs of autosomes and one pair of sex chromosomes. Like other mammals, humans have an XY sex-determination system, so that females have the sex chromosomes XX and males have XY.[131]
A rough and incomplete
By comparing the parts of the genome that are not under natural selection and which therefore accumulate mutations at a fairly steady rate, it is possible to reconstruct a genetic tree incorporating the entire human species since the last shared ancestor. Each time a certain mutation (SNP) appears in an individual and is passed on to his or her descendants, a
Human accelerated regions, first described in August 2006,[140][141] are a set of 49 segments of the human genome that are conserved throughout vertebrate evolution but are strikingly different in humans. They are named according to their degree of difference between humans and their nearest animal relative (chimpanzees) (HAR1 showing the largest degree of human-chimpanzee differences). Found by scanning through genomic databases of multiple species, some of these highly mutated areas may contribute to human-specific traits.[citation needed]
The forces of natural selection have continued to operate on human populations, with evidence that certain regions of the genome display directional selection in the past 15,000 years.[142]
Life cycle
As with other mammals,
The zygote divides inside the female's uterus to become an embryo, which over a period of 38 weeks (9 months) of gestation becomes a fetus. After this span of time, the fully grown fetus is birthed from the woman's body and breathes independently as an infant for the first time. At this point, most modern cultures recognize the baby as a person entitled to the full protection of the law, though some jurisdictions extend various levels of personhood earlier to human fetuses while they remain in the uterus.
Compared with other species, human childbirth is dangerous. Painful labors lasting 24 hours or more are not uncommon and sometimes lead to the death of the mother, the child or both.
In developed countries, infants are typically 3–4 kg (7–9 lb) in weight and 50–60 cm (20–24 in) in height at birth.
Evidence-based studies indicate that the life span of an individual depends on two major factors,
Diet
Until the development of agriculture approximately 10,000 years ago, Homo sapiens employed a hunter-gatherer method as their sole means of food collection. This involved combining stationary food sources (such as fruits, grains, tubers, and mushrooms, insect larvae and aquatic mollusks) with
In general, humans can survive for two to eight weeks without food, depending on stored body fat. Survival without water is usually limited to three or four days. About 36 million humans die every year from causes directly or indirectly related to starvation.
Biological variation
No two humans—not even
Most current
The human body's ability to
There is biological variation in the human species—with traits such as
The hue of human skin and hair is determined by the presence of
Structure of variation
Within the human species, the greatest degree of genetic
Males typically have larger
Humans of the same sex are 99.9% genetically identical. There is extremely little variation between human geographical populations, and most of the variation that does occur is at the personal level within local areas, and not between populations.[179][210][211] Of the 0.1% of human genetic differentiation, 85% exists within any randomly chosen local population, be they Italians, Koreans, or Kurds. Two randomly chosen Koreans may be genetically as different as a Korean and an Italian. Any ethnic group contains 85% of the human genetic diversity of the world. Genetic data shows that no matter how population groups are defined, two people from the same population group are about as different from each other as two people from any two different population groups.[179][212][213][214]
Current genetic research has demonstrated that humans on the
Geographical distribution of human variation is complex and constantly shifts through time which reflects complicated human evolutionary history. Most human biological variation is clinally distributed and blends gradually from one area to the next. Groups of people around the world have different frequencies of polymorphic genes. Furthermore, different traits are non-concordant and each have different clinal distribution. Adaptability varies both from person to person and from population to population. The most efficient adaptive responses are found in geographical populations where the environmental stimuli are the strongest (e.g. Tibetans are highly adapted to high altitudes). The clinal geographic genetic variation is further complicated by the migration and mixing between human populations which has been occurring since prehistoric times.[179][218][219][220][221][222]
Human variation is highly non-concordant: most of the genes do not cluster together and are not inherited together. Skin and hair color are not correlated to height, weight, or athletic ability. Human species do not share the same patterns of variation through geography. Skin color varies with latitude and certain people are tall or have brown hair. There is a statistical correlation between particular features in a population, but different features are not expressed or inherited together. Thus, genes which code for superficial physical traits—such as skin color, hair color, or height—represent a minuscule and insignificant portion of the human genome and do not correlate with genetic affinity. Dark-skinned populations that are found in Africa, Australia, and South Asia are not closely related to each other.
Due to practices of group
Psychology
The human brain, the focal point of the
Generally regarded as more capable of these higher order activities, the human brain is believed to be more "intelligent" in general than that of any other known species. While some non-human species are capable of creating structures and
Sleep and dreaming
Humans are generally
Consciousness and thought
Humans are one of the relatively few species to have sufficient self-awareness to recognize themselves in a mirror.[239] Around 18 months most human children are aware that the mirror image is not another person.[240]
The human brain
The physical aspects of the mind and brain, and by extension of the nervous system, are studied in the field of neurology, the more behavioral in the field of psychology, and a sometimes loosely defined area between in the field of psychiatry, which treats mental illness and behavioral disorders. Psychology does not necessarily refer to the brain or nervous system, and can be framed purely in terms of phenomenological or information processing theories of the mind. Increasingly, however, an understanding of brain functions is being included in psychological theory and practice, particularly in areas such as artificial intelligence, neuropsychology, and cognitive neuroscience.
The nature of thought is central to psychology and related fields.
Some philosophers divide consciousness into phenomenal consciousness, which is experience itself, and access consciousness, which is the processing of the things in experience.
Motivation and emotion
Motivation is the driving force of desire behind all deliberate
Happiness, or the state of being happy, is a human emotional condition. The definition of happiness is a common philosophical topic. Some people might define it as the best condition that a human can have—a condition of
Emotion has a significant influence on, or can even be said to control, human behavior, though historically many cultures and philosophers have for various reasons discouraged allowing this influence to go unchecked. Emotional experiences perceived as
In modern scientific thought, certain refined emotions are considered a complex neural trait innate in a variety of
Sexuality and love
This section needs additional citations for verification. (February 2020) |
For humans, sexuality has important social functions: it creates physical intimacy, bonds and hierarchies among individuals, besides ensuring biological
Human choices in acting on sexuality are commonly influenced by social norms which vary between cultures. Restrictions are often determined by religious beliefs or social customs. People can fall anywhere along a continuous scale of sexual orientation.[243] There is considerably more evidence supporting nonsocial, biological causes of sexual orientation than social ones, especially for males.[244] Recent research, including neurology and genetics, suggests that other aspects of human sexuality are biologically influenced as well.[245]
Behavior
This section needs additional citations for verification. (July 2018) |
Human society statistics | |
---|---|
World population[29] | 8.1 billion |
Population density[29][246] | 16/km2 (41/sq mi) by total area 54/km2 (141/sq mi) by land area |
Largest cities[247]
|
|
Most widely spoken native languages[248] | |
Most popular religions[249] | Christianity, Islam, Hinduism, Buddhism, Sikhism, Judaism |
GDP (nominal) [citation needed] |
US$ 36,356,240 million (US$5,797 per capita) |
GDP (PPP) [citation needed] |
$51,656,251 million IND ($8,236 per capita) |
Humans are highly social beings and tend to live in large complex social groups. More than any other creature, humans are capable of using systems of communication for self-expression, the exchange of ideas, and
Culture is defined here as patterns of complex symbolic behavior, i.e. all behavior that is not innate but which has to be learned through social interaction with others; such as the use of distinctive
Language
While many species communicate, language is unique to humans, a defining feature of humanity, and a cultural universal. Unlike the limited systems of other animals, human language is open—an infinite number of meanings can be produced by combining a limited number of symbols. Human language also has the capacity of displacement, using words to represent things and happenings that are not presently or locally occurring, but reside in the shared imagination of interlocutors.[130] Language differs from other forms of communication in that it is modality independent; the same meanings can be conveyed through different media, auditively in speech, visually by sign language or writing, and even through tactile media such as braille. Language is central to the communication between humans, and to the sense of identity that unites nations, cultures and ethnic groups. The invention of writing systems at least five thousand years ago allowed the preservation of language on material objects, and was a major technological advancement. The science of linguistics describes the structure and function of language and the relationship between languages. There are approximately six thousand different languages currently in use, including sign languages, and many thousands more that are extinct.[250]
Gender roles
The sexual division of humans into male, female, and in some societies other genders
Kinship
All human societies organize, recognize and classify types of social relationships based on relations between parents and children (
Ethnicity
Humans often form ethnic groups, such groups tend to be larger than kinship networks and be organized around a common identity defined variously in terms of shared ancestry and history, shared cultural norms and language, or shared biological phenotype. Such ideologies of shared characteristics are often perpetuated in the form of powerful, compelling narratives that give legitimacy and continuity to the set of shared values. Ethnic groupings often correspond to some level of political organization such as the
Society, government, and politics
Society is the system of organizations and institutions arising from interaction between humans. Within a society people can be divided into different groups according to their income, wealth, power, reputation, etc., but the structure of social stratification and the degree of social mobility differs, especially between modern and traditional societies.[258] A state is an organized political community occupying a definite territory, having an organized government, and possessing internal and external sovereignty. Recognition of the state's claim to independence by other states, enabling it to enter into international agreements, is often important to the establishment of its statehood. The "state" can also be defined in terms of domestic conditions, specifically, as conceptualized by Max Weber, "a state is a human community that (successfully) claims the monopoly of the 'legitimate' use of physical force within a given territory."[259]
Government can be defined as the political means of creating and enforcing laws; typically via a bureaucratic hierarchy. Politics is the process by which decisions are made within groups; this process often involves conflict as well as compromise. Although the term is generally applied to behavior within governments, politics is also observed in all human group interactions, including corporate, academic, and religious institutions. Many different political systems exist, as do many different ways of understanding them, and many definitions overlap. Examples of governments include monarchy, Communist state, military dictatorship, theocracy, and liberal democracy, the last of which is considered dominant today. All of these issues have a direct relationship with economics.
Trade and economics
Trade is the voluntary exchange of goods and services, and is a form of
Economics is a social science which studies the production, distribution, trade, and consumption of goods and services. Economics focuses on measurable variables, and is broadly divided into two main branches: microeconomics, which deals with individual agents, such as households and businesses, and macroeconomics, which considers the economy as a whole, in which case it considers aggregate supply and demand for money, capital and commodities. Aspects receiving particular attention in economics are resource allocation, production, distribution, trade, and competition. Economic logic is increasingly applied to any problem that involves choice under scarcity or determining economic value.
War
War is a state of organized armed conflict between
There have been a wide variety of
Material culture and technology
Stone tools were used by proto-humans at least 2.5 million years ago.[261] The controlled use of fire began around 1.5 million years ago. Since then, humans have made major advances, developing complex technology to create tools to aid their lives and allowing for other advancements in culture. Major leaps in technology include the discovery of agriculture—what is known as the Neolithic Revolution, and the invention of automated machines in the Industrial Revolution.
Body culture
Throughout history, humans have altered their appearance by wearing clothing[262] and adornments, by trimming or shaving hair or by means of body modifications.
Body modification is the deliberate altering of the
Philosophy and self-reflection
Philosophy is a discipline or field of study involving the investigation, analysis, and development of ideas at a general, abstract, or fundamental level. It is the discipline searching for a general understanding of reality, reasoning and values. Major fields of philosophy include logic, metaphysics, epistemology, philosophy of mind, and axiology (which includes ethics and aesthetics). Philosophy covers a very wide range of approaches, and is used to refer to a worldview, to a perspective on an issue, or to the positions argued for by a particular philosopher or school of philosophy.
Religion and spirituality
Religion is generally defined as a
Although the exact level of religiosity can be hard to measure,
Art, music, and literature
This section needs additional citations for verification. (March 2018) |
Humans have been producing
Music is a natural
Science
Another unique aspect of human culture and thought is the development of complex methods for acquiring knowledge through observation, quantification, and verification.[citation needed] The scientific method has been developed to acquire knowledge of the physical world and the rules, processes and principles of which it consists, and combined with mathematics it enables the prediction of complex patterns of causality and consequence.[citation needed] An understanding of mathematics is unique to humans, although other species of animal have some numerical cognition.[269]
All of science can be divided into three major branches, the
See also
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Further reading
- Freeman, Scott; Jon C. Herron (2007). Evolutionary Analysis (4th ed.). Pearson Education, Inc. ISBN 0-13-227584-8. pp. 757–61.
- Reich, David (2018). Who We Are And How We Got Here – Ancient DNA and the New Science of the Human Past. ISBN 978-1101870327.
External links
- Archaeology Info
- Homo sapiens – The Smithsonian Institution's Human Origins Program
- "Homo sapiens Linnaeus, 1758" at the Encyclopedia of Life
- View the human genome on Ensembl
- Human Timeline (Interactive) – Smithsonian, National Museum of Natural History (August 2016).
- Media related to Homo sapiens at Wikimedia Commons